The oldest Inocelliidae (Raphidioptera) from the Eocene of western North America

2019 ◽  
Vol 151 (04) ◽  
pp. 521-530 ◽  
Author(s):  
Vladimir N. Makarkin ◽  
S. Bruce Archibald ◽  
James E. Jepson

AbstractOne new genus of Inocelliidae (Raphidioptera) with one new species and one undetermined specimen is described from the Eocene of North America: Paraksenocellia borealis new genus, new species from the early Eocene (Ypresian) Okanagan Highlands shale at Driftwood Canyon, British Columbia, Canada (a forewing), and Paraksenocellia species from the middle Eocene (Lutetian) of the Coal Creek Member of the Kishenehn Formation, northwestern Montana, United States of America (a hind wing). These are the oldest records of the family. The new genus possesses many character states that are rare in Inocelliidae, e.g., a very long pterostigma extending to ScP in both the forewings and hind wings; the forewing subcostal space has three crossveins; the forewing and hind wing AA1 are deeply forked; the crossvein between CuA and CuP is located far distad the crossvein 1r-m. Paraksenocellia is confidently a member of the Inocelliidae, as it possesses a proximal shift of the basal crossvein 1r-m (connecting R and M) in the forewing and the loss of the basal crossvein 1r-m in the hind wing, both apomorphies of the family. It shares some character states with the Mesozoic Mesoraphidiidae, which we consider to be mostly stem-group plesiomorphies.

1991 ◽  
Vol 65 (5) ◽  
pp. 742-755 ◽  
Author(s):  
A. W. Potter

Cocks and Rong (1989) subdivided the species of the Ordovician brachiopod genus Bimuria into two groups, those with a simple, or not undercut, cardinal process and those with an undercut cardinal process. They placed the second group in a new genus, Cooperea, and grouped Cooperea with Craspedelia in the new subfamily Craspedeliinae of the family Sowerbyellidae. However, the cardinal process of Bimuria ranges from not undercut to undercut within at least two species. Cooperea is therefore placed in synonymy with Bimuria, and Craspedelia is returned to the Bimuriidae.Cocks and Rong (1989) inferred that, with rare exceptions, the cardinal process of the Plectambonitacea evolved from simple to trifid to undercut. The evidence from Bimuria indicates, however, that the undercut process developed more than once and, in Bimuria, directly from the simple type.As an adjunct to the above discussion three new species are described: Bimuria gilbertella, Craspedelia intonsa, and Craspedelia sp. 1. Craspedelia is a rarely reported genus documented here for the first time in western North America (northern California). Additional evidence is introduced that the ratio Lmusbv/Lebv decreased from middle to late Ordovician species of Bimuria.


Zootaxa ◽  
2009 ◽  
Vol 1983 (1) ◽  
pp. 63-65 ◽  
Author(s):  
FASHENG LI ◽  
XINGYUE LIU

The psocid genus Dorypteryx Aaron is a small group in the family Psyllipsocidae, comprising only three species until now. The first Dorypteryx species, D. pallida, was described from North America by Aaron (1883), and subsequently recorded in Europe by Dessart (1976). Smithers (1958) described Dolopteryx domestica from Zimbabwe as the only species in a new genus. However, this genus was later synonymized with Dorypteryx by Lienhard (1977), who re-defined Dorypteryx and redescribed the two known species. After that revision, a third species, D. longipennis, was described by Smithers (1991) from Australia on imported specimens. Despite the low species diversity of Dorypteryx, the geographical range of this genus is rather wide, from the West Palaearctic, Afrotropical, Australian, Nearctic, and Neotropical realms. However, until now Dorypteryx had not been found in the Oriental realm. In the present paper, the genus Dorypteryx is recorded from a domestic habitat in Yunnan Province, China. Based on the contribution by Li (2002), there are three genera and 14 species of Psyllipsocidae in China (Pseudopsyllipsocus Li, Psyllipsocus Selys-Longchamps, and Psocathropos Ribaga). This discovery of Dorypteryx represents the first record of this genus in China, and also in Asia and the Oriental realm.The types of the new species described below, all females, are preserved in alcohol and deposited in the Entomological Museum of China Agricultural University (CAU), Beijing. The terminology of the adult generally follows Smithers (1972).


1964 ◽  
Vol 21 (5) ◽  
pp. 933-939 ◽  
Author(s):  
Richard H. Rosenblatt

A new species, Pholis clemensi, referred to the family Pholidae, is named and described from 12 specimens taken in southern British Columbia waters and the Strait of Juan de Fuca. Pholis clemensi is compared with other members of the genus, and a key is given to the North American species.


Zootaxa ◽  
2021 ◽  
Vol 4980 (1) ◽  
pp. 142-156
Author(s):  
VLADIMIR N. MAKARKIN ◽  
S. BRUCE ARCHIBALD ◽  
ROLF W. MATHEWES

The osmylid subfamily Protosmylinae is revised based on our emended diagnosis: Petrushevskia Martynova, 1958 and Mesosmylidus Jepson et al., 2012 are excluded (both considered Osmylidae incertae sedis), and Sogjuta Martynova, 1958 is transferred to it from the Mesosmylininae. The late Eocene genus Protosmylus Krüger, 1913 is considered a junior synonym of Osmylidia Cockerell, 1908 based on a distinct apomorphy (deeply forked MA in the hind wing), syn. nov. Three new species of Osmylidia from the early Eocene of North America are described: O. donnae sp. nov. from Quilchena, O. glastrai sp. nov. from Republic, Washington, USA, and an unnamed species of Osmylidia is reported from Driftwood Canyon Provincial Park, British Columbia (all localities of the Okanagan Highlands series), and O. taliae sp. nov. from the Green River Formation of Colorado, USA. 


2002 ◽  
Vol 76 (1) ◽  
pp. 76-94 ◽  
Author(s):  
Frederick A. Sundberg ◽  
Linda B. McCollum

Kochaspids are an informal group of ptychopariid trilobites that were both abundant and widespread in the early Middle Cambrian of North America. Based on the reassociation of pygidia and cranidia of some kochaspids, Kochiella Poulsen, 1927, is redefined and Hadrocephalites n. gen. is proposed. Hadrocephalites includes taxa previously assigned by Rasetti and Palmer to Schistometopus Resser, 1938a. Schistometopus is considered nomen dubium. Representatives of Kochiella and Hadrocephalites from the Pioche Shale and Carrara Formation of Nevada are described, including the new species Kochiella rasettii, K. brevaspis, Hadrocephalites lyndonensis, and H. rhytidodes. Other kochaspids previously assigned to Kochaspis Resser, 1935; Eiffelaspis Chang, 1963; Schistometopus; and Kochiella are discussed and some are reassigned. The type specimens of Kochiella augusta (Walcott, 1886); K. crito (Walcott, 1917b); K. chares (Walcott, 1917a); K. mansfieldi Resser, 1939; K. arenosa Resser, 1939; Hadrocephalites carina (Walcott, 1917b), and H. cecinna (Walcott, 1917b) are re-illustrated.


1975 ◽  
Vol 49 (S7) ◽  
pp. 1-74 ◽  
Author(s):  
Dennis R. Kolata

Echinoderms of the Middle Ordovician Platteville and lower Galena Groups of north-central Illinois and south-central Wisconsin are represented by at least seven classes including the Homoiostelea, Cystoidea, Crinoidea, Stelleroidea, Edrioasteroidea, Cyclocystoidea, Echinoidea, and possibly Holothuroidea. The most abundant and diverse echinoderms are monocyclic and dicyclic inadunate and camerate crinoids, of which the cupulocrinids and glyptocrinids are best represented. Thirteen new species and one new genus of crinoids are described:Cremacrinus guttenbergensis, Anulocrinus forrestonensis, Isotomocrinus minutus, Carabocrinus oogyi, Dendrocrinus? springeri, Merocrinus britonensis, Cupulocrinus plattevillensis, C. molanderi, Reteocrinus rocktonensis, R. spinosus, Traskocrinus mahlburgi(n. gen. et sp.),Rhaphanocrinus buckleyi, Glyptocrinus charltoni, andG. pustulosis.One new species of rhombiferan cystoid,Coronocystis durandensis, is also described. Echinoids of the family Bothriocidaridae (Bothriocidaris solemin. sp. andNeobothriocidaris templetonin. sp.) are reported for the first time from North America and appear to be the earliest recorded occurrence. Remains of a third bothriocidarid characterized by coalesced podial pores have also been discovered. Light and scanning electron microscopy of well preserved cyclocystoid specimens reveal a number of previously undescribed morphologic features of the central disc and submarginal ring that may shed some light on their functional morphology.Cupulocrinus gracilisRamsbottom from the Upper Drummuck Group Starfish Bed at Thraive Glen near Girvan, Scotland, is here reassigned toC. drummuckensisn. sp.Echinoderms are associated with an abundant and diverse fauna consisting largely of suspension feeders, primarily strophomenid and orthid brachiopods and trepostome and cryptostome bryozoans. Other common groups include corals, trilobites, sponges, and mollusks. Filamentous brown, foliose red and siphonaceous green algae are associated with the fauna at some localities.Carbonate rocks of both the limestone and dolostone facies of the Platteville (Briton, Walgreen and Forreston Members) and lower Galene Groups in the study area consist primarily of abundant shelly invertebrates occurring as whole and broken, unabraded, commonly articulated remains “floating” in a calcisiltite matrix that is highly bioturbated. Carbonate sediments appear to have been deposited in a near-to below wave base, highly stable, open marine environment characterized by low depositional slopes, good circulation and low terrigenous influx.


1987 ◽  
Vol 61 (5) ◽  
pp. 1047-1057 ◽  
Author(s):  
Wang Cheng-Yuan ◽  
Scott M. Ritter ◽  
David L. Clark

The well-exposed and fossiliferous Permian carbonates in China have yielded Early and Late Permian species of the Sweetognathus complex that permit worldwide stratigraphic evaluation of members of the group. The sporadic appearance of species of Sweetognathus and related genera throughout the Permian in western North America and Iran, in particular, may represent iterative evolution and homeomorphy. The pectiniform element morphologies of the several species are interpreted as most important for evolutionary studies and document a partial Permian biostratigraphy in China that aids in the interpretation of less complete sequences elsewhere. A new genus, Pseudosweetognathus, and four new species, Pseudosweetognathus costatus, Sweetognathus subsymmetricus, S. paraguizhouensis, and Iranognathus nudus, are described.


1995 ◽  
Vol 69 (1) ◽  
pp. 110-122 ◽  
Author(s):  
M. J. Orchard

Exceptionally well preserved Lower Triassic conodonts from Oman include an array ofNeospathodusspecies, many of which are new. Those from the upper Lower Triassic, or Spathian, are described in conjunction with a restudy of conodont type material from Chios, Greece, and an assessment of contemporary collections from Pakistan and western North America. The taxonomic scope of three key species from Chios,Neospathodus homeri, N. triangularis, andN. gondolelloides, is revised. Seven new species are described from Oman:N. abruptus, N. brevissimus, N. brochus, N. crassatus, N. curtatus, N. pusillus, andN. symmetricus;and one new species,N. clinatus, is described from Pakistan. The alliedIcriospathodus collinsoniis also described from Oman. The occurrence and range ofNeospathodusspecies are presented in the context of the ammonoid succession in the Spathian of North America. BothN. homeriandN. triangularis, as revised, have shorter ranges and are more age diagnostic than previously thought.Neospathodus gondolelloidesis a distinct taxon, and not synonymous withChiosella timorensis.Five informal faunal divisions are identified based onNeospathodusand allied species. In ascending stratigraphic order, these are typified byIcriospathodus collinsoni, Neospathodus homeri, N. triangularis, N. symmetricus, andN. gondolelloides.Oman collections represent three of these faunas, which occur also in theColumbitesthroughProhungarites/Subcolumbitesammonoid beds of western U.S.A.


2004 ◽  
Vol 136 (6) ◽  
pp. 823-834
Author(s):  
J. Donald Lafontaine ◽  
James T. Troubridge

AbstractAlastriagen. nov. is described, and Callierges tropicalis Schaus is transferred to the genus. Alastria chicosp. nov. is described from western North America and Alastria machosp. nov. is described from Costa Rica. We provide illustrations of the adults and genitalia of all three species, as well as the male genitalia of two related genera, Nedra Clarke and Actinotia Hübner.


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