Taxonomy and phylogeny of the Triassic bivalve families Mysidiellidae Cox, 1964 and Healeyidae new family

2008 ◽  
Vol 82 (3) ◽  
pp. 555-564 ◽  
Author(s):  
M. Hautmann
Keyword(s):  
Outer Shell ◽  
Junior Synonym ◽  
Late Triassic ◽  
Late Paleozoic ◽  
Great Similarity ◽  
Shell Layer ◽  
Shell Microstructure ◽  
General Shape ◽  
New Family

The Mysidiellidae are morphologically isolated among Triassic bivalves but share important characters with Late Paleozoic Ambonychioidea. Apart from a great similarity in the general shape of the shell, the most primitive mysidiellid genus Promysidiella resembles ambonychioids in the presence of a duplivincular-opisthodetic ligament system. Within the Mysidiellidae, this ligament type evolved into the transitional ligament system that characterizes Late Triassic Mysidiella. The phyletic polarity indicates that this evolution probably took place by paedomorphosis. New examinations of the shell microstructure of Mysidiella demonstrate the presence of simple prismatic and possibly foliated structures in the calcitic outer shell layer, which further supports an ambonychioid affinity. Therefore, the Mysidiellidae are removed from the Mytiloidea and assigned to the Ambonychioidea. The poorly known genus Protopis, which was originally included in the Mysidiellidae, probably had a parivincular ligament system and was hence a member of the Heteroconchia. Joannina, which was previously considered a junior synonym of Protopis, is re-established. The hinge margin of Joannina carries a well developed nymph but lacks teeth. These characters as well as its modioliform shape, anterior shell lobe, and pronounced diagonal carina link Joannina with the Late Triassic genus Healeya (Modiomorphoidea). Both taxa are herein placed in the new family Healeyidae, which differs from the morphologically similar Kalenteridae in the absence of elaborated hinge teeth. Protopis, as well as the recently described genera Leidapoconcha, Waijiaoella, and Qingyaniola, are tentatively assigned to the Healeyidae.

Links between shell chemistry and microstructure – A case study using Arctica islandica

2020 ◽  
Author(s):  
Ellen Schnabel ◽  
Kotaro Shirai ◽  
Naoko Murakami-Sugihara ◽  
Klaus Peter Jochum ◽  
Nils Höche ◽  
...  
Keyword(s):  
Environmental Control ◽  
Formation Rate ◽  
Strong Relationship ◽  
Outer Shell ◽  
Ventral Margin ◽  
Shell Layer ◽  
Shell Microstructure ◽  
Arctica Islandica ◽  
Growth Increments ◽  
Strong Control

<p>Bivalves offer outstanding potential as environmental archives. However, vital effects exert a strong control on the incorporation of many trace and minor elements into the shell so that their use as environmental proxies is currently limited. Furthermore, Sr and Mg show a strong relationship to the micrometer-sized shell architecture (shell microstructure), i.e., near growth lines, which are typically dominated by irregular simple/spherulitic prismatic microstructures, the concentrations of these elements are significantly higher than in portions between growth lines (= growth increments, which are microstructurally more complex). In contrast, Ba is uncoupled from the prevailing shell microstructure. To shed more light on these issues, we conducted a combined element chemical (in-situ analysis by means of LA-ICP-MS) and microstructural analyses (using SEM) of shells of <em>Arctica islandica</em> collected alive in NE Iceland.</p><p>According to our findings, (1) contemporaneous shell portions in the hinge and ventral margin (both belonging to the outer shell layer) within individual specimens showed nearly identical Sr/Ca and Mg/Ca values, but Ba/Ca was 1.5 – 2.5 times higher in the ventral margin than in the hinge. (2) In agreement with previous studies, Sr and Mg were strongly elevated near annual growth lines. (3) Along an isochronous transect from the inner portion of the outer shell layer near the myostracum toward the outer shell surface (in the ventral margin), Si/Ca values increased, on average, by 75% ± 11%, whereas Na/Ca values decreased by 7% ± 1%. Along this transect, the shell microstructure gradually changed from crossed-acicular to homogeneous suggesting that Si and Na are linked to the prevailing nanometer-sized shell architecture or underlying physicochemical processes controlling their formation. (4) In the hinge, Ba/Ca, Sr/Ca, Mn/Ca and Mg/Ca attained highest values along the axis of maximum growth, but gradually decreased in slower growing (contemporaneous) shell portions away from that axis. (5) In contemporaneous shell portions (in either the hinge or the ventral margin), the concentration of some elements varied significantly among specimens, whereas others showed little variability. For example, in similar and contemporaneous shell portions of different specimens, Na/Ca values exhibited only little variation (17.4 – 23.7 mmol/mol), whereas Sr/Ca and B/Ca differed more severely (0.3 – 1.6 mmol/mol and 0.04 – 0.07 mmol/mol, respectively; both within growth increments). Despite these inter-specimen chemical differences, the shell microstructure remained largely invariant.</p><p>Our findings firstly suggest that the extrapallial fluid, if it exists at all, is chemically inhomogeneous. This could result from differences in the efficiency of transmembrane ion transport or to differences in shell formation rate along the growing margin (e.g., faster growth in the outer portion of the outer shell layer than in portions closer to the myostracum). Secondly, chemical differences among specimens may be attributed to physiological differences. Thirdly, some elements such as Ba are uncoupled to microstructural properties, but co-vary strongly among specimens suggesting an environmental control on the uptake and incorporation of this element into the shell.</p>

Evolution and phylogenetic significance of cardioidean shell microstructure (Mollusca, Bivalvia)

2001 ◽  
Vol 75 (3) ◽  
pp. 607-643 ◽  
Author(s):  
Jay A. Schneider ◽  
Joseph G. Carter
Keyword(s):  
Phylogenetic Relationships ◽  
Outer Shell ◽  
Data Sets ◽  
Shell Layer ◽  
Shell Microstructure ◽  
Data Set ◽  
Fracture Control ◽  
Selection For ◽  
Morphologic Data ◽  
Combined Data

The shell microstructure of Carboniferous and Triassic permophorids; Triassic and Recent carditids; Devonian, Carboniferous, and Triassic crassatelloideans; and Jurassic through Recent cardioideans is examined in a phylogenetic context, using separate microstructural and morphologic data sets, as well as a combined data set. The microstructural and morphologic data sets are significantly incongruent, but the combined data set suggests that modiomorphoideans (modiomorphids and permophorids) are basal to crassatelloideans; crassatelloideans are basal to carditids (includingSeptocardia), and carditids are basal to cardiids. On the other hand, the possibility of direct permophorid ancestry for the carditid-cardiid clade cannot be excluded, as suggested by the retention of permophorid-like matted (transitional nacreous-porcelaneous) structure in some early carditids and cardiids. In the absence of stratigraphic data and other evidence for phylogenetic relationships, shell microstructure offers limited potential for assessing subfamily-level phylogenetic relationships within the Cardioidea. This is because of microstructural convergences reflecting biomechanical adaptations for fracture control and abrasion resistance, and possibly also selection for metabolic economy of secretion in tropical, oligotrophic habitats. General evolutionary trends in cardiid shell microstructure are nevertheless apparent: Cretaceous cardiids completely replaced an ancestral laminar, matted structure in their inner shell layer with non-laminar porcelaneous structures; evolved better defined CL structure, stronger reflection of the shell margins, and increased thickness or secondary loss of the ancestral prismatic outer shell layer; and, inProtocardia(Pachycardium)stantoni, added inductural deposition. Some Cenozoic cardiids then evolved wider first-order crossed lamellae, non-denticular composite prisms, composite fibrous prisms, ontogenetic submergence of a juvenile non-denticular composite prismatic outer shell layer into the CL middle shell layer, or ontogenetic submergence of the inner part of a juvenile fibrous prismatic outer shell layer into the CL middle shell layer.The shell microstructure ofHemidonax donaciformisis unusual for a cardioidean, and suggests closer affinities with the superfamily Tellinoidea than with the superfamily Cardioidea.Extensive inductural deposits inProtocardia(Pachycardium)stantoniraise the possibility that photosymbiosis evolved among some Mesozoic members of the Protocardiinae, thereby increasing the likelihood that this feature has evolved several times independently in the Cardiidae.Cemented, calcareous periostracal granules or spines are known to occur in modiolopsoideans, mytiloideans, modiomorphids, permophorids, trigonioids, astartids, cardiids, myoids, pholadomyoids, and septibranchoids. Consequently, the presence of these structures is not necessarily indicative of close anomalodesmatan affinities.

Shell microstructure of the Late Carboniferous rostroconch molluscApotocardium lanterna(Branson, 1965)

2003 ◽  
Vol 77 (4) ◽  
pp. 655-673 ◽  
Author(s):  
Nicole S. Rogalla ◽  
Joseph G. Carter ◽  
John Pojeta
Keyword(s):  
Sensu Stricto ◽  
Outer Shell ◽  
Late Carboniferous ◽  
Shell Layer ◽  
Shell Microstructure

The Late Carboniferous bransoniid conocardioideanApotocardium lanterna(Branson, 1965) had an entirely aragonitic shell with a finely prismatic outer shell layer, a predominantly crossed lamellar to complex crossed lamellar middle shell layer, and an “inner” shell layer of finely textured porcelaneous and/or matted structure. This “inner” layer is probably homologous with the inner part of the middle shell layer and the inner layersensu strictoof bivalved molluscs. Shell morphological and microstructural convergences between conocardioids and living heart cockles suggest that at least some conocardioids may have farmed algal endosymbionts in their posterior mantle margins. This symbiosis may have helped conocardioids compete with the biomechanically more efficient bivalves during the latter part of the Paleozoic.

Condensing lenses and shell microstructure in Corculum (Mollusca: Bivalvia)

1997 ◽  
Vol 71 (1) ◽  
pp. 56-61 ◽  
Author(s):  
J. G. Carter ◽  
J. A. Schneider
Keyword(s):  
Light Transmission ◽  
Outer Shell ◽  
Shell Layer ◽  
Shell Microstructure ◽  
Microstructural Modification

The microstructure of the non-window portions of the shell of Corculum cardissa resembles other Fraginae, with predominantly fibrous prismatic outer, branching crossed lamellar middle, and complex crossed lamellar inner layers. Both the anterior and posterior windows in its shell reflect reduced pigmentation and incursion of the outer shell layer, but the posterior windows involve deeper incursion plus reduction of the outer and middle sublayers of the outer shell layer and microstructural modification of the middle shell layer to enhance light transmission. The planoconvex shape of the posterior windows has more likely evolved to direct and focus light toward the deeper, zooxanthellae-rich gills and anterior mantle, than to merely disperse light.

Nacre in an early gryphaeid bivalve (Mollusca)

1994 ◽  
Vol 68 (6) ◽  
pp. 1405-1408 ◽  
Author(s):  
Christopher A. McRoberts ◽  
Joseph G. Carter
Keyword(s):  
Outer Layer ◽  
Outer Shell ◽  
Late Triassic ◽  
Left Valve ◽  
Shell Microstructure ◽  
Shell Surface ◽  
Prismatic Structure ◽  
Dark Material ◽  
Foliated Structure

McRoberts (1992, figs. 4.13, 4.14, 6.8) illustrated the shell microstructure of late Triassic Gryphaea (Gryphaea) arcuataeformis Kiparisova, 1936, and Gryphaea (Gryphaea) nevadensis McRoberts, 1992. McRoberts (1992, p. 33) described the left valve of G. arcuataeformis as showing “neomorphosed calcite with multiple laminae of ?prismatic structure perpendicular to [the] outer shell surface ….” He described the left valve of G. nevadensis as consisting of two distinct layers of neomorphosed calcite:“…an outer layer with ?prismatic structure occasionally with bands of dark material (?micritic matrix), and a much thinner inner layer with ?cross-foliated structure ….” Subsequent study has shown these microstructural diagnoses to be inaccurate. They are revised as follows.

Correcting a 135-year error: Limulidae Leach, 1819 (Chelicerata, Xiphosura) is the proper authority, not Limulidae Zittel, 1885

2021 ◽  
pp. 1-2
Author(s):  
Philip M. Novack-Gottshall ◽  
Roy E. Plotnick
Keyword(s):  
Horseshoe Crab ◽  
Marine Species ◽  
Limulus Polyphemus ◽  
Junior Synonym ◽  
Living Fossil ◽  
New Family ◽  
The Family ◽  
The World ◽  
Modern Genus

The horseshoe crab Limulus polyphemus (Linnaeus, 1758) is a famous species, renowned as a ‘living fossil’ (Owen, 1873; Barthel, 1974; Kin and Błażejowski, 2014) for its apparently little-changed morphology for many millions of years. The genus Limulus Müller, 1785 was used by Leach (1819, p. 536) as the basis of a new family Limulidae and synonymized it with Polyphemus Lamarck, 1801 (Lamarck's proposed but later unaccepted replacement for Limulus, as discussed by Van der Hoeven, 1838, p. 8) and Xyphotheca Gronovius, 1764 (later changed to Xiphosura Gronovius, 1764, another junior synonym of Limulus). He also included the valid modern genus Tachypleus Leach, 1819 in the family. The primary authority of Leach (1819) is widely recognized in the neontological literature (e.g., Dunlop et al., 2012; Smith et al., 2017). It is also the authority recognized in the World Register of Marine Species (WoRMS Editorial Board, 2021).

scholarly journals New data on the Dogger - Cenomanian stratigraphy of Tripolitza series in Central Crete

2018 ◽  
Vol 34 (2) ◽  
pp. 565
Author(s):  
Α. ΖΑΜΠΕΤΑΚΗ - ΛΕΚΚΑ ◽  
Α. ΑΛΕΞΟΠΟΥΛΟΣ
Keyword(s):  
Early Cretaceous ◽  
Late Eocene ◽  
Depositional Environments ◽  
Late Triassic ◽  
Late Paleozoic ◽  
Carbonate Microfacies ◽  
Aegean Islands ◽  
Lower Aptian ◽  
Sedimentation Conditions

Tripolitza series represents the eastern part of the Gavrovo - Tripolitza platform. It outcrops in central and southeastern Peloponnesus, Crete and the Aegean islands. Its stratigraphie column starts with a volcano-sedimentary, clastic sequences (the Tyros beds), of Late Paleozoic to Late Triassic age, followed by a carbonate series of Late Triassic to Late Eocene age and a tertiary flysch. On account of intense dolomitisation as well as of rarity of fossils, the stratigraphy of Tripolitza series is not so well known as the Gavrovo series. Recent investigation attempts to complete the puzzle of Tripolitza's stratigraphie column and to reconstruct the paleogeographic sedimentation conditions. In this paper, we study some stratigraphie sections, which are taken in different places in northern – central Crete(Fig.l). New data about the stratigraphy and the sedimentation conditions of the Tripolitza series during Late Dogger to Cenomanian, complete older ones and provide information about the paleogeographic evolution of the platform. Late Dogger is determinated by Pfenderina salernitana. It is overlain by a carbonate series containing Cladocoropsis mirabilis and Macroporella sellii, dating Early Malm. Early Malm (Oxfordian - Early Kimmeridgian) is characterized by the presence of Cladocoropsis mirabilis, Kurnubia palastiniensis, Neokilianina rahonensis, Parurgonina caelinensis. Late Malm (Late Kimmeridgian - Portlandian) is characterized by Clypeina jurassica and Kurnubia palastiniensis. Early Cretaceous (Valanginian - Barremian) is determinated by Salpingoporella katzeri and Orbitolinopsis capuensis. Early Aptian is determinated by Palorbitolina lenticularis, Salpingoporella dinarica, Debarina hahounerensis, Pseudocyclammina hedbergi. Late(?) Aptian comprises Sabaudia minuta, Cuneolina hensoni, Cuneolina laurentii, Glomospira urgoniana. Albian is characterized by the disappearence of Cuneolina hensoni and Cuneolina laurentii, while Praechrysalidina infracretacea and Cretacicladus minervini are present. Late Albian is determined by the presence of "Coskinolina" bronnimanni. Early Cenomanian is not determinated by characteristic microfossils. Upper Cenomanian is overlain in comformity with upper Albian - lower Cenomanian carbonates. It is characterized by Chrysalidina gradata, Pseudorhapydionina dubia, Pseudorhapydionina laurinensis, Nummoloculina heimi, Broeckina balcanica, Nezzazata gyra, Biconcava bentori, Trochospira anvimelechi. Sedimentation took place in a peritidal environment. We observe alternations of subtidal and intratidal to supratidal conditions of sedimentation. Comparison between the carbonate microfacies of different sections representing synchronous deposits on different places of the platform, show lateral differentiation of depositional environments, from subtidal to supratidal, even supported short and local emersive episodes, (compare lower Aptian deposits of Profitis Ilias and Pinakianou sections, upper Albian deposits of Kythia and Karouzanos sections in present paper, as well as upper Cenomanian deposits of Karouzanos section in this paper, Louloudaki section (ZAMBETAKIS-LEKKAS et al. 1995 and Vitina section ZAMBETAKIS et al. 1988, Varassova section BERNIER & FLEURY 1980, Gavrovo mountain I.G.R.S. & I.F.P. 1966)(Fig.2). Similar sedimentation characterize the perimediterranean platforms during this period (SARTONI & CRESCENTI 1962, DE CASTRO 1962, FARINACCI & RADOICIC 1964, GUSIC 1969, GUSIC, NIKLER & SOKAC 1971, VELIC 1977, CHIOCCHINI et. al.1979, LUPERTO SINNI & MASSE 1993).

scholarly journals A rare case of an evolutionary late and ephemeral biomineralization: tunicates with composite calcareous skeletons

2020 ◽  
Vol 94 (4) ◽  
pp. 748-757
Author(s):  
Jobst Wendt
Keyword(s):  
New Species ◽  
New Genus ◽  
Lower Jurassic ◽  
Late Triassic ◽  
Cambrian Explosion ◽  
New Order ◽  
Colonial Ascidian ◽  
New Family ◽  
The Family ◽  
Almost All

AbstractIn contrast to almost all other invertebrate phyla that constructed biomineralized skeletons during the “Cambrian explosion” and maintained them during the entire fossil record, ascidian tunicates evolved this protective and stabilizing advantage only during the Permian, although soft-bodied representatives of this subphylum made their first appearance already in the early Cambrian. It remains enigmatic why these compound calcareous skeletons persisted only until the Late Triassic, subsequently followed by less-rigid internal skeletons from the Lower Jurassic onwards, which consist of scattered isolated spicules only. In addition to recently described aragonitic ascidian exoskeletons from the Permian and Triassic, new discoveries of similar, but colonial ascidian compound endoskeletons in the lower Carnian exhibit a short-living branch of this group, which moreover contain the first indubitable calcareous spicules. The latter are embedded in the solid endoskeleton, which is composed of polygonal aragonitic plates with smooth outer and zigzag lined inner boundaries. They consist of irregular, parallel (orthogonal), or fan-shaped (clinogonal) arrangements of acicular aragonite crystals. The following taxa are described as new: order Cassianomorpha new order with the family Cassianosomidae new family and the genus Toscanisoma new genus with the species T. multipartitum new species and T. triplicatum new species.UUID: http://zoobank.org/03555353-cdab-42e8-8e99-9bfce15fa249

Generic concepts in the Platycrinitidae Austin and Austin, 1842 (class Crinoidea)

2009 ◽  
Vol 83 (5) ◽  
pp. 694-717 ◽  
Author(s):  
William I. Ausich ◽  
Thomas W. Kammer
Keyword(s):  
Type Species ◽  
Western Europe ◽  
Sensu Stricto ◽  
Species Level ◽  
Junior Synonym ◽  
Late Paleozoic ◽  
New Genera ◽  
The Family ◽  
Nomina Dubia ◽  
Stratigraphic Ranges

Platycrinitesis traditionally one of the more recognizable crinoids, a camerate crinoid with very few if any fixed brachials or interradials and a helically twisted column. Accordingly, many taxa have been assigned to this genus. With a better understanding of the Platycrinitidae, these characters actually unite the family Platycrinitidae rather than the genus. Further, use of different genus-diagnostic characters in Western Europe versus North America has resulted in a confused systematics for this important late Paleozoic family. Here, we objectively define genera within the Platycrinitidae and assign all species to either newly defined or newly named genera. A phylogenetic hypothesis, incorporating both parsimony-based character analysis and stratigraphic ranges, of the genera within the Platycrinitidae is presented.With consideration of the type species,Platycrinites laevisMiller, 1821,Platycrinitessensu stricto is distinguished fromPlatycrinitessensu lato, which is used for species that cannot be assigned with confidence to any objectively defined genus. New genera areArtaocrinusn. gen.,Collicrinusn. gen.,Elegantocrinusn. gen., andLaticrinusn. gen.; andExsulacrinusBowsher and Strimple, 1986 is designated a junior synonym ofPlatycrinitess.s.Collicrinus shumardin. gen. and sp.,Laticrinus owenin. gen. and sp., andLaticrinus wachsmuthin. gen. and sp. are described; andPlatycrinites formosus approximatus(Miller and Gurley, 1896a) is designated a junior synonym ofPlatycrinites formosus(Miller and Gurley, 1895a), which is reassigned here toCollicrinusn. gen.Platycrinitess.s. now includes 14 species and species-level taxa, and 76 species are assigned toPlatycrinitess.l. Ten species are designated nomina dubia, as are taxa based solely on columnals or pluricolumnals. Two species are designated nomina nuda, and two are transferred to genera outside of the Platycrinitidae. In addition, twenty-seven species and four open-nomenclature taxa are each reassigned to a different genus.

Early Triassic (Spathian) post-extinction microconchids from western Pangea

2013 ◽  
Vol 87 (1) ◽  
pp. 159-165 ◽  
Author(s):  
Michał Zatoń ◽  
Paul D. Taylor ◽  
Olev Vinn
Keyword(s):  
Lower Triassic ◽  
Late Triassic ◽  
Early Triassic ◽  
Mass Extinctions ◽  
Shell Microstructure ◽  
Slow Recovery ◽  
Dominant Component ◽  
Beaver Dam ◽  
Low Diversity ◽  
Water Facies

A new microconchid tentaculitoid,Microconchus utahensisnew species, is described from the Lower Triassic (Spathian) Virgin Formation of two localities (Hurricane Cliffs and Beaver Dam Mountains) near St George, Utah. This small encrusting tubeworm, previously referred to erroneously asSpirorbis, has a laminated shell microstructure containing minute pores (punctae). The population from deeper water facies of the Beaver Dam Mountains is more abundant than that from Hurricane Cliffs and the tubes are significantly larger in size. Although represented by only one species (M. utahensis), microconchids are by far the most dominant component of the otherwise impoverished sclerobiont assemblage of the Virgin Formation, which also includes rare cemented bivalves and probable foraminifers. Whereas the remainder of the Virgin fauna is quite diverse, the low diversity of encrusters suggests a slow recovery from end-Permian mass extinctions. Indeed, more typically Mesozoic sclerobiont assemblages dominated by cyclostome bryozoans and serpulid polychaetes did not appear until the Late Triassic, probably Rhaetian.

Sign in / Sign up

Export Citation Format

Share Document