Kochiella Poulsen, 1927, and Hadrocephalites new genus (Trilobita: Ptychopariida) from the early Middle Cambrian of western North America

2002 ◽  
Vol 76 (1) ◽  
pp. 76-94 ◽  
Author(s):  
Frederick A. Sundberg ◽  
Linda B. McCollum

Kochaspids are an informal group of ptychopariid trilobites that were both abundant and widespread in the early Middle Cambrian of North America. Based on the reassociation of pygidia and cranidia of some kochaspids, Kochiella Poulsen, 1927, is redefined and Hadrocephalites n. gen. is proposed. Hadrocephalites includes taxa previously assigned by Rasetti and Palmer to Schistometopus Resser, 1938a. Schistometopus is considered nomen dubium. Representatives of Kochiella and Hadrocephalites from the Pioche Shale and Carrara Formation of Nevada are described, including the new species Kochiella rasettii, K. brevaspis, Hadrocephalites lyndonensis, and H. rhytidodes. Other kochaspids previously assigned to Kochaspis Resser, 1935; Eiffelaspis Chang, 1963; Schistometopus; and Kochiella are discussed and some are reassigned. The type specimens of Kochiella augusta (Walcott, 1886); K. crito (Walcott, 1917b); K. chares (Walcott, 1917a); K. mansfieldi Resser, 1939; K. arenosa Resser, 1939; Hadrocephalites carina (Walcott, 1917b), and H. cecinna (Walcott, 1917b) are re-illustrated.

1988 ◽  
Vol 62 (2) ◽  
pp. 218-233 ◽  
Author(s):  
John Mark Malinky

Concepts of the family Hyolithidae Nicholson fide Fisher and the genera Hyolithes Eichwald and Orthotheca Novak have been expanded through time to encompass a variety of morphologically dissimilar shells. The Hyolithidae is here considered to include only those hyolithid species which have a rounded (convex) dorsum; slopes on the dorsum are inflated, and the venter may be flat or slightly inflated. Hyolithes encompasses species which possess a low dorsum and a prominent longitudinal sulcus along each edge of the dorsum; the ligula is short and the apertural rim is flared. The emended concept of Orthotheca includes only those species of orthothecid hyoliths which have a subtriangular transverse outline and longitudinal lirae covering the shell on both dorsum and venter.Eighteen species of Hyolithes and one species of Orthotheca from the Appalachian region and Western Interior were reexamined in light of more modern taxonomic concepts and standards of quality for type material. Reexamination of type specimens of H. similis Walcott from the Lower Cambrian of Newfoundland, H. whitei Resser from the Lower Cambrian of Nevada, H. billingsi Walcott from the Lower Cambrian of Nevada, H. gallatinensis Resser from the Upper Cambrian of Wyoming, and H. partitus Resser from the Middle Cambrian of Alabama indicates that none of these species represents Hyolithes. Hyolithes similis is here included under the new genus Similotheca, in the new family Similothecidae. Hyolithes whitei is designated as the type species of the new genus Nevadotheca, to which H. billingsi may also belong. Hyolithes gallatinensis is referred to Burithes Missarzhevsky with question, and H. partitus may represent Joachimilites Marek. The type or types of H. attenuatus Walcott, H. cecrops Walcott, H. comptus Howell, H. cowanensis Resser, H. curticei Resser, H. idahoensis Resser, H. prolixus Resser, H. resseri Howell, H. shaleri Walcott, H. terranovicus Walcott, and H. wanneri Resser and Howell lack shells and/or other taxonomically important features such as a complete aperture, rendering the diagnoses of these species incomplete. Their names should only be used for the type specimens until better preserved topotypes become available for study. Morphology of the types of H.? corrugatus Walcott and “Orthotheca” sola Resser does not support placement in the Hyolitha; the affinities of these species are uncertain.


1987 ◽  
Vol 61 (5) ◽  
pp. 1047-1057 ◽  
Author(s):  
Wang Cheng-Yuan ◽  
Scott M. Ritter ◽  
David L. Clark

The well-exposed and fossiliferous Permian carbonates in China have yielded Early and Late Permian species of the Sweetognathus complex that permit worldwide stratigraphic evaluation of members of the group. The sporadic appearance of species of Sweetognathus and related genera throughout the Permian in western North America and Iran, in particular, may represent iterative evolution and homeomorphy. The pectiniform element morphologies of the several species are interpreted as most important for evolutionary studies and document a partial Permian biostratigraphy in China that aids in the interpretation of less complete sequences elsewhere. A new genus, Pseudosweetognathus, and four new species, Pseudosweetognathus costatus, Sweetognathus subsymmetricus, S. paraguizhouensis, and Iranognathus nudus, are described.


Zootaxa ◽  
2020 ◽  
Vol 4809 (1) ◽  
pp. 111-131
Author(s):  
STEVEN FEND ◽  
PILAR RODRIGUEZ

Four new species of the lumbriculid genus Eremidrilus Fend & Rodriguez, 2003 are described from Idaho, Montana, and New Mexico. The new species all have a filiform proboscis, club-shaped atria in X, each with two functional vasa deferentia, and spermathecae paired in both XI and XII. Male pores of Eremidrilus artzaini n. sp., E. humboldti n. sp., and E. gilita n. sp. are all associated with distinctive porophores, but these structures are absent in E. montanensis n. sp. The new species are easily distinguished from California Eremidrilus species, all of which have spermathecae only in XI. New observations of type specimens of Eremidrilus allegheniensis (Cook, 1971), a species with two spermathecal segments, known only from eastern USA, confirm differences in atrial morphology, and also document unusual spermathecal porophores and ventral glands. 


2004 ◽  
Vol 136 (6) ◽  
pp. 823-834
Author(s):  
J. Donald Lafontaine ◽  
James T. Troubridge

AbstractAlastriagen. nov. is described, and Callierges tropicalis Schaus is transferred to the genus. Alastria chicosp. nov. is described from western North America and Alastria machosp. nov. is described from Costa Rica. We provide illustrations of the adults and genitalia of all three species, as well as the male genitalia of two related genera, Nedra Clarke and Actinotia Hübner.


2019 ◽  
Vol 151 (04) ◽  
pp. 521-530 ◽  
Author(s):  
Vladimir N. Makarkin ◽  
S. Bruce Archibald ◽  
James E. Jepson

AbstractOne new genus of Inocelliidae (Raphidioptera) with one new species and one undetermined specimen is described from the Eocene of North America: Paraksenocellia borealis new genus, new species from the early Eocene (Ypresian) Okanagan Highlands shale at Driftwood Canyon, British Columbia, Canada (a forewing), and Paraksenocellia species from the middle Eocene (Lutetian) of the Coal Creek Member of the Kishenehn Formation, northwestern Montana, United States of America (a hind wing). These are the oldest records of the family. The new genus possesses many character states that are rare in Inocelliidae, e.g., a very long pterostigma extending to ScP in both the forewings and hind wings; the forewing subcostal space has three crossveins; the forewing and hind wing AA1 are deeply forked; the crossvein between CuA and CuP is located far distad the crossvein 1r-m. Paraksenocellia is confidently a member of the Inocelliidae, as it possesses a proximal shift of the basal crossvein 1r-m (connecting R and M) in the forewing and the loss of the basal crossvein 1r-m in the hind wing, both apomorphies of the family. It shares some character states with the Mesozoic Mesoraphidiidae, which we consider to be mostly stem-group plesiomorphies.


2015 ◽  
Vol 89 (3) ◽  
pp. 411-423 ◽  
Author(s):  
Simon Conway Morris ◽  
Paul A. Selden ◽  
Glade Gunther ◽  
Paul G. Jamison ◽  
Richard A. Robison

AbstractCambrian strata of the Laurentian craton contain numerous examples of Burgess Shale–type faunas. Although displaying a more or less concentric distribution around the cratonal margin, most faunal occurrences are in present-day western North America, extending from the Northwest Territories to California. Nevertheless, the soft-bodied and lightly skeletalized fossils in most of these Lagerstätten are highly sporadic. Here, we extend knowledge of such Middle Cambrian occurrences in Utah with reports of four taxa. An arthropod from the Marjum Formation, Dytikosicula desmatae gen. et sp. nov., is a putative megacheiran. It is most similar to Dicranocaris guntherorum, best known from the younger Wheeler Formation, but differs primarily in the arrangement of pleurae and overall size. Along with a specimen of ?Yohoia sp, a new species of Yohoia, Y. utahana sp. nov., is described. It differs from the type and only known species, Y. tenuis, principally in its larger size and shorter exopods; it is the first description of this genus from outside the Burgess Shale. A new species of a stem-group lophotrochozoan from the Spence Shale, Wiwaxia herka sp. nov., possesses a palisade of dorso-lateral spines that are more robust and numerous than the type species of Wiwaxia, W. corrugata. Another notable taxon is Eldonia ludwigi from the Marjum Formation, which is interpreted as a primitive ambulacrarian (assigned to the cambroernids) and a new specimen of the ?cnidarian Cambrorhytium from the Wheeler Shale is illustrated.


1987 ◽  
Vol 61 (3) ◽  
pp. 595-603 ◽  
Author(s):  
David W. Krause

Baiotomeus is a new genus of ptilodontid multituberculate from the late Torrejonian Land-Mammal Age (late middle Paleocene) of western North America. Baiotomeus douglassi (Simpson), the type species, has been assigned, at one time or another, to Ptilodus, Mimetodon, and Neoplagiaulax. In addition, a second, new species, B. lamberti, from three localities in the Medicine Rocks area of southeastern Montana is reported here. The Medicine Rocks localities are tentatively assigned a latest Torrejonian age, younger than localities yielding B. douglassi and younger than previously suggested on the basis of plesiadapid primates.Multituberculates appear to have attained their highest species richness during the Torrejonian Land-Mammal Age, but the discovery of B. lamberti illustrates that our knowledge of multituberculate diversity is incomplete from even that interval of time.


1986 ◽  
Vol 118 (10) ◽  
pp. 991-1057 ◽  
Author(s):  
Valerie M. Behan-Pelletier

AbstractRepresentatives of the oribatid mite family Ceratozetidae of subarctic western North America, including 14 species in eight genera, are treated. A new genus Laminizetes, and eight new species, Diapterobates siccatus, Trichoribates ogilviensis, Laminizetes fortispinosus, Ceratozetes inupiaq, C. kutchin, C. fjellbergi, Sphaerozetes firthensis, and Melanozetes tanana, are proposed, and Dentizetes rudentiger Hammer, Diapterobates humeralis (Hermann), Neogymnobates luteus (Hammer), Trichoribates striatus Hammer, Sphaerozetes castaneus Hammer, and Melanozetes meridianus Sellnick are redescribed. Immatures of Dentizetes rudentiger and Sphaerozetes firthensis are described. A key to the adults of the 31 species of Ceratozetidae recorded from the western North American arctic and subarctic is given. Relationships among the 12 genera in the Ceratozetidae recorded from the North American arctic and subarctic are discussed.


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