John Lloyd Williams was an authority on the arctic-alpine flora of Snowdonia during the late nineteenth century when plant collecting was at its height, but unlike other botanists and plant collectors he did not fully pursue the fashionable trend of forming a complete herbarium. His diligent plant-hunting in a comparatively little explored part of Snowdonia led to his discovering a new site for the rare Killarney fern (Trichomanes speciosum), a feat which was considered a major achievement at the time. For most part of the nineteenth century plant distribution, classification and forming herbaria, had been paramount in the learning of botany in Britain resulting in little attention being made to other aspects of the subject. However, towards the end of the century many botanists turned their attention to studying plant physiology, a subject which had advanced significantly in German laboratories. Rivalry between botanists working on similar projects became inevitable in the race to be first in print as Lloyd Williams soon realized when undertaking his major study on the cytology of marine algae.
