Coastal dunes occur along the majority of coastlines worldwide, often as parts of barrier islands, and are one of the most dynamic areas on earth. Sufficient supply of sand and strong onshore winds create a dynamic topography of building dunes near the shoreline and often a system of older dunes and troughs further inland. Coastal dunes have significant economic value for, among other things, tourism and fishing, as they protect inland areas from storms and high-water events. They also have critical ecological value, as they provide habitat for many animals, especially nesting seabirds, and shelter- sensitive oyster and seagrass beds in bays, as well as wetlands, and marshes from intense storms. Some of the earliest studies of plant succession, such as Cowles work in 1899, “The ecological relations of the vegetation on the sand dunes of Lake Michigan,” were conducted on coastal dunes and remain prominent for environmental research—especially as sea levels rise and human-related disturbances increase.