A late Quaternary vegetation history from Lily Lake, Chilkat Peninsula, southeast Alaska

1990 ◽  
Vol 68 (5) ◽  
pp. 1106-1112 ◽  
Author(s):  
Les C. Cwynar

Pollen and plant macrofossil analyses of a radiocarbon-dated core from Lily Lake on the Chilkat Peninsula, southeast Alaska, provide evidence for the history of the Pacific Coastal Forest. A Pinus contorta woodland with an abundance of herbs and ferns initially became established after deglaciation (10 870 years BP). By 10 330 years BP, Alnus viridis had become a significant component of the pine woodland while herbs and other shrubs declined. At 9480 years BP, Picea sitchensis and Populus were locally present and P. contorta populations declined. Tsuga heterophylla, which dominates the modern vegetation, became locally established at 7880 years BP. The last major component, Tsuga mertensiana, joined the vegetation about 6710 years BP. The modern closed forest, dominated by T. heterophylla and P. sitchensis, became established about 2870 years BP. The arrival times for the dominant species are compatible with dates from other studies that suggest a northward migration along the coast. Key words: fossil pollen, Alaska, Quaternary, plant migration, coastal forest.

2012 ◽  
Vol 78 (3) ◽  
pp. 572-582 ◽  
Author(s):  
Terri Lacourse ◽  
J. Michelle Delepine ◽  
Elizabeth H. Hoffman ◽  
Rolf W. Mathewes

AbstractPollen and conifer stomata analyses of lake sediments from Hippa Island on the north coast of British Columbia were used to reconstruct the vegetation history of this small hypermaritime island. Between 14,000 and 13,230 cal yr BP, the island supported diverse herb–shrub communities dominated by Cyperaceae, Artemisia and Salix. Pinus contorta and Picea sitchensis stomata indicate that these conifers were present among the herb–shrub communities, likely as scattered individuals. Transition to open P. contorta woodland by 13,000 cal yr BP was followed by increases in Alnus viridis, Alnus rubra and P. sitchensis. After 12,000 cal yr BP, Pinus-dominated communities were replaced by dense P. sitchensis and Tsuga heterophylla forest with Lysichiton americanus and fern understory. Thuja plicata stomata indicate that this species was present by 8700 cal yr BP, but the pollen record suggests that its populations did not expand to dominate regional rainforests, along with Tsuga and Picea, until after 6600 cal yr BP. Conifer stomata indicate that species may be locally present for hundreds to thousands of years before pollen exceed thresholds routinely used to infer local species arrival. When combined, pollen and conifer stomata can provide a more accurate record of paleovegetation than either when used alone.


2010 ◽  
Vol 73 (2) ◽  
pp. 259-268 ◽  
Author(s):  
Thomas A. Ager ◽  
Paul E. Carrara ◽  
Jane L. Smith ◽  
Victoria Anne ◽  
Joni Johnson

An AMS radiocarbon-dated pollen record from a peat deposit on Mitkof Island, southeastern Alaska provides a vegetation history spanning ∼12,900 cal yr BP to the present. Late Wisconsin glaciers covered the entire island; deglaciation occurred > 15,400 cal yr BP. The earliest known vegetation to develop on the island (∼12,900 cal yr BP) was pine woodland (Pinus contorta) with alder (Alnus), sedges (Cyperaceae) and ferns (Polypodiaceae type). By ∼12,240 cal yr BP, Sitka spruce (Picea sitchensis) began to colonize the island while pine woodland declined. By ∼11,200 cal yr BP, mountain hemlock (Tsuga mertensiana) began to spread across the island. Sitka spruce-mountain hemlock forests dominated the lowland landscapes of the island until ∼10,180 cal yr BP, when western hemlock (Tsuga heterophylla) began to colonize, and soon became the dominant tree species. Rising percentages of pine, sedge, and sphagnum after ∼7100 cal yr BP may reflect an expansion of peat bog habitats as regional climate began to shift to cooler, wetter conditions. A decline in alders at that time suggests that coastal forests had spread into the island's uplands, replacing large areas of alder thickets. Cedars (Chamaecyparis nootkatensis, Thuja plicata) appeared on Mitkof Island during the late Holocene.


2001 ◽  
Vol 79 (4) ◽  
pp. 389-397 ◽  
Author(s):  
Hugh J Barclay

Leaf angle distributions are important in assessing both the flexibility of a plant's response to differing daily and seasonal sun angles and also the variability in the proportion of total leaf area visible in remotely sensed images. Leaf angle distributions are presented for six conifer species, Abies grandis (Dougl. ex D. Don) Lindl., Thuja plicata Donn. ex D. Don, Tsuga heterophylla (Raf.) Sarg., Pseudotsuga menziesii (Mirb.) Franco, Picea sitchensis (Bong.) Carr. and Pinus contorta Dougl. ex Loud. var. latifolia. The leaf angles were calculated by measuring four foliar quantities, and then the distributions of leaf angles are cast in three forms: distributions of (i) the angle of the long axis of the leaf from the vertical for the range 0–180°; (ii) the angle of the long axis of the leaf for the range 0–90°; and (iii) the angle of the plane of the leaf for the range 0–90°. Each of these are fit to the ellipsoidal distribution to test the hypothesis that leaf angles in conifers are sufficiently random to fit the ellipsoidal distribution. The fit was generally better for planar angles and for longitudinal angles between 0° and 90° than for longitudinal angles between 0° and 180°. The fit was also better for Tsuga heterophylla, Pseudotsuga menziesii, Picea sitchensis, and Pinus contorta than for Abies grandis and Thuja plicata. This is probably because Abies and Thuja are more shade tolerant than the other species, and so the leaves in Abies and Thuja are preferentially oriented near the horizontal and are much less random than for the other species. Comparisons of distributions on individual twigs, whole branches, entire trees, and groups of trees were done to test the hypothesis that angle distributions will depend on scale, and these comparisons indicated that the apparent randomness and goodness-of-fit increased on passing to each larger unit (twigs up to groups of trees).Key words: conifer, leaf angles, ellipsoidal distribution.


1997 ◽  
Vol 12 (4) ◽  
pp. 115-121 ◽  
Author(s):  
Andris Eglitis ◽  
Paul E. Hennon

Abstract This study describes feeding damage by porcupines (Erethizon dorsatum) in precommercially thinned young growth stands of Sitka spruce (Picea sitchensis) and western hemlock (Tsuga heterophylla) on Mitkof Island in central southeast Alaska. We examined 641 trees from 54 sampling plots along transect lines in three 12 to 20 yr old stands. Porcupine feeding was monitored each spring and fall from 1985 to 1987. Four categories of feeding damage are described: complete girdling of the bole, partial girdling (bole scars), branch clipping, and "tasting wounds" (small basal bole scars). Sitka spruce, the primary crop tree in these thinned stands, sustained significantly higher damage (52% of trees affected) than western hemlock (26% of trees affected). Porcupine feeding was greater on taller than shorter Sitka spruce. Although only 8 of 59 trees initially girdled in 1985 were killed, many later sustained additional feeding damage. Following the 1987 season 3 yr after thinning, nearly 30% of the spruce and 14% of the western hemlock crop trees had been partially or completely girdled. Issues deserving future attention include the role of thinning in predisposing stands to porcupine damage, methods of population assessment, and mechanisms of host selection by porcupines. West. J. Appl. For. 12(4):115-121.


1977 ◽  
Vol 8 (3) ◽  
pp. 282-306 ◽  
Author(s):  
Calvin J. Heusser

Quaternary deposits on the Pacific slope of Washington range in age from the earliest known interglaciation, the Alderton, through the Holocene. Pollen stratigraphy of these deposits is represented by 12 major pollen zones and is ostensibly continuous through Zone 8 over more than 47,000 radiocarbon yr. Before this, the stratigraphy is discontinuous and the chronology less certain. Environments over the time span of the deposits are reconstructed by the comparison of fossil and modern pollen assemblages and the use of relevant meteorological data. The Alderton Interglaciation is characterized by forests of Douglas fir (Pseudotsuga menziesii), alder (Alnus), and fir (Abies). During the next younger interglaciation, the Puyallup, forests were mostly of pine, apparently lodgepole (Pinus contorta), except midway in the interval when fir, western hemlock (Tsuga heterophylla), and Douglas fir temporarily replaced much of the pine. Vegetation outside the limits of Salmon Springs ice (>47,00034,000 yr BP) varied chiefly between park tundra and forests of western hemlock, spruce (Picea), and pine. The Salmon Springs nonglacial interval at the type locality records early park tundra followed by forests of pine and of fir. During the Olympia Interglaciation (34,000–28,000 yr BP), pine invaded the Puget Lowland, whereas western hemlock and spruce became manifest on the Olympic Peninsula. Park tundra was widespread during the Fraser Glaciation (28,000–10,000 yr BP) with pine becoming more important from about 15,000 to 10,000 yr BP. Holocene vegetation consisted first of open communities of Douglas fir and alder; later, closed forests succeeded, formed principally of western hemlock on the Olympic Peninsula and of western hemlock and Douglas fir in the Puget Lowland. Over the length of the reconstructed environmental record, climate shifted between cool and humid or relatively warm, semihumid forest types and cold, relatively dry tundra or park tundra types. During times of glaciation, average July temperatures are estimated to have been at least 7°C lower than today. Only during the Alderton Interglaciation and during the Holocene were temperatures higher for protracted periods than at present.


1978 ◽  
Vol 56 (13) ◽  
pp. 1510-1517 ◽  
Author(s):  
Calvin J. Heusser

Modern pollen rain was studied at 98 sites distributed in a belt transect some 250 km wide running a distance of 425 km between La Push on the Pacific coast and Grand Coulee on the Columbia Plateau. Sites are located in a variety of plant communities from the Picea sitchensis Zone near sea level to the Alpine Tundra Zone at 2285 m in the Cascades. The purposes of the study are (1) to establish the nature of the pollen rain and its relation to vegetation sources and (2) to develop further the pool of potential analogs of fossil pollen spectra for use in the reconstruction of Quaternary environments.Relative frequencies are shown for the pollen of Pinus, Picea, Abies, Tsuga, Pseudotsuga, Cupressaceae, Alnus, Gramineae, Cyperaceae, Chenopodiaceae, Artemisia, and Compositae. Results indicate the pollen rain to be related to source vegetation at the collection sites except for certain instances of infiltration by extraneous types, of which Pinus is most conspicuous. Alnus is the dominant pollen in the Puget Lowland and the western Coast and Cascade Ranges; Tsuga heterophylla is most prominent in the Coast Range; and Pinus ranks foremost in the eastern Cascades and on the Columbia Plateau. On the Columbia Plateau, the Gramineae, Artemisia, Compositae, and Chenopodiaceae are also distinctive. Subalpine forest types and nonarboreals are generally characteristic of the Timberline and Alpine Tundra Zones.


1996 ◽  
Vol 46 (2) ◽  
pp. 161-175 ◽  
Author(s):  
Barbara C. S. Hansen ◽  
Daniel R. Engstrom

A 13,000-year history of late-Quaternary vegetational and climatic change has been derived from lacustrine sediments from Pleasant Island in the Glacier Bay region of southeastern Alaska. Early arrival of lodgepole pine and mountain hemlock, indicated by the presence of pollen and conifer stomata, suggests expansion from refugia in the Alexander Archipelago. A short-term climatic reversal, possibly correlated with the European Younger Dryas, is inferred from the expansion of tundra elements and deposition of inorganic sediments between 10,600 and 9900 14C yr B.P. Two peat cores from the lake catchment verify Holocene vegetation changes and aid in the separation of biogenic from climatic forces affecting vegetation history. Differences in pollen representation among the three cores illustrate the variation among pollen-collecting substrates, as well as the spatial heterogeneity of peatland development and its dependence on local hydrology. Initial peat accumulation and soil paludification, occasioned by increases in temperature and precipitation in the early Holocene, allowed western and mountain hemlock to replace sitka spruce 8500–8000 yr B. P. Open muskeg became widespread about 7000 yr B. P. and allowed lodgepole pine to reinvade the region after a 2000-yr absence. The extensive replacement of fen elements by bog taxa at 3400 yr B. P. suggests increased paludification due to changing hydrologic conditions; its correlation with the upland expansion of Tsuga heterophylla suggests the onset of a cooler/wetter Neoglacial climate in southeastern Alaska.


2002 ◽  
Vol 58 (3) ◽  
pp. 215-225 ◽  
Author(s):  
Colin J. Long ◽  
Cathy Whitlock

AbstractHigh-resolution charcoal and pollen analyses were used to reconstruct a 4600-yr-long history of fire and vegetation near Taylor Lake in the wettest forests of coastal Oregon. Today, fires in these forests are rare because the season of ignition does not coincide with months of dry fuels. From ca. 4600 to 2700 cal yr B.P. fire episodes occurred at intervals of 140±30 yr while forest vegetation was dominated by disturbance-adapted taxa such as Alnus rubra. From ca. 2700 cal yr B.P. to the present, fire episodes have become less common, occurring at intervals of 240±30 yr, and fire-sensitive forest taxa, such as Tsuga heterophylla and Picea sitchensis, have become more prominent. Fire occurrence during the mid-Holocene was similar to that of the more xeric forests in the eastern Coast Range and suggests that summer drought was widespread. After ca. 2700 cal yr B.P., a decrease in fire episode frequency suggests that cooler conditions and possibly increased summer fog allowed the establishment of present-day Picea sitchensis forests within the watershed. These results provide evidence that fire has been an important disturbance agent in the Coast Range of Oregon, and variations in fire frequency and climate have led to the establishment of present-day forests.


1986 ◽  
Vol 25 (1) ◽  
pp. 100-120 ◽  
Author(s):  
Dorothy M. Peteet

Seventy surface pollen samples from coastal forest, coastal meadow, muskeg, tree line, and alpine tundra communities form a basis for interpreting fossil pollen assemblages in the Malaspina Glacier district, Alaska. Poflen and macrofossil analyses of three radiocarbon-dated fossil sections from Icy Cape indicate that vegetational changes resulting from plant succession can be distinguished from those of migrational and climatic origin. Vegetation of the early Holocene xerothermic interval (10,000–7600 yr B.P.) was dominated by Alnus communities. Wetter conditions ensued, enabling generative muskeg surfaces to develop and first Picea sitchensis, then Tsuga heterophylla to expand from areas southeastward. Climatic cooling in more recent millennia (3500 yr B.P. to the present) is indicated by the appearance and persistent growth of Tsuga mertensiana and Selaginella selaginoides along this portion of the Gulf of Alaska coastline.


1998 ◽  
Vol 28 (10) ◽  
pp. 1501-1508 ◽  
Author(s):  
L M Trummer ◽  
P E Hennon ◽  
E M Hansen ◽  
P S Muir

Amodel was developed to predict the severity of dwarf mistletoe (Arceuthobium tsugense (Rosendahl) G.N. Jones) in western hemlock trees (Tsuga heterophylla (Raf.) Sarg.) that developed within forests of Southeast Alaska that experienced near-catastrophic windthrow in the late 1800s. The model suggests that the degree of dwarf mistletoe severity on western hemlock trees was significantly and positively correlated with levels of dwarf mistletoe infection and basal area (m2/ha) in large and small residual trees that survived the wind disturbance. No significant relationships were found between severity level and any other factors, including site productivity, density of coexisting Sitka spruce (Picea sitchensis (Bong.) Carr.), or slope. The model demonstrates the overriding importance of infected residual trees to predict future severity of dwarf mistletoe; greater size and infection level of residual trees results in greater dwarf mistletoe levels on regenerating hemlock crop trees. The model, derived from 76 plots on Kuiu Island, was tested in 18 plots on Chichagof Island, providing a preliminary validation. Slower rates of dwarf mistletoe spread and intensification in forests of southeastern Alaska, as compared with similar coastal forests south of Alaska, provide an opportunity for managers to manipulate the parasite to desired levels in managed forests.


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