The factors determining individual tree growth response are examined during the 4 years following thinning in experiments in even-aged, 8- or 12-year-old regrowth Eucalyptusregnans F. Muell. forest at two sites in southern Australia. At one site, a vigorous understorey dominated by a sedge developed after the thinning. At that site, light-use efficiency by the trees was unaffected by thinning and the aboveground biomass production by the trees in the thinned stand was substantially less than that in the unthinned stand. At the other site, little understorey developed, light-use efficiency by trees in the thinned stand was greater than that in the unthinned stand, and aboveground biomass production was unaffected by thinning even though the leaf weight of the thinned stand was far below that of the unthinned stand. Where the understorey developed, it was concluded that it competed successfully with the trees for water, thereby reducing production in the thinned stand when compared with the unthinned stand. The individual tree growth response that occurred in the thinned stand at that site appeared to be due soley to the extra light available to individual trees following the canopy opening. Where the understorey did not develop, it was concluded that individual tree growth response was due not only to the extra light available to individual trees but also to the increased availability of belowground resources, most probably soil water. Application of a pre-existing stand growth model suggested that at that site the tendency for increased growth resulting from extra water availability in the thinned stand was just balanced by decreased growth due to lower radiation absorption by the reduced canopy, so that net production was unaffected by thinning.
Predicting individual tree growth using stand-level simulation, diameter distribution, and Bayesian calibration
