Evolution is conceptualized as a multi-level phenomenon (subunit–organization–organizational field–national economy) that links organizational and ecological systems. Analysis of population and community-level evolution emphasizes the roles of institutional change (e.g., industry deregulation, globalization, market reforms), technological innovation cycles (e.g., technological discontinuities, dominant designs), entrepreneurs, and social movements as triggers of organizational variation. Institutional and technological change transforms the dynamics of organizational communities by shifting the boundaries of organizational forms, destabilizing or reinforcing existing community structures, giving rise to consensus and/or conflict oriented social movements, and creating opportunities for entrepreneurs and venture capitalists to shape new organizational forms.