A new large-bodied, rupicolous Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) allied to Cnemaspis heteropholis Bauer, 2002 from the Central Western Ghats of Karnataka, India 

Zootaxa ◽  
2020 ◽  
Vol 4801 (1) ◽  
pp. 57-84
Author(s):  
AKSHAY KHANDEKAR ◽  
TEJAS THACKERAY ◽  
SAUNAK PAL ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Body Size ◽  
Natural History ◽  
Western Ghats ◽  
Field Survey ◽  
Sequence Divergence ◽  
Colour Pattern ◽  
Central Western Ghats ◽  
Survey Effort

We describe a new large-bodied, rupicolous Cnemaspis from near Sakleshpur in the Central Western Ghats of Karnataka, India. Cnemaspis magnifica sp. nov. is allied to C. heteropholis and can be diagnosed from all other Indian Cnemaspis by the absence of spine-like scales on flank; heterogeneous dorsal pholidosis, presence of only femoral pores in males, homogenous dorsal pholidosis of tail with small, smooth, regularly arranged, flattened, subimbricate scales, without whorls of enlarged tubercles, median row of sub-caudals enlarged and smooth. The new species differs from its closest congener C. heteropholis in body size, the number of longitudinal ventral scales, number of lamellae under digit I and IV of manus and pes, and subtle colour pattern differences; besides 9.1 % uncorrected ND2 sequence divergence. We provide the first description of topotypic male of C. heteropholis along with data on its distribution and natural history. The discovery of Cnemaspis magnifica sp. nov. with minimal field survey effort indicates the need for further herpetological exploration in the region. 

A new small-bodied, polymorphic Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) allied to C. monticola Manamendra-Arachchi, Batuwita & Pethiyagoda, 2007 from the Central Western Ghats of Karnataka, India

Zootaxa ◽  
2021 ◽  
Vol 4950 (3) ◽  
pp. 501-527
Author(s):  
AKSHAY KHANDEKAR ◽  
TEJAS THACKERAY ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Western Ghats ◽  
Field Survey ◽  
Small Body ◽  
Sequence Divergence ◽  
Systematic Sampling ◽  
Small Body Size ◽  
Central Western Ghats ◽  
Survey Effort ◽  
Dorsal Tubercle

We describe a new small-bodied, polymorphic Cnemaspis from near Hongadahalla village, Sakleshpur in the Central Western Ghats of Karnataka, India. Cnemaspis schalleri sp. nov. is allied to C. monticola and can be diagnosed from all other Indian Cnemaspis by its small body size; presence of spine-like tubercles on flank; heterogeneous dorsal pholidosis; presence of both precloacal and femoral pores in males; dorsal pholidosis of tail heterogeneous, composed of small, weakly keeled, flattened, sub-imbricate scales intermixed with enlarged, strongly keeled, distinctly pointed, conical tubercles forming whorls; slightly enlarged median row of sub-caudal scales, smooth at anterior half and strongly keeled posteriorly. The new species differs from C. monticola based on the number of dorsal tubercle rows at mid-body, the number of paravertebral tubercles, the number of precloacal pores and poreless scales between precloacal and femoral pores; besides 8.9–10.7 % uncorrected ND2 sequence divergence. We also provide the first description of topotypic males of C. monticola along with the first data on live colouration and natural history. Cnemaspis schalleri sp. nov. is the second new Cnemaspis described from around Sakleshpur within a span of less than a year with minimal field survey effort. It is almost certain than the dedicated systematic sampling across the Central Western Ghats will result in the discovery of many undescribed Cnemaspis lineages. 

Two new Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) from the Shevaroy massif, Tamil Nadu, India, with a preliminary ND2 phylogeny of Indian Cnemaspis

Zootaxa ◽  
2019 ◽  
Vol 4609 (1) ◽  
pp. 68
Author(s):  
AKSHAY KHANDEKAR ◽  
NIKHIL GAITONDE ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Body Size ◽  
South Asian ◽  
Western Ghats ◽  
Tamil Nadu ◽  
Elevation Gradient ◽  
Sequence Divergence ◽  
Colour Pattern ◽  
Peninsular India ◽  
The Western Ghats

We present a preliminary ND2 phylogeny of South Asian Cnemaspis, recovering a number of deeply divergent clades within Indian Cnemaspis, endemic to the southern and northern Western Ghats besides the Mysore Plateau and hills of Tamil Nadu. There are a number of unnamed lineages that are >5% divergent on ND2 across the phylogeny, including three from the gracilis clade on an elevation gradient (800–1400 m asl.) around Yercaud in the Shevaroy massif, Salem district, Tamil Nadu. We describe two of these as new species— Cnemaspis shevaroyensis sp. nov. and Cnemaspis thackerayi sp. nov. are both allied to Cnemaspis gracilis and can be diagnosed from all other Indian Cnemaspis by the absence of spine-like scales on flank, heterogeneous dorsal pholidosis, presence of femoral and precloacal pores, tail with enlarged, strongly keeled, conical tubercles forming whorls, a median row of enlarged and smooth sub-caudals. They differ from C. gracilis and each other in body size, the number of tubercles around midbody, the number of tubercles in paravertebral rows, the number of femoral and precloacal pores, the number of poreless scales in-between precloacal pores and between femoral and precloacal pores, and subtle colour pattern differences; besides uncorrected mitochondrial sequence divergence (7.9–16.6 %). We also provide a description of Cnemaspis yercaudensis from its type locality and an additional locality. The discovery of two endemic species and a third unnamed divergent lineage from an isolated massif in peninsular India outside the Western Ghats indicate that many other such understudied hill ranges may harbour high endemic biodiversity. 

A new cryptic Cnemaspis Strauch (Squamata: Gekkonidae) from an isolated granite hill on the Mysore Plateau, Karnataka, India

Zootaxa ◽  
2020 ◽  
Vol 4845 (4) ◽  
pp. 509-528
Author(s):  
AKSHAY KHANDEKAR ◽  
TEJAS THACKERAY ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Body Size ◽  
Endemic Species ◽  
Small Body ◽  
Sequence Divergence ◽  
Colour Pattern ◽  
Continuous Series ◽  
Small Body Size ◽  
Undescribed Species ◽  
The Many

We describe a cryptic new species of the Cnemaspis mysoriensis complex from an isolated rocky hillock near Yadiyur, Mandya District of south Karnataka, India. Cnemaspis stellapulvis sp. nov. can be diagnosed from all other peninsular Indian congeners by its small body size (SVL <32 mm), presence of spine-like tubercles on the flanks, heterogeneous dorsal pholidosis, two or three femoral pores on each thigh separated on either side by eight poreless scales from a continuous series of two precloacal pores in males, tail with enlarged, strongly keeled, conical tubercles forming whorls, median row of sub-caudals smooth and slightly enlarged, and a distinct colour pattern. The new species can be diagnosed from members of C. mysoriensis clade by the number of femoral and precloacal pores and number of poreless scales separating these series, the number of dorsal tubercles rows at mid-body, the number of ventral scales across the belly at mid-body and subtle colour pattern differences; besides 13.4–21.4 % uncorrected ND2 sequence divergence. In this paper, we define the Cnemaspis mysoriensis clade and provide a diagnosis and comparison against peninsular Indian congeners. We also provide major diagnostic characters for members of the C. mysoriensis clade. The discovery of yet another endemic species of Cnemaspis from one of the many isolated granite rocky hills on Mysore Plateau highlights the possibilities of presence of many more cryptic undescribed species. 

A new species of South Asian Cnemaspis (Squamata: Gekkonidae) from the Eastern Ghats, India

Zootaxa ◽  
2020 ◽  
Vol 4802 (3) ◽  
pp. 449-462
Author(s):  
ISHAN AGARWAL ◽  
AARON M BAUER ◽  
AKSHAY KHANDEKAR
Keyword(s):  
New Species ◽  
Body Size ◽  
South Asian ◽  
Western Ghats ◽  
Small Body ◽  
Eastern Ghats ◽  
Evergreen Forest ◽  
Colour Pattern ◽  
The Western Ghats ◽  
A New Species

We describe a new species from within the South Asian clade of the polyphyletic gekkonid genus Cnemaspis from the Velikonda Range, in the Eastern Ghats, Andhra Pradesh, India. The new species is the smallest known Indian gekkonid and can be diagnosed from all Indian congeners by its small body size (SVL <29 mm), the absence of spine-like scales on flank, heterogeneous dorsal pholidosis, presence of precloacal pores and no femoral pores in males, tail with enlarged, strongly keeled, conical tubercles forming whorls, median row of sub-caudals smooth and slightly enlarged, and a distinct colour pattern. This is the 12th species of Cnemaspis to be described from outside the Western Ghats and is the first endemic from the Velikonda region of the Eastern Ghats. The new species is only known from its type locality, a lowland riparian habitat (<200 m asl.) in dry evergreen forest at the base of a ~1200 m asl. peak. This discovery extends the range of the genus in peninsular India by ~130–150 kilometres to the northeast and suggests the genus is likely to be even more widely distributed than understood. The new species is clearly allied to C. mysoriensis, C. otai and C. yercaudensis with which it shares superficial colour pattern and similar body size, but is the first species from outside the Western Ghats in which males lack femoral pores. 

A new medium-sized rupicolous Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) of the C. bangara clade from granite boulder habitats in Krishnagiri, Tamil Nadu, India

Zootaxa ◽  
2021 ◽  
Vol 4969 (2) ◽  
pp. 351-366
Author(s):  
ISHAN AGARWAL ◽  
TEJAS THACKERAY ◽  
AKSHAY KHANDEKAR
Keyword(s):  
New Species ◽  
Tamil Nadu ◽  
Sequence Divergence ◽  
Colour Pattern ◽  
Systematic Sampling ◽  
Anterior Portion ◽  
True Diversity ◽  
Granite Boulder ◽  
Paravertebral Region ◽  
Fourth Member

We describe a new species of the Cnemaspis bangara clade from a rocky hillock near Kugai Periya Mariamman Temple, Krishnagiri fort, Krishnagiri District of Tamil Nadu, India. Cnemaspis krishnagiriensis sp. nov. can be diagnosed from all other peninsular Indian congeners by its medium body size (maximum SVL 40 mm), heterogeneous dorsal pholidosis, spine-like tubercles absent on flanks; presence of four femoral pores on each thigh separated on either side by nine or 10 poreless scales from three continuous precloacal pores in males; tail with enlarged, strongly keeled, conical tubercles forming four whorls on anterior portion, six tubercles in first whorl, four tubercles in second to fourth whorls, enlarged tubercles restricted to only paravertebral region on rest of the tail; median row of sub-caudals smooth and distinctly enlarged, and a unique colour pattern. The new species can be diagnosed from members of C. bangara clade by the number of dorsal tubercles rows at mid-body, the number of enlarged tubercles in paravertebral rows, the number of ventral scales across belly at mid-body, the number longitudinal ventral scales from mental to cloaca, the number of femoral and precloacal pores and poreless scales separating these series, and subtle colour pattern differences; besides 9.2–17.6 % uncorrected ND2 sequence divergence. Cnemaspis krishnagiriensis sp. nov. is the fourth member of the recently described bangara clade and is the first that is known to be distributed <900 m asl.. The discovery of yet another endemic species of Cnemaspis from lower elevations of the Mysore Plateau once again highlights the need of dedicated systematic sampling to uncover the true diversity of Cnemaspis. We also address a nomenclatural issue related to the recently described Cnemaspis stellapulvis Khandekar, Thackeray & Agarwal. 

A new species of Philautus Gistel (Amphibia: Anura: Rhacophoridae) from southern Western Ghats, India

Zootaxa ◽  
2007 ◽  
Vol 1621 (1) ◽  
pp. 1-16 ◽  
Author(s):  
KOTAMBYLU VASUDEVA GURURAJA ◽  
K. P. DINESH ◽  
MUHAMED JAFER PALOT ◽  
C. RADHAKRISHNAN ◽  
T. V. RAMACHANDRA
Keyword(s):  
New Species ◽  
Natural History ◽  
Western Ghats ◽  
Posterior Part ◽  
Upper Eyelid ◽  
Golden Yellow ◽  
Reserve Forest ◽  
Southern Western Ghats ◽  
Kerala State ◽  
A New Species

A new species of the oriental shrub frog genus Philautus is described from Kakkayam Reserve Forest of Calicut district, Kerala state, in the southern Western Ghats. This species is distinguished from congeners by the combination of characters such as body small, elongate, squat and flat; head arched, wider than long; snout short rounded, equal or sub equal to diameter of eye; canthus rostralis rounded; tympanum indistinct but visible; eyes protruding, pupil with striking golden yellow dentition like marks; belly granular; vocal sac unpigmented; fleshy brown to cream yellow dorsum with two distinct golden yellow lateral bands bordered by dark brown from upper eyelid to the posterior part of flanks. A description of the advertisement calls and ecology and natural history notes are also provided for the new species which so far is only known from the type locality.

Another new Cyrtodactylus (Squamata: Gekkonidae) from Guwahati, Assam, India

Zootaxa ◽  
2020 ◽  
Vol 4732 (3) ◽  
pp. 375-392
Author(s):  
JAYADITYA PURKAYASTHA ◽  
MADHURIMA DAS ◽  
SANATH CHANDRA BOHRA ◽  
AARON M. BAUER ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Additional Data ◽  
Sequence Data ◽  
Sequence Divergence ◽  
Northeast India ◽  
The State ◽  
Colour Pattern ◽  
Mitochondrial Sequence ◽  
Guwahati City ◽  
A New Species

We describe a new species of Cyrtodactylus from Guwahati city in the state of Assam, India and provide additional data on the recently described Cyrtodactylus guwahatiensis. Cyrtodactylus urbanus sp. nov. falls in the newly defined khasiensis group within the Indo-Burma clade of Cyrtodactylus and is the poorly supported sister taxon to Cyrtodactylus khasiensis. The new species differs from other members of the khasiensis group in mitochondrial sequence data (12.5–17.1 % uncorrected pairwise ND2 sequence divergence) as well as aspects of morphology including the number and arrangement of precloacal pores in males, the number of mid-ventral scales and paravertebral tubercles, and colour pattern. This is the second Cyrtodactylus endemic to the Guwahati region, the fourth from Assam and the twelfth from Northeast India. 

A novel small-bodied rupicolous Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) from the northern Western Ghats, Maharashtra, India, with comments on the status of C. indraneildasii, Bauer 2000

Zootaxa ◽  
2021 ◽  
Vol 4969 (2) ◽  
pp. 331-350
Author(s):  
AKSHAY KHANDEKAR ◽  
TEJAS THACKERAY ◽  
ISHAN AGARWAL
Keyword(s):  
New Species ◽  
Body Size ◽  
Western Ghats ◽  
Small Body ◽  
Junior Synonym ◽  
Small Body Size ◽  
The Status ◽  
Dorsal Tubercle

We describe a distinct new rupicolous Cnemaspis based on four adult specimens collected from the Kedareshwar caves on Harishchandragad mountain, from the northern Western Ghats, approximately 1200 m asl. in Ahmednagar district of Maharashtra state, India. Cnemaspis uttaraghati sp. nov. can be diagnosed from all other peninsular Indian congeners by its small body size (maximum SVL < 39 mm), heterogeneous dorsal pholidosis, presence of spine-like tubercles on flanks; presence of only femoral pores in males; tail with enlarged, strongly keeled, conical tubercles forming whorls; median row of sub-caudal scales smooth, slightly enlarged, regularly arranged with condition of two slightly larger scales alternating with a large divided scale. The new species is likely to be allied to the C. girii clade and can be diagnosed from its members by its slightly larger body size, presence of spine-like tubercles on flanks, the number of enlarged tubercles in paravertebral rows between forelimb and hindlimb insertions, the number of dorsal tubercle rows at mid-body, the number of ventral scales across belly at mid-body, the number of longitudinal ventral scales from mental to cloaca, and the number of lamellae under digit IV of manus. We also provide a detailed morphological dataset for C. goaensis and C. indraneildasii based on topotypic material and specimens collected from additional localities to confirm C. indraneildasii is a junior synonym of C. goaensis, and provide an updated diagnosis for C. goaensis. 

A new species of Raorchestes (Amphibia: Anura: Rhacophoridae) from mid-elevation evergreen forests of the southern Western Ghats, India

Zootaxa ◽  
2012 ◽  
Vol 3410 (1) ◽  
pp. 19 ◽  
Author(s):  
KADABA SHAMANNA SESHADRI ◽  
KOTAMBYLU VASUDEVA GURURAJA ◽  
NEELAVARA ANANTHARAM ARAVIND
Keyword(s):  
New Species ◽  
Natural History ◽  
Western Ghats ◽  
Advertisement Call ◽  
Closely Related Species ◽  
Evergreen Forests ◽  
Hill Region ◽  
Inner Surface ◽  
Southern Western Ghats ◽  
A New Species

A new species of the shrub frog genus Raorchestes Biju, Souche, Dubois, Dutta and Bossuyt is described as Raorchesteskakachi sp. nov. from Agastyamalai hill region in the southern Western Ghats, India. The small sized Raorchestes (male:24.7–25.8 mm, n = 3 and female: 24.3–34.1 mm, n = 3) is distinguished from all other known congeners by the followingsuite of characters. Snout oval in dorsal view; tympanum indistinct; head wider than long; moderate webbing in feet;colour on dorsum varying from ivory to brown, blotches of dark brown on flanks, brown mottling on throat reducing to-wards vent; inner and outer surface of thigh, inner surface of shank and inner surface of tarsus with a distinct dark brownhorizontal band which extends upto first three toes on upper surface. A detailed description, advertisement call features, ecology, natural history notes and comparison with closely related species are provided for the new species.

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