AbstractMotivated by a seminal paper of Kesten et al. (Ann. Probab., 3(1), 1–31, 1975) we consider a branching process with a conditional geometric offspring distribution with i.i.d. random environmental parameters An, n ≥ 1 and with one immigrant in each generation. In contrast to above mentioned paper we assume that the environment is long-tailed, that is that the distribution F of $\xi _{n}:=\log ((1-A_{n})/A_{n})$
ξ
n
:
=
log
(
(
1
−
A
n
)
/
A
n
)
is long-tailed. We prove that although the offspring distribution is light-tailed, the environment itself can produce extremely heavy tails of the distribution of the population size in the n th generation which becomes even heavier with increase of n. More precisely, we prove that, for all n, the distribution tail $\mathbb {P}(Z_{n} \ge m)$
ℙ
(
Z
n
≥
m
)
of the n th population size Zn is asymptotically equivalent to $n\overline F(\log m)$
n
F
¯
(
log
m
)
as m grows. In this way we generalise Bhattacharya and Palmowski (Stat. Probab. Lett., 154, 108550, 2019) who proved this result in the case n = 1 for regularly varying environment F with parameter α > 1. Further, for a subcritical branching process with subexponentially distributed ξn, we provide the asymptotics for the distribution tail $\mathbb {P}(Z_{n}>m)$
ℙ
(
Z
n
>
m
)
which are valid uniformly for all n, and also for the stationary tail distribution. Then we establish the “principle of a single atypical environment” which says that the main cause for the number of particles to be large is the presence of a single very small environmental parameter Ak.