scholarly journals Twenty years of drought‐mediated change in snag populations in mixed‐conifer and ponderosa pine forests in Northern Arizona

2021 ◽  
Vol 8 (1) ◽  
Author(s):  
Joseph L. Ganey ◽  
Jose M. Iniguez ◽  
Scott C. Vojta ◽  
Amy R. Iniguez
Keyword(s):  
Pinus Ponderosa ◽  
Ponderosa Pine ◽  
Tree Mortality ◽  
Forest Types ◽  
Mixed Conifer ◽  
Size Classes ◽  
Ponderosa Pine Forests ◽  
Northern Arizona ◽  
Mortality Event ◽  
Over Time

Abstract Background Snags (standing dead trees) are important biological legacies in forest systems, providing numerous resources as well as a record of recent tree mortality. From 1997 to 2017, we monitored snag populations in drought-influenced mixed-conifer and ponderosa pine (Pinus ponderosa) forests in northern Arizona. Results Snag density increased significantly in both forest types. This increase was driven largely by a pulse in snag recruitment that occurred between 2002 and 2007, following an extreme drought year in 2002, with snag recruitment returning to pre-pulse levels in subsequent time periods. Some later years during the study also were warmer and/or drier than average, but these years were not as extreme as 2002 and did not trigger the same level of snag recruitment. Snag recruitment was not equal across tree species and size classes, resulting in significant changes in species composition and size-class distributions of snag populations in both forest types. Because trees were far more abundant than snags in these forests, the effect of this mortality pulse on tree populations was far smaller than its effect on snag populations. Snag loss rates increased over time during the study, even though many snags were newly recruited. This may reflect the increasing prevalence of white fir snags and/or snags in the smaller size classes, which generally decay faster than snags of other species or larger snags. Thus, although total numbers of snags increased, many of the newly recruited snags may not persist long enough to be valuable as nesting substrates for native wildlife. Conclusions Increases in snag abundance appeared to be due to a short-term tree mortality “event” rather than a longer-term pattern of elevated tree mortality. This mortality event followed a dry and extremely warm year (2002) embedded within a longer-term megadrought. Climate models suggest that years like 2002 may occur with increasing frequency in the southwestern U.S. Such years may result in additional mortality pulses, which in turn may strongly affect trajectories in abundance, structure, and composition of snag populations. Relative effects on tree populations likely will be smaller, but, over time, also could be significant.

scholarly journals Trends in Snag Populations in Drought-Stressed Mixed-Conifer and Ponderosa Pine Forests (1997–2007)

2012 ◽  
Vol 2012 ◽  
pp. 1-8 ◽  
Author(s):  
Joseph L. Ganey ◽  
Scott C. Vojta
Keyword(s):  
Pinus Ponderosa ◽  
Ponderosa Pine ◽  
Tree Mortality ◽  
Management Practices ◽  
Pine Forests ◽  
Conifer Forest ◽  
Mixed Conifer ◽  
Ecological Processes ◽  
Ponderosa Pine Forests ◽  
Class Composition

Snags provide important biological legacies, resources for numerous species of native wildlife, and contribute to decay dynamics and ecological processes in forested ecosystems. We monitored trends in snag populations from 1997 to 2007 in drought-stressed mixed-conifer and ponderosa pine (Pinus ponderosaDougl.exLaws) forests, northern Arizona. Median snag density increased by 75 and 90% in mixed-conifer and ponderosa pine forests, respectively, over this time period. Increased snag density was driven primarily by a large pulse in drought-mediated tree mortality from 2002 to 2007, following a smaller pulse from 1997 to 2002. Decay-class composition and size-class composition of snag populations changed in both forest types, and species composition changed in mixed-conifer forest. Increases in snag abundance may benefit some species of native wildlife in the short-term by providing increased foraging and nesting resources, but these increases may be unsustainable in the long term. Observed changes in snag recruitment and fall rates during the study illustrate the difficulty involved in modeling dynamics of those populations in an era of climate change and changing land management practices.

Ponderosa pine mortality following fire in northern Arizona

2003 ◽  
Vol 12 (1) ◽  
pp. 7 ◽  
Author(s):  
Charles W. McHugh ◽  
Thomas E. Kolb
Keyword(s):  
Pinus Ponderosa ◽  
Prescribed Fire ◽  
Ponderosa Pine ◽  
Tree Mortality ◽  
Variable Model ◽  
Individual Tree ◽  
Logistic Regression Models ◽  
Ponderosa Pine Forests ◽  
Northern Arizona ◽  
Crown Volume

Sampling of 1367 trees was conducted in the Side wildfire (4 May 1996), Bridger-Knoll wildfire (20 June 1996) and Dauber prescribed fire (9 September 1995) in northern Arizona ponderosa pine forests (Pinus ponderosa). Tree mortality was assessed for 3 years after each fire. Three-year post-fire mortality was 32.4% in the Side wildfire, 18.0% in the Dauber prescribed fire, and 13.9% in the Bridger-Knoll wildfire. In the Dauber and Side fires, 95% and 94% of 3-year post-fire mortality occurred by year 2, versus 76% in the Bridger-Knoll wildfire. Compared with trees that lived for 3 years after fire, dead trees in all fires had more crown scorch, crown consumption, bole scorch, ground char, and bark beetle attacks. Logistic regression models were used to provide insight on factors associated with tree mortality after fire. A model using total crown damage by fire (scorch + consumption) and bole char severity as independent variables was the best two-variable model for predicting individual tree mortality for all fires. The amount of total crown damage associated with the onset of tree mortality decreased as bole char severity increased. Models using diameter at breast height (dbh) and crown volume damage suggested that tree mortality decreased as dbh increased in the Dauber prescribed fire where trees were smallest, and tree mortality increased as dbh increased in the Side and Bridger-Knoll wildfires where trees were largest. Moreover, a U-shaped dbh–mortality distribution for all fires suggested higher mortality for the smallest and largest trees compared with intermediate-size trees. We concluded that tree mortality is strongly influenced by interaction between crown damage and bole char severity, and differences in resistance to fire among different-sized trees can vary among sites.

Corrigendum to: Ponderosa pine mortality following fire in northern Arizona

2003 ◽  
Vol 12 (2) ◽  
pp. 245 ◽  
Author(s):  
Charles W. McHugh ◽  
Thomas E. Kolb
Keyword(s):  
Pinus Ponderosa ◽  
Prescribed Fire ◽  
Ponderosa Pine ◽  
Tree Mortality ◽  
Variable Model ◽  
Individual Tree ◽  
Logistic Regression Models ◽  
Ponderosa Pine Forests ◽  
Northern Arizona ◽  
Crown Volume

Sampling of 1367 trees was conducted in the Side wildfire (4 May 1996), Bridger-Knoll wildfire (20 June 1996) and Dauber prescribed fire (9 September 1995) in northern Arizona ponderosa pine forests (Pinus ponderosa). Tree mortality was assessed for 3 years after each fire. Three-year post-fire mortality was 32.4% in the Side wildfire, 18.0% in the Dauber prescribed fire, and 13.9% in the Bridger-Knoll wildfire. In the Dauber and Side fires, 95% and 94% of 3-year post-fire mortality occurred by year 2, versus 76% in the Bridger-Knoll wildfire. Compared with trees that lived for 3 years after fire, dead trees in all fires had more crown scorch, crown consumption, bole scorch, ground char, and bark beetle attacks. Logistic regression models were used to provide insight on factors associated with tree mortality after fire. A model using total crown damage by fire (scorch + consumption) and bole char severity as independent variables was the best two-variable model for predicting individual tree mortality for all fires. The amount of total crown damage associated with the onset of tree mortality decreased as bole char severity increased. Models using diameter at breast height (dbh) and crown volume damage suggested that tree mortality decreased as dbh increased in the Dauber prescribed fire where trees were smallest, and tree mortality increased as dbh increased in the Side and Bridger-Knoll wildfires where trees were largest. Moreover, a U-shaped dbh–mortality distribution for all fires suggested higher mortality for the smallest and largest trees compared with intermediate-size trees. We concluded that tree mortality is strongly influenced by interaction between crown damage and bole char severity, and differences in resistance to fire among different-sized trees can vary among sites.

scholarly journals Verbenone Inhibits Attraction of Ips pini (Coleoptera: Curculionidae) to Pheromone-Baited Traps in Northern Arizona

2020 ◽  
Vol 113 (6) ◽  
pp. 3017-3020
Author(s):  
Monica L Gaylord ◽  
Stephen R McKelvey ◽  
Christopher J Fettig ◽  
Joel D McMillin
Keyword(s):  
Aggregation Pheromone ◽  
Pinus Ponderosa ◽  
Ponderosa Pine ◽  
Tree Mortality ◽  
Treatment Options ◽  
Current Treatment ◽  
Northern Arizona ◽  
Pine Engraver ◽  
Baited Traps ◽  
Significant Difference

Abstract Recent outbreaks of engraver beetles, Ips spp. De Geer (Coleoptera: Curculionidae; Scolytinae), in ponderosa pine, Pinus ponderosa var. scopulorum Engelm. (Pinales: Pinaceae), forests of northern Arizona have resulted in widespread tree mortality. Current treatment options, such as spraying individual P. ponderosa with insecticides or deep watering of P. ponderosa in urban and periurban settings, are limited in applicability and scale. Thinning stands to increase tree vigor is also recommended, but appropriate timing is crucial. Antiaggregation pheromones, widely used to protect high-value trees or areas against attacks by several species of Dendroctonus Erichson (Coleoptera: Curculionidae; Scolytinae), would provide a feasible alternative with less environmental impacts than current treatments. We evaluated the efficacy of the antiaggregation pheromone verbenone (4,6,6-trimethylbicyclo[3.1.1]hept-3-en-2-one) in reducing attraction of pine engraver, I. pini (Say), to funnel traps baited with their aggregation pheromone in two trapping assays. Treatments included 1) unbaited control, 2) aggregation pheromone (bait), 3) bait with verbenone deployed from a pouch, and 4) bait with verbenone deployed from a flowable and biodegradable formulation (SPLAT Verb, ISCA Technologies Inc., Riverside, CA). Unbaited traps caught no beetles. In both assays, baited traps caught significantly more I. pini than traps with either formulation of verbenone, and no significant difference was observed between the verbenone pouch and SPLAT Verb. In the second assay, we also examined responses of Temnochila chlorodia (Mannerheim) (Coleoptera: Trogositidae), a common bark beetle predator. Traps containing verbenone pouches caught significantly fewer T. chlorodia than the baited control and SPLAT Verb treatments. We conclude that verbenone shows promise for reducing tree mortality from I. pini.

Mark-recapture estimation of snag standing rates in Northern Arizona mixed-conifer and ponderosa pine forests

2015 ◽  
Vol 79 (8) ◽  
pp. 1369-1377 ◽  
Author(s):  
Joseph L. Ganey ◽  
Gary C. White ◽  
Jeffrey S. Jenness ◽  
Scott C. Vojta
Keyword(s):  
Ponderosa Pine ◽  
Pine Forests ◽  
Mixed Conifer ◽  
Mark Recapture ◽  
Ponderosa Pine Forests ◽  
Northern Arizona

scholarly journals Fire Histoy Determination in the Mixed Conifer/Aspen Community of Bryce Canyon National Park

1993 ◽  
Vol 17 ◽  
pp. 31-35
Author(s):  
Michael Jenkins
Keyword(s):  
Pinus Ponderosa ◽  
Ponderosa Pine ◽  
National Park ◽  
Fire History ◽  
Fire Regime ◽  
Mixed Conifer ◽  
Major Objective ◽  
Ponderosa Pine Forests ◽  
Fire Scar ◽  
European Settlement

The major objective of this ongoing study is to document vegetative changes resulting from alteration of the fire regime in the mixed conifer/aspen communities of Bryce Canyon National Park. Previous fire history studies have documented fire return intervals using fire scar analysis of ponderosa pine Pinus ponderosa in the park (Buchannan and Tolman 1983: Wight 1989) and for the Paunsaugunt Plateau (Stein 1988). Numerous other studies have similarly documented the fire regime in pre-European settlement ponderosa pine forests in western North America. The study is being conducted in the more mesic mixed conifer communities at the south end of Bryce Canyon National Park and will specifically document vegetative changes suggested by Roberts et al. (1992) resulting from suppression of frequent low intensity surface fires and overgrazing.

Seasonal fire effects on mixed-conifer forest structure and ponderosa pine resin properties

2006 ◽  
Vol 36 (1) ◽  
pp. 238-254 ◽  
Author(s):  
Daniel DB Perrakis ◽  
James K Agee
Keyword(s):  
Pinus Ponderosa ◽  
Forest Structure ◽  
Ponderosa Pine ◽  
Bark Beetles ◽  
Conifer Forest ◽  
Mixed Conifer ◽  
Pine Resin ◽  
Ponderosa Pine Forests ◽  
Fuel Loads ◽  
Resin Defenses

This study examined the effects of spring and fall restoration burning in an old-growth mixed-conifer – ponderosa pine (Pinus ponderosa Dougl. ex P. & C. Laws.) forest in southern Oregon. Variables measured include fuel loads, forest structure indices, mortality of large ponderosa pines, and pine resin defenses. One year after treatment, reductions in surface fuel loads and changes to forest structure parameters suggested that burning treatments could meet restoration objectives, with fall burns being somewhat more effective than spring burns. However, mortality of pre settlement pines was significantly higher in fall burns than in spring burns, and both were higher than in unburned controls. Bark beetles (Coleoptera: Scolytidae) were important mortality agents within 2 years after burning. Resin defenses (pressure and flow) were variable over the 2 years of postburn study but showed no evidence of decrease in burned trees; rather, resin defenses were significantly higher in burned trees than in controls at several measurement dates. While increased beetle attacks have previously been documented following burning, there has been much less research on resin responses to fire. These findings suggest that current models of beetle–host interactions do not properly explain the effects of prescribed fire in ponderosa pine forests.

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