extinct species
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Author(s):  
Ajit Pal Singh

Dogs are considered to be the most loyal companion of Mankind since ages. They are considered to be the decedents of wolfs. They hold their decent from antient extinct species of wolfs and recently available grey wolfs. Owing to its loyalty and affectionate nature they are the first domesticated animal in man’s world for 15000 years ago. Firstly, they were used by hunters during the time of hunting and were used as to gather the hunts, as in those days’ agriculture was not in practice. This long relationship between human and dogs made them quite loyal and adaptable to human needs. They can eat anything which a man can provide to them and happily accommodates with Human. There are large number of dogs breed which is available in various countries having their own individual traits. They vary in their shape, size, colours, behaviour and their sensory capabilities. Some of dogs are calm and some are aggressive in nature, due to which their utility changes, as they perform many roles for mankind, such as hunting, herding, pulling loads, protection, assisting police and the military, companionship, therapy and aiding disabled people. This relationship with mankind has earned them a title “Man’s best friend”. There are many popular dog breeds in India such as 1) Beagles 2) German Shepherd 3) Great Dane, 4) Labrador Retriever, 5) Boxer, 6) Rottweiller, 7) Pug, 8) Golden Retriever, 9) Cocker Spaniel 10) Dachshund etc.


2021 ◽  
Author(s):  
Michael Vincent Westbury ◽  
Eline D Lorenzen

(1) Within evolutionary biology, mitochondrial genomes (mitogenomes) provide useful insights at both population and species level. Several approaches are available to assemble mitogenomes. However, most are not suitable for divergent, extinct species, due to the requirement of a reference mitogenome from a conspecific or close relative, and relatively high-quality DNA. (2) Iterative mapping can overcome the lack of a close reference sequence, and has been applied to an array of extinct species. Despite its widespread use, the accuracy of the reconstructed assemblies are yet to be comprehensively assessed. Here, we investigated the influence of mapping software (BWA or MITObim), parameters, and bait reference phylogenetic distance on the accuracy of the reconstructed assembly using two simulated datasets: (i) spotted hyena and various mammalian bait references, and (ii) southern cassowary and various avian bait references. Specifically, we assessed the accuracy of results through pairwise distance (PWD) to the reference conspecific mitogenome, number of incorrectly inserted base pairs (bp), and total length of the reconstructed assembly. (3) We found large discrepancies in the accuracy of reconstructed assemblies using different mapping software, parameters, and bait references. PWD to the reference conspecific mitogenome, which reflected the level of incorrect base calls, was consistently higher with BWA than MITObim. The same was observed for the number of incorrectly inserted bp. In contrast, the total sequence length was lower. Overall, the most accurate results were obtained with MITObim using mismatch values of 3 or 5, and the phylogenetically closest bait reference sequence. Accuracy could be further improved by combining results from multiple bait references. (4) We present the first comprehensive investigation of how mapping software, parameters, and bait reference influence mitogenome reconstruction from ancient DNA through iterative mapping. Our study provides information on how mitogenomes are best reconstructed from divergent, short-read data. By obtaining the most accurate reconstruction possible, one can be more confident as to the reliability of downstream analyses, and the evolutionary inferences made from them.


2021 ◽  
Vol 11 (1) ◽  
Author(s):  
Mairin A. Balisi ◽  
Abhinav K. Sharma ◽  
Carrie M. Howard ◽  
Christopher A. Shaw ◽  
Robert Klapper ◽  
...  

AbstractReconstructing the behavior of extinct species is challenging, particularly for those with no living analogues. However, damage preserved as paleopathologies on bone can record how an animal moved in life, potentially reflecting behavioral patterns. Here, we assess hypothesized etiologies of pathology in a pelvis and associated right femur of a Smilodon fatalis saber-toothed cat, one of the best-studied species from the Pleistocene-age Rancho La Brea asphalt seeps, California, USA, using visualization by computed tomography (CT). The pelvis exhibits massive destruction of the right hip socket that was interpreted, for nearly a century, to have developed from trauma and infection. CT imaging reveals instead that the pathological distortions characterize chronic remodeling that began at birth and led to degeneration of the joint over the animal’s life. These results suggest that this individual suffered from hip dysplasia, a congenital condition common in domestic dogs and cats. This individual reached adulthood but could not have hunted properly nor defended territory on its own, likely relying on a social group for feeding and protection. While extant social felids are rare, these fossils and others with similar pathologies are consistent with a spectrum of social strategies in Smilodon supported by a predominance of previous studies.


Fossil Record ◽  
2021 ◽  
Vol 24 (2) ◽  
pp. 339-346
Author(s):  
Valentine Bouju ◽  
Simon Rosse-Guillevic ◽  
Marion Griffon ◽  
Błażej Bojarski ◽  
Jacek Szwedo ◽  
...  

Abstract. A new, extinct species of Allodia Winnertz is described from early Miocene amber of Ethiopia. Allodia paleoafricana sp. nov. is mostly characterized by the scutum with strong anteromarginal, dorsocentral, and lateral setae and the wing with the stem of the M-fork slightly shorter than the vein r–m and the base of the M4–CuA fork aligned with the base of r–m. The assignment to any of the two subgenera Allodia stricto sensu or Brachycampta Winnertz remains equivocal as the fossil intermingles traits found in both taxa. Allodia is known mostly from the Palearctic region, while only a few species have been described from Africa. In this regard, the new fossil species from Ethiopia brings significant new information regarding the Afrotropical distribution and natural history of the genus.


2021 ◽  
Author(s):  
Filipa Coutinho Soares ◽  
Ricardo Faustino de Lima ◽  
Jorge Mestre Palmeirim ◽  
Pedro Cardoso ◽  
Ana S. L. Rodrigues

Aim: We analyse the functional consequences of the changes in species composition resulting from extinctions and introductions on oceanic island bird assemblages. Specifically, we ask if introduced species have compensated the functional loss resulting from species extinctions. Location: Seventy-four oceanic islands (>100 km2) in the Atlantic, Pacific and Indian Oceans. Time period: Late Holocene. Major taxa studied: Terrestrial and freshwater bird species. Methods: We compiled a species list per island (extinct and extant, native and introduced), and then compiled traits per species. We used single-trait analyses to assess the effects of past species extinctions and introductions on functional composition. Then, we used probabilistic hypervolumes in trait space to calculate functional richness and evenness of original versus present avifaunas of each island (and net change), and to estimate functional originality of extinct and introduced species. Results: The net effects of extinctions and introductions were: an increase in average species richness per island (alpha diversity), yet a decline in diversity across all islands (gamma diversity); an average increase in the prevalence of most functional traits (23 out of 35) yet an average decline functional richness and evenness, associated with the fact that extinct species were functionally more original (when compared to extant natives) than introduced species. Main conclusions: Introduced species are on average offsetting (and even surpassing) the losses of extinct species per island in terms of species richness, and they are increasing the prevalence of most functional traits. However, they are not compensating the loss of functional richness due to extinctions. Current island bird assemblages are becoming functionally poorer, having lost original species and being composed of functionally more homogeneous species. This is likely to have cascading repercussions on the functioning of island ecosystems.


2021 ◽  
Author(s):  
Emanuel Tschopp ◽  
James G Napoli ◽  
Lukardis C M Wencker ◽  
Massimo Delfino ◽  
Paul Upchurch

Abstract Generally, the species is considered to be the only naturally occurring taxon. However, species recognised and defined using different species delimitation criteria cannot readily be compared, impacting studies of biodiversity through Deep Time. This comparability issue is particularly marked when comparing extant with extinct species, because the only available data for species delimitation in fossils is derived from their preserved morphology, which is generally restricted to osteology in vertebrates. Here, we quantify intraspecific, intrageneric, and intergeneric osteological variability in extant species of lacertid lizards using pairwise dissimilarity scores based on a dataset of 253 discrete osteological characters for 99 specimens referred to 24 species. Variability is always significantly lower intraspecifically than between individuals belonging to distinct species of a single genus, which is in turn significantly lower than intergeneric variability. Average values of intraspecific variability and associated standard deviations are consistent (with few exceptions), with an overall average within a species of 0.208 changes per character scored. Application of the same methods to six extinct lacertid species (represented by 40 fossil specimens) revealed that intraspecific osteological variability is inconsistent, which can at least in part be attributed to different researchers having unequal expectations of the skeletal dissimilarity within species units. Such a divergent interpretation of intraspecific and interspecific variability among extant and extinct species reinforces the incomparability of the species unit. Lacertidae is an example where extant species recognised and defined based on a number of delimitation criteria show comparable and consistent intraspecific osteological variability. Here, as well as in equivalent cases, application of those skeletal dissimilarity values to palaeontological species delimitation potentially provides a way to ameliorate inconsistencies created by the use of morphology to define species.


2021 ◽  
Vol 48 (2) ◽  
pp. 244-262
Author(s):  
A. Urry

The study of extinction was rooted in Victorian naturalists’ practices of observation and collection, but presented a challenge to the discipline’s increasing emphasis on empiricism and precision. This paper traces the role of witness testimony and hearsay accounts in early studies of extinction, as preserved in the notebooks of Cambridge zoology professor, Alfred Newton. Beginning in 1850s, Newton and his collaborators sought to trace the histories of suspected extinct species such as the British great bustard and the great auk of Iceland. With its subject absent by definition, the study of extinction relied on hearsay and rumour as well as evidence gleaned from past published accounts. Through methodical attempts to collate diverse and contradictory sources, from eyewitnesses to newspapers to local folklore and gossip, Newton demonstrated the inextricability of human activities from the practice of studying extinction. These attempts to resolve social evidence into scientific certainty were time and again frustrated by the uncertain epistemic status of his sources.


2021 ◽  
Vol 21 (1) ◽  
Author(s):  
Simon G. Scarpetta ◽  
David T. Ledesma ◽  
Christopher J. Bell

Abstract Background Alligator lizards (Gerrhonotinae) are a well-known group of extant North American lizard. Although many fossils were previously referred to Gerrhonotinae, most of those fossils are isolated and fragmentary cranial elements that could not be placed in a precise phylogenetic context, and only a handful of known fossils are articulated skulls. The fossil record has provided limited information on the biogeography and phylogeny of Gerrhonotinae. Results We redescribe a nearly complete articulated fossil skull from the Pliocene sediments of the Anza-Borrego Desert in southern California, and refer the specimen to the alligator lizard genus Elgaria. The fossil is a representative of a newly described species, Elgaria peludoverde. We created a morphological matrix to assess the phylogeny of alligator lizards and facilitate identifications of fossil gerrhonotines. The matrix contains a considerably expanded taxonomic sample relative to previous morphological studies of gerrhonotines, and we sampled two specimens for many species to partially account for intraspecific variation. Specimen-based phylogenetic analyses of our dataset using Bayesian inference and parsimony inferred that Elgaria peludoverde is part of crown Elgaria. The new species is potentially related to the extant species Elgaria kingii and Elgaria paucicarinata, but that relationship was not strongly supported, probably because of extensive variation among Elgaria. We explored several alternative biogeographic scenarios implied by the geographic and temporal occurrence of the new species and its potential phylogenetic placements. Conclusions Elgaria peludoverde is the first described extinct species of Elgaria and provides new information on the biogeographic history and diversification of Elgaria. Our research expands the understanding of phylogenetic relationships and biogeography of alligator lizards and strengthens the foundation of future investigations. The osteological data and phylogenetic matrix that we provided will be critical for future efforts to place fossil gerrhonotines. Despite limited intraspecific sampled sizes, we encountered substantial variation among gerrhonotines, demonstrating the value of exploring patterns of variation for morphological phylogenetics and for the phylogenetic placement of fossils. Future osteological investigations on the species we examined and on species we did not examine will continue to augment our knowledge of patterns of variation in alligator lizards and aid in phylogenetics and fossil placement.


Author(s):  
K E Jones ◽  
R J Brocklehurst ◽  
S E Pierce

Abstract Deciphering the biological function of rare or extinct species is key to understanding evolutionary patterns across the tree of life. While soft tissues are vital determinants of joint function, they are rarely available for study. Therefore, extracting functional signals from skeletons, which are more widely available via museum collections, has become a priority for the field of comparative biomechanics. While most work has focused on the limb skeleton, the axial skeleton plays a critical role in body support, respiration, and locomotion, and is therefore of central importance for understanding broad-scale functional evolution. Here, we describe and experimentally validate AutoBend, an automated approach to estimating intervertebral joint function from bony vertebral columns. AutoBend calculates osteological range of motion (oROM) by automatically manipulating digitally articulated vertebrae while incorporating multiple constraints on motion, including both bony intersection and the role of soft tissues by restricting excessive strain in both centrum and zygapophyseal articulations. Using AutoBend and biomechanical data from cadaveric experiments on cats and tegus, we validate important modeling parameters required for oROM estimation, including the degree of zygapophyseal disarticulation, and the location of the center of rotation. Based on our validation, we apply a model with the center of rotation located within the vertebral disc, no joint translation, around 50% strain permitted in both zygapophyses and discs, and a small amount of vertebral intersection permitted. Our approach successfully reconstructs magnitudes and craniocaudal patterns of motion obtained from ex vivo experiments, supporting its potential utility. It also performs better than more typical methods that rely solely on bony intersection, emphasizing the importance of accounting for soft tissues. We estimated the sensitivity of the analyses to vertebral model reconstruction by varying joint spacing, degree of overlap, and the impact of landmark placement. The effect of these factors was small relative to biological variation craniocaudally and between bending directions. Within, we also present a new approach for estimating joint stiffness directly from oROM and morphometric measurements that can successfully reconstruct the craniocaudal patterns, but not magnitudes, derived from experimental data. Together, this work represents a significant step forward for understanding vertebral function in difficult-to-study (e.g., rare or extinct) species, paving the way for a broader understanding of patterns of functional evolution in the axial skeleton.


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