<p>The physical and mechanical processes rooted in the hydrated, serpentinized mantle above subduction zones (the &#8220;cold nose&#8221;) remain debated and poorly understood, despite fundamental consequences on the elastic loading of the seismogenic interface. The fluids crossing this interface are expected to generate nests of seismicity and at the same time weaken the interface hanging wall through serpentinization and metasomatic processes. Ultramafic and jadeitite samples from two natural laboratories where such fossil settings are now visible at the Earth&#8217;s surface are used here to document multi-scale deformation mechanisms and fluid-rock interaction processes. Field relationships enable tracking the pathways followed by the fluids during HP metamorphism. Petrographic, geochemical, geochronological and microstructural observations demonstrate the complex interplay between brittle and plastic deformation processes throughout the gradual hydration of the cold nose mantle over millions of years. Changes in bulk rock geochemical and paragenetic sequence also reveal the evolution of the composition of the fluid source through time. These results shed light on the geometry of the cold nose above the interface, with implications for volatile budget estimates, rheology of the plate interface (including the various types of seismicity) as well as the interpretation of Vp/Vs ratios from active subduction settings worldwide.</p>