<p>The Middle Eocene Climatic Optimum (MECO) represents an episode of widespread&#160;warming occurring ~40 million years ago. It is characterized by gradual warming over&#160;a period of 500,000 years, leading to a rise in ocean temperatures of about 5&#176; C in the&#160;mid and high-latitudes (Sluijs et al., 2013). Contrary to the traditional understanding&#160;and consensus that accommodation space or downstream factors control stratigraphic&#160;architecture in fluvial successions, we test the hypothesis that upstream factors, rather&#160;than downstream factors, control fluvial architecture through changes in the median&#160;grain size, sediment supply and water discharge with paleoslope being a measurable&#160;proxy to quantify these changes. We test our hypothesis utilizing the natural system&#160;of the Escanilla sediment routing system, encompassing the Middle Eocene Climatic&#160;Optimum. The Escanilla system is an overall prograding system, consisting of 1000 m&#160;thick alluvial and fluvial deposits at the southern-margin of the Tremp-Graus Basin in&#160;the south/central Pyrenees, Spain. Multiple lateral measurements for grain size&#160;distributions and cross-set measurements, flow direction and channel geometry are&#160;taken close to the source (Coll de Vent), at an intermediate location (Lascuarre), and&#160;at a distal part (Olson) of the system for paleohydraulic reconstructions. Drone flight&#160;missions are also undertaken to capture aerial photographs of the field area, which&#160;are required for the construction of 3D photogrammetric models. At Olson, alternating sequences of laterally continuous amalgamated channel bodies and several small&#160;sequences of vertically stacked isolated channel bodies have been identified.&#160;Preliminary results show distinct values of median grain size, dune height, flow depth&#160;and paleoslope for the amalgamated and vertically stacked isolated channel&#160;sequences across the MECO; the addition of our 3D models provide further insight&#160;into the lateral connectivity of the amalgamated units. Our results suggest different&#160;paleohydraulic conditions during the deposition of amalgamated and nonamalgamated&#160;units. This data will also be supported by numerical simulations carried&#160;out to better understand the response of fluvial systems to changes in upstream&#160;factors.</p>