Ingensalinae subfam. nov. (Hemiptera: Fulgoromorpha: Fulgoroidea: Inoderbidae), a new planthopper subfamily from mid-Cretaceous Kachin amber from Myanmar

The second genus and species of recently established planthopper family Inoderbidae, Ingensala xiai gen. et sp. nov., is described based on a well-preserved specimen from mid-Cretaceous Kachin (Burmese) amber, and it can be definitely attributed to Inoderbidae mainly based on its head structure, pronotum, and mesonotum without median and lateral carinae and tegmen venation. Ingensala gen. nov. is superficially similar to Eofulgoridium regarding its venation pattern, rather than to the Inoderbidae type genus Inoderbe, and further confirmed that Inoderbidae might descend from the Jurassic planthopper family Fulgoridiidae. The early fork of CuA and the stem CuA bearing many branches also can be found in Jurassic Qiyangiricaniidae and Eocene Weiwoboidae. Ingensala gen. nov. also superficially resembles some Tropiduchidae: Tropiduchinae. The new genus differs from the type genus Inoderbe to a large extent according to its wide head, frons without fastigium, antennae not so elongate, the tectiform condition of wings' position in repose, large, broad and translucent tegmen, triangular basal cell, single CuA1, legs covered with short setae, and the lack of filamentous wax on body. Therefore, two new subfamilies (Inoderbinae stat. nov. and Ingensalinae subfam. nov.) are established for these two genera respectively. The diversification in planthoppers could be the result of pressure of environmental changes during the mid-Cretaceous, and Inoderbidae provides more information for us to understand the Cretaceous stage of Fulgoroidea evolution and diversification.

A mid-Cretaceous fungus, Philothysanus burmanicus gen. et sp. nov. (Ascomycota: Spheciophilaceae), on a thrip (Thysanoptera) in Burmese amber

Male and female thalli of a mid-Cretaceous fungus attached to a thrip (Thysanoptera) in Burmese amber are described as Philothysanus burmanicus gen. et sp. nov. in the family Spheciophilaceae (Ascomycota). Both spermatia and ascospores are present. Currently, Philothysanus burmanicus gen. et sp. nov., together with the previously described Spheciophilia adercia Poinar, represent the oldest known ectoparasitic fungi of insects and their age can be used as a reference point in future phylogenetic studies on the origin of fungi with this life style.

Extant Genus in the Mesozoic: Paleoplatyura Meunier (Diptera: Keroplatidae) Found in the Cretaceous Amber of Myanmar

Three new species of Paleoplatyura Meunier, 1899, i.e., Paleoplatyura agnieszkae sp. nov., P. miae sp. nov., and P. magnifica sp. nov., are described and figured. The concept of the genus is briefly discussed, and its systematic position is clarified. A key to fossil species is provided. The genus Paleoplatyura is described from the Eocene Baltic amber. It is concluded that, in Baltic amber, this group is represented only by the type species, and the identity of the other two species is problematic. No additional specimens have been found so far in this amber. Therefore, the presence of as many as three new species in Burmese amber, certainly belonging to Paleoplatyura, is a confirmation of its occurrence already in the Mesozoic.

Just a Fragment of Undescribed Diversity: Twenty New Oriental and Palearctic Species of Sciaroidea (Diptera), including DNA Sequence Data and Two New Fossil Genera

The following 17 extant new species of Sciaroidea (Diptera: Bibionomorpha) are described: Bolitophila nikolae Ševčík sp. nov. (Bolitophilidae, Taiwan), Catocha jingfui sp. nov. (Cecidomyiidae, Taiwan), Catocha manmiaoe sp. nov. (Cecidomyiidae, Taiwan), Catocha shengfengi sp. nov. (Cecidomyiidae, Taiwan), Planetella taiwanensis sp. nov. (Cecidomyiidae, Taiwan), Diadocidia pseudospinusola sp. nov. (Diadocidiidae, Taiwan), Asioditomyia bruneicola sp. nov. (Ditomyiidae, Brunei), Asioditomyia lacii sp. nov. (Ditomyiidae, Taiwan), Ditomyia asiatica sp. nov. (Ditomyiidae, Thailand), Chetoneura davidi sp. nov. (Keroplatidae, Brunei), Euceroplatus mantici sp. nov. (Keroplatidae, Thailand), Setostylus fangshuoi sp. nov. (Keroplatidae, Taiwan), Platyceridion yunfui sp. nov. (Keroplatidae, Hainan), Terocelion adami sp. nov. (Keroplatidae, Taiwan), Hadroneura martini sp. nov. (Mycetophilidae, Taiwan), Paratinia furcata sp. nov. (Mycetophilidae, Czech Republic, Slovakia), and Nepaletricha sikorai sp. nov. (Sciaroidea incertae sedis, Thailand). Two new genera are described from the mid-Cretaceous Burmese amber, Burmasymmerus gen. nov. (Ditomyiidae, type species Burmasymmerus korneliae sp. nov., including also B. wieslawi sp. nov.), representing the first record of the family Ditomyiidae from the Mesozoic, and Burmatricha gen. nov. (Sciaroidea incertae sedis, type species Burmatricha mesozoica sp. nov.). Molecular phylogeny of Ditomyiidae, based on two DNA markers (28S, COI), as well as that of Catocha Haliday, 1833, based on the mitochondrial COI and 16S fragments, are also presented.

Prototeleia Talamas, Popovici, Shih & Ren: A new genus of Platygastridae from Burmese amber

A new genus and species of parasitoid wasp is described from Burmese amber: Prototeleia kleiogen. nov., sp. nov. (Platygastroidea, Platygastridae). Prototeleia kleio exhibits multiple characters that we consider to be plesiomorphic for Platygastridae. These characters are analyzed and discussed in the context of extant and fossil taxa.

On the Nosodendridae from mid-Cretaceous amber of northern Myanmar (Coleoptera: Nosodendroidea)

Nosodendridae is a small polyphagan beetle family with a sparse fossil record. Herein, the fossil Nosodendridae from mid-Cretaceous Burmese amber (ca. 99 Ma) are systematically reviewed. Nosodendron cretaceum Deng et al. is transferred into Archaenosodendron Li & Cai gen. nov., as A. cretaceum (Deng et al.) comb. nov., primarily based on the morphology of prosternum. Three new species of Archaenosodendron from Burmese amber, A. explanatum Li & Cai sp. nov., A. remotidens Li & Cai sp. nov., and A. angulare Li & Cai sp. nov., are also described and illustrated. A key to nosodendrid genera and species from Burmese amber is provided.

The Darwin wasps (Hymenoptera: Ichneumonidae) of Burmese amber

Three new genera and five new species of Darwin wasps (Ichneumonidae) are described from the mid-Cretaceous Burmese (Kachin) amber: Heteropimpla pulverulenta sp. nov., Rasnichneumon alexandri gen. et sp. nov., R. gracilis sp. nov., Heteroichneumon rasnitsyni gen. et sp. nov., and Rogichneumon braconidicus gen. et sp. nov. The genus Heteropimpla is transferred to Novichneumoninae. The subfamily Novichneumoninae, comprising 6 genera with 8 species, is endemic to Burmese amber, and no other subfamilies of Ichneumonidae are known in the Burmese assemblage. Ichneumonids are extremely rare in Burmese amber, as well as in other Cretaceous amber faunas. However, nine known specimens represent nine different species (one is not formally described due to poor preservation), indicating a serious paleontological record gap for Burmese Darwin wasps.

An aberrant colydiine-like tenebrionoid beetle from mid-Cretaceous Burmese amber (Coleoptera: Tenebrionoidea: Zopheridae)

A new fossil species of the superfamily Tenebrionoidea, Coslonatus rasnitsyni gen. et sp. nov., is described and illustrated from the mid-Cretaceous Burmese amber. The new genus is tentatively placed in the extant subfamily Colydiinae (Zopheridae) based primarily on the 4-segmented tarsi, clubbed antennae, and all freely connected abdominal ventrites. Coslonatus is unique in Colydiinae in having dorsally located antennal insertions and markedly long antennae, suggesting a unique ecological niche of the new species.