scholarly journals Speciation and infrageneric classification in the planktonic dinoflagellate Tripos (Gonyaulacales, Dinophyceae)

2020 ◽  
Author(s):  
Fernando Gomez
Keyword(s):  
Life Cycle ◽  
Intraspecific Variability ◽  
Taxonomic Revision ◽  
Marine Species ◽  
Molecular Data ◽  
Infrageneric Classification ◽  
Numerous Species ◽  
The Past ◽  
Infraspecific Taxa

The genus Tripos , formerly marine species of Ceratium , is the dinoflagellate with the greatest number of species and infraspecific taxa (~800) due to the high morphological intraspecific variability of numerous species. In the past, the species of Tripos were proposed into distinct genera. A re-instatement of this generic split is not justified due the difficulties to circumscribe the basal subgenera Amphiceratium and Archaeceratium , and the polyphyletic character of Biceratium . A new infrageneric classification is proposed. The subgenus Amphiceratium is dismembered after the classification of the sections Inflata and partially Fusiformia into Archaeceratium . The subgenus Tripos (autonym) replaces other names such as Tripoceratium or Orthoceratium . Based on the original descriptions, the records of T. furca and T. lineatus correspond to T. eugrammus and T. furca , respectively, and T. macroceros has been reported as T. contrarius . The names T. belone and T. carriensis have been misapplied for T. pacificus and T. volans , respectively. Tripos arcuatus , T. gracilis , T. inclinatus , T. scapiformis and T. subcontortus are revived to replace T. euarcuatus , T. declinatus , T. horridus , T. longirostrum and T. contortus , respectively. The species T. ramakrishnae and T. fusus var. schuettii were described from individuals infected by endoparasites. Tripos rotundatus comb. nov . is proposed for C. digitatum var. rotundatum. As a result of taxonomic revision, Tripos is restricted to 57 correct species, although the speciation and synonymy is largely incomplete due to lack of studies in the life cycle and molecular data.

Speciation and Infrageneric Classification in the Planktonic Dinoflagellate Tripos (Gonyaulacales, Dinophyceae)

2021 ◽  
Vol 01 ◽  
Author(s):  
Fernando Gómez
Keyword(s):  
Life Cycle ◽  
Mediterranean Sea ◽  
Intraspecific Variability ◽  
Marine Species ◽  
Molecular Data ◽  
Infrageneric Classification ◽  
Numerous Species ◽  
The Past ◽  
Infraspecific Taxa

Background: The genus Tripos, formerly marine species of Ceratium, is the dinoflagellate with the greatest number of species and infraspecific taxa (~800) due to the high morphological intraspecific variability of numerous species. In the past, the species of Tripos were proposed into distinct genera. Objective: To propose an infrageneric classification and to review the taxonomy and nomenclature of each taxon in order to establish the correct species and synonymy. Method: Observations from the Mediterranean Sea, and Atlantic and Pacific Oceans, a review of the original descriptions and further literature, and the available molecular data. Results: re-instatement of this generic split is not justified due the difficulties to circumscribe the basal subgenera Amphiceratium and Archaeceratium, and the polyphyletic character of Biceratium. The subgenus Amphiceratium is dismembered after the classification of the sections Inflata and partially Fusiformia into Archaeceratium. The subgenus Tripos (autonym) replaces other names such as Tripoceratium or Orthoceratium. Based on the original descriptions, the records of T. furca and T. lineatus correspond to T. eugrammus and T. furca, respectively, and T. macroceros has been reported as T. contrarius. The names T. belone and T. carriensis have been misapplied for T. pacificus and T. volans, respectively. Tripos arcuatus, T. gracilis, T. inclinatus, T. scapiformis and T. subcontortus are revived to replace T. euarcuatus, T. declinatus, T. horridus, T. longirostrum and T. contortus, respectively. The species T. ramakrishnae and T. fusus var. schuettii were described from individuals infected by endoparasites. Tripos rotundatus comb. nov. is proposed for C. digitatum var. rotundatum. Conclusion: Tripos is restricted to 57 correct species, although the speciation and synonymy is largely incomplete due to lack of studies in the life cycle and molecular data.

A first step towards the revision of Cicurina: redescription of type specimens of 60 troglobitic species of the subgenus Cicurella (Araneae: Dictynidae), and a first visual assessment of their distribution

Zootaxa ◽  
2009 ◽  
Vol 2002 (1) ◽  
pp. 1-67 ◽  
Author(s):  
PIERRE PAQUIN ◽  
NADINE DUPÉRRÉ
Keyword(s):  
Conservation Status ◽  
Intraspecific Variability ◽  
Taxonomic Revision ◽  
Visual Assessment ◽  
Nominal Species ◽  
Type Specimens ◽  
Numerous Species ◽  
Morphological Differences ◽  
The U.S ◽  
Conservation And Management

The intraspecific variability recently documented in the genus Cicurina strongly suggests a reassessment of the taxonomy, particularly for the troglobitic members. Adult troglobitic Cicurina (subgenus Cicurella) are rare and most of the 60 nominal species of troglobitic Cicurina (Cicurella) were originally described upon the examination of only one or two females, resulting in numerous species differing only in minor variations of the female genitalia. In many cases, such morphological differences could also be interpreted as intraspecific variability. We present the first step of the taxonomic revision of the genus with the redescription of all troglobitic species of the subgenus Cicurella based on type specimens to provide a reliable and comparable morphological basis: Cicurina bandera Gertsch 1992, C. bandida Gertsch 1992, C. baronia Gertsch 1992, C. barri Gertsch 1992, C. browni Gertsch 1992, C. brunsi Cokendolpher 2004, C. bullis Cokendolpher 2004, C. buwata Chamberlin & Ivie 1940, C. caliga Cokendolpher & Reddell 2001, C. caverna Gertsch 1992, C. coahuila Gertsch 1971, C. coryelli Gertsch 1992, C. cueva Gertsch 1992, C. delrio Gertsch 1992, C. ezelliGertsch 1992, C. gruta Gertsch 1992, C. holsingeri Gertsch 1992, C. hoodensis Cokendolpher & Reddell 2001, C. leona Gertsch 1992, C. loftini Cokendolpher 2004, C. machete Gertsch 1992, C. madla Gertsch 1992, C. maya Gertsch 1977, C. mckenziei Gertsch 1992, C. medina Gertsch 1992, C. menardia Gertsch 1992, C. mirifica Gertsch 1992, C. mixmaster Cokendolpher & Reddell 2001, C. neovespera Cokendolpher 2004, C. obscura Gertsch 1992, C. orellia Gertsch 1992, C. pablo Gertsch 1992, C. pastura Gertsch 1992, C. patei Gertsch 1992, C. platypus Cokendolpher 2004, C. porteri Gertsch 1992, C. puentecilla Gertsch 1992, C. rainesi Gertsch 1992, C. reclusa Gertsch 1992, C. reddelli Gertsch 1992, C. reyesi Gertsch 1992, C. russelli Gertsch 1992, C. sansaba Gertsch 1992, C. selecta Gertsch 1992, C. serena Gertsch 1992, C. sheari Gertsch 1992, C. sprousei Gertsch 1992, C. stowersi Gertsch 1992, C. suttoni Gertsch 1992, C. travisae Gertsch 1992, C. troglobia Cokendolpher 2004, C. ubicki Gertsch 1992, C. uvalde Gertsch 1992, C. venefica Gertsch 1992, C. venii Gertsch 1992, C. vespera Gertsch 1992, C. vibora Gertsch 1992, C. wartoni Gertsch 1992, C. watersi Gertsch 1992 and C. wiltoni Gertsch 1992. We provide, in a series of maps, a first visual assessment of the distribution of these troglobites. Several problems became evident from our examination of the data, including potential synonymies, conflicting distributions, damaged types, potential mislabeling, species known only from one sex, misevaluation of intraspecific variability, and unknown location of type localities. A reliable taxonomic basis for the eyeless members of this genus is particularly important for cave conservation and management because the particular conservation status of these troglobites, particularly for four species that are included on the U.S. Federal list of endangered species.

scholarly journals Taxonomic problems in the subgenus Meloehelea Wirth of the genus Atrichopogon Kieffer (Diptera: Ceratopogonidae) inferred from both morphological and molecular characters

2008 ◽  
Vol 19 (1) ◽  
pp. 1-12 ◽  
Author(s):  
Andrea Tóthová ◽  
Jan Knoz ◽  
Radim Sonnek ◽  
Josef Bryja ◽  
Jaromír Vaňhara
Keyword(s):  
Morphological Variability ◽  
Intraspecific Variability ◽  
Taxonomic Revision ◽  
Morphological Characters ◽  
Valid Species ◽  
European Species ◽  
Rdna Sequences ◽  
16S Rdna Sequences ◽  
Molecular Characters

The classification of Meloehelea (Ceratopogonidae) species is based on morphological characters. The taxonomic revision of generally well-known species recovered interesting facts on possible geographic intraspecific variability or on the presence of an additional valid species. In this study, 5 European species and 2 species from the U.S.A. and Canada have been compared based on their morphological characters. Subsequently, the 16S rDNA sequences analyses of well known European species have been performed to confirm the characters mentioned in the determination key ofthis subgenus. Although, this study was limited by the number of examined non-European specimens, it provides some interesting facts on the possible morphological variability of well-known species and also an initial phylogenetic backbone for the progressive reconstruction of infrageneric relationships within the genus Atrichopogon.

scholarly journals Taxonomic Revision and Classification of Extant Holococcolithophores Previously Placed in the Genus Anthosphaera Kamptner emend. Kleijne 1991

2021 ◽  
Vol 59 (3-4) ◽  
pp. 121-139
Author(s):  
Odysseas A. Archontikis ◽  
Jeremy R. Young ◽  
Lluïsa Cros
Keyword(s):  
Life Cycle ◽  
Type Species ◽  
Taxonomic Revision ◽  
Species Level ◽  
New Combinations ◽  
Distinctive Group

The genus Anthosphaera Kamptner emend. Kleijne is one of the most taxonomically confusing modern coccolithophores and its species level taxonomy has long been in a state of flux. Based on the review of imaged specimens from our collections, we attempt to rectify the nomenclatural problems and elucidate the obfuscated taxonomy of the genus. Review of included formally and informally described species shows that they are a distinctive group with shared characters, including ten different morphotypes of probable species level. Two of these, including the type species A. fragaria, have been shown to form life-cycle associations with heterococcoliths of the Syracosphaera molischii type. Hence, all species are transferred to Syracosphaera and the new combinations S. periperforata, S. lafourcadii, and S. origami are proposed. In addition, various informally described morphotypes are now formally described as Syracosphaera molischii var. pertusa, S. periperforata var. cylindrata, S. periperforata var. tridentata, S. rotaconica, and S. elevata. urn:lsid:zoobank.org:pub:0E5D4BD7-BC3B-4D30-B319-964AC887DDDE

Phylogeny, major clades and infrageneric classification of Corymbia (Myrtaceae), based on nuclear ribosomal DNA and morphology

2009 ◽  
Vol 22 (5) ◽  
pp. 384 ◽  
Author(s):  
Carlos Parra-O. ◽  
Michael J. Bayly ◽  
Andrew Drinnan ◽  
Frank Udovicic ◽  
Pauline Ladiges
Keyword(s):  
Ribosomal Dna ◽  
Convergent Evolution ◽  
Morphological Characters ◽  
Molecular Data ◽  
Nuclear Ribosomal Dna ◽  
Molecular Evidence ◽  
Nuclear Rdna ◽  
Infrageneric Classification ◽  
Taxonomic Categories

Phylogenetic relationships of sections and species within Corymbia (Myrtaceae), the bloodwood eucalypts, were evaluated by using combined analyses of nuclear rDNA (ETS + ITS) and morphological characters. Combining morphological characters with molecular data provided resolution of relationships within Corymbia. The analyses supported the monophyly of the genus and recognition of the following two major clades, treated here as new subgenera: subgenus Corymbia, including informal sections recognised by Hill and Johnson (1995), namely Rufaria (red bloodwoods), Apteria and Fundoria; and subgenus Blakella, including sections Politaria (spotted gums), Cadagaria, Blakearia (paper-fruited bloodwoods or ghost gums) and Ochraria (yellow bloodwoods). Hill and Johnson’s section Rufaria is monophyletic if Apteria and Fundoria are included. It is evident that, among the red bloodwoods, series are not monophyletic and several morphological characters result from convergent evolution. There was strong morphological and molecular evidence that the three species of red bloodwoods that occur in south-western Western Australia (series Gummiferae: C. calophylla and C. haematoxylon, and series Ficifoliae: C. ficifolia) form a monophyletic group, separate from the eastern C. gummifera (series Gummiferae), which is probably sister to the clade of all other red bloodwoods. Phylogenetic results supported recognition of new taxonomic categories within Corymbia, and these are formalised here.

Corrigendum to: Phylogeny, major clades and infrageneric classification of Corymbia (Myrtaceae), based on nuclear ribosomal DNA and morphology

2010 ◽  
Vol 23 (2) ◽  
Author(s):  
Carlos Parra-O. ◽  
Michael J. Bayly ◽  
Andrew Drinnan ◽  
Frank Udovicic ◽  
Pauline Ladiges
Keyword(s):  
Ribosomal Dna ◽  
Convergent Evolution ◽  
Morphological Characters ◽  
Molecular Data ◽  
Nuclear Ribosomal Dna ◽  
Molecular Evidence ◽  
Nuclear Rdna ◽  
Infrageneric Classification ◽  
Taxonomic Categories

Phylogenetic relationships of sections and species within Corymbia (Myrtaceae), the bloodwood eucalypts, were evaluated by using combined analyses of nuclear rDNA (ETS + ITS) and morphological characters. Combining morphological characters with molecular data provided resolution of relationships within Corymbia. The analyses supported the monophyly of the genus and recognition of the following two major clades, treated here as new subgenera: subgenus Corymbia, including informal sections recognised by Hill and Johnson (1995), namely Rufaria (red bloodwoods), Apteria and Fundoria; and subgenus Blakella, including sections Politaria (spotted gums), Cadagaria, Blakearia (paper-fruited bloodwoods or ghost gums) and Ochraria (yellow bloodwoods). Hill and Johnson's section Rufaria is monophyletic if Apteria and Fundoria are included. It is evident that, among the red bloodwoods, series are not monophyletic and several morphological characters result from convergent evolution. There was strong morphological and molecular evidence that the three species of red bloodwoods that occur in south-western Western Australia (series Gummiferae: C. calophylla and C. haematoxylon, and series Ficifoliae: C. ficifolia) form a monophyletic group, separate from the eastern C. gummifera (series Gummiferae), which is probably sister to the clade of all other red bloodwoods. Phylogenetic results supported recognition of new taxonomic categories within Corymbia, and these are formalised here.

Comparative leaf micromorphology of Drypetes and Putranjiva (Putranjivaceae) and its taxonomic significance

2020 ◽  
Author(s):  
Akeem B Kadiri ◽  
Alexandra N Muellner-Riehl
Keyword(s):  
Molecular Data ◽  
Leaf Epidermis ◽  
Infrageneric Classification ◽  
Leaf Micromorphology ◽  
Stomatal Complexes ◽  
Branching Patterns ◽  
Doubtful Species ◽  
Epidermal Features ◽  
Micromorphological Characters

Abstract Putranjivaceae are a pantropically distributed but poorly known glucosinulate-producing family of three genera (Drypetes, Putranjiva and Sibangea), previously included in Euphorbiaceae subfamily Phyllanthoideae. Characters of the leaf epidermis were previously suggested as being of infrafamilial taxonomic relevance, but epidermal features of the three genera have not so far been compared in the light of the phylogenetic relationships recovered by molecular data. Therefore, we compared the leaf micromorphology of 44 (out of c. 200) Drypetes spp. and two (out of three) Putranjiva spp. using light microscopy. Our findings revealed that there are more stomata types than previously reported in both genera. We also present a new record of stomatal complexes and epidermal cell characteristics. The cell wall ornamentation character on both surfaces of the epidermis is useful for distinguishing D. aframensis subsp. aframensis. Based on a combination of the studied characters, doubtful species of Drypetes and Putranjiva are recognizable as belonging to the two genera, and this will require taxonomic and nomenclatural adjustments. Other taxonomically useful characters are trichome distribution and types and leaf areole shape and tertiary veins branching patterns that appear to be relatively constant across the two genera. Oil droplets located within the cell lumen in Putranjiva differentiates it from Drypetes, where they inwardly streak the epidermal walls among other features. Thus, our study provides evidence that the investigated leaf micromorphological characters are useful for distinguishing the species of the two genera, and they corroborate the existing infrageneric classification of Drypetes that was based on exomorphology.

Centipede systematics: progress and problems*

Zootaxa ◽  
2007 ◽  
Vol 1668 (1) ◽  
pp. 327-341 ◽  
Author(s):  
GREGORY D. EDGECOMBE
Keyword(s):  
Species Delimitation ◽  
Middle Devonian ◽  
Phylogenetic Analyses ◽  
Molecular Data ◽  
Ribosomal Genes ◽  
Model Species ◽  
Crown Group ◽  
Web Based ◽  
The Past

Breakthroughs in centipede systematics over the past 25 years have included: a stable morphology-based cladogram for ordinal interrelationships that is largely congruent with well-sampled nuclear ribosomal genes; the discovery of mid Palaeozoic crown-group fossils, including Silurian-Devonian stem-group Scutigeromorpha and an extinct order in the Middle Devonian; and, a web-based catalogue of all centipede species globally. Challenges include species delimitation in several groups, conflict between different kinds of molecular data (nuclear coding genes versus ribosomal genes), the inter-familial relationships and classification of the Geophilomorpha in particular, and effecting a synthesis between microanatomical studies of selected ‘model’ species and dense taxonomic sampling in numerical phylogenetic analyses.

Resolving phylogenetic and classical nomenclature: A revision of Seraphsidae Jung, 1974 (Gastropoda: Neostromboidae)

Zootaxa ◽  
2021 ◽  
Vol 4990 (3) ◽  
pp. 401-453
Author(s):  
STEPHEN J. MAXWELL ◽  
TASMIN L. RYMER ◽  
BRADLEY C. CONGDON
Keyword(s):  
Taxonomic Revision ◽  
First Record ◽  
Molecular Data ◽  
The Family ◽  
Structural Differences ◽  
Integrated Taxonomy ◽  
Taxonomic Approach ◽  
Current Species ◽  
A New Species

This revision demonstrates that an integrated taxonomic approach to classical taxonomic practice can lead to increased internal cladistic resolution within a clade, including the recognition of new taxa at all nomenclatural levels. In particular, this revision has two aims: 1) to complete an α-taxonomic revision of Seraphsidae (Stromboidea); and 2) to resolve the infrafamilial relationships within Seraphsidae using morphological cladistics. An annotated synonymy was generated for each taxon, the precedence of names determined, and revised descriptions formulated. Character sets that reflect the synapomorphies within the major subclades of Seraphsidae were coded and a cladogram generated using maximum likelihood within the tnt program with default settings. Four forms of material and evidence were used in this systematic review: 1) the type material for each taxon; 2) non-type physical material; 3) published images; and 4) literary references for specimen localities without illustration. The current morphologically-based classification of Seraphsidae was found to be sound in terms of current species delimitations. Regardless of this, the use of an integrated taxonomy improved understanding of the internal cladistic relationships within Seraphsidae, which led to higher resolution of the internal cladistic arrangements and taxonomic delimitation. Furthermore, this increased resolution is now reflected in the family nomenclature. Two new subfamilies are proposed, Seraphsinae and Pseudoterebellinae, which are used to distinguish the two clades within Seraphsidae based on form: members of Seraphsinae are involute, while members of Pseudoterebellinae are evolute. Pseudoterebellum is proposed as a new genus to show the discontinuity in the fossil record and highlight the structural differences between it and Terebellum. A new species from Jamaica, Seraphs kaindraperi, is described, being the first record from that location and that geological period and is morphologically distinct from other described taxa. This revision included Mauryna within Seraphsidae on the grounds that it provides a basal link to the Seraphsidae sister taxa Semiterebellum and Terebellopsis, all of which are currently contained within Rostellariidae, and all three clades may in time be brought together under one family. All clades have been constructed to be in compliance with both the ICZN and PhyloCode. It was not possible within the context of this revision to test the results of the α-taxonomic findings against phylogenies generated with molecular data. This was due to the high number of extinct taxa within the Seraphsidae.  

Phylogeny and taxonomic revision of Deronectina Galewski, 1994 (Coleoptera: Dytiscidae: Hydroporinae: Hydroporini)

Zootaxa ◽  
2018 ◽  
Vol 4474 (1) ◽  
pp. 1 ◽  
Author(s):  
HANS FERY ◽  
IGNACIO RIBERA
Keyword(s):  
Molecular Phylogeny ◽  
Type Species ◽  
Taxonomic Revision ◽  
Molecular Data ◽  
Neotropical Region ◽  
Mitochondrial Coi ◽  
The North ◽  
Current Classification ◽  
Taxonomic Changes

The subtribe Deronectina Galewski, 1994 (Dytiscidae, Hydroporinae, Hydroporini) is distributed in the Nearctic, in the north of the Neotropical region, and in the Palaearctic and Afrotropical regions. It is currently composed of 194 species and 13 subspecies in eight genera: Amurodytes Fery & Petrov, 2013, Boreonectes Angus, 2010, Deronectes Sharp, 1882, Nebrioporus Régimbart, 1906, Oreodytes Seidlitz, 1887, Scarodytes Gozis, 1914, Stictotarsus Zimmermann, 1919, and Trichonectes Guignot, 1941. We present a morphological and a molecular phylogeny of the species of the subtribe, and a revision of their taxonomy to accommodate our phylogenetic results. The morphological phylogeny is based on the study of 54 characters of the adults of 189 species and 2 subspecies, of which 114 species and the 2 subspecies were coded in the morphological matrix. For the molecular phylogeny we investigated 115 species and 11 subspecies, using a combination of fragments of four mitochondrial (COI, 16S rRNA, tRNA-Leu and NAD1) and two nuclear genes (18S rRNA and H3), analysed with maximum likelihood and Bayesian methods. For both datasets we included the type species of all genus-group taxa. The morphological, molecular and combined phylogenies mostly agree with the current classification of the group, but in some cases our results are in contradiction with established genera. Most remarkable are the polyphyly of Stictotarsus and Nebrioporus, the low support for the monophyly and internal phylogeny of Oreodytes, and the low support for the monophyly of Deronectina with molecular data. Thus, we introduce some taxonomic changes in the current classification to accommodate the generic concepts to our phylogenetic results. Nine new genera are established: Clarkhydrus n. gen. (type species Hydroporus roffii Clark, 1862), Hornectes n. gen. (type species Hydroporus quadrimaculatus Horn, 1883), Iberonectes n. gen. (type species Deronectes bertrandi Legros, 1956), Larsonectes n. gen. (type species Potamonectes minipi Larson, 1991), Leconectes n. gen. (type species Hydroporus striatellus LeConte, 1852), Mystonectes n. gen. (type species Deronectes neomexicanus Zimmerman & Smith, 1975), Nectoboreus n. gen. (type species Hydroporus aequinoctialis Clark, 1862), Nectomimus n. gen. (type species Oreodytes okulovi Lafer, 1988), and Zaitzevhydrus n. gen. (type species Hydroporus formaster Zaitzev, 1908). Three genera are reinstated as valid: Deuteronectes Guignot, 1945 (stat. rest.) (type species Hydroporus picturatus Horn, 1883), Nectoporus Guignot, 1950 (stat. rest.) (type species Hydroporus abbreviatus Fall, 1923), and Neonectes J. Balfour-Browne, 1940 (stat. rest.) (type species Hydroporus natrix Sharp, 1884). Thirty-six new combinations for species and subspecies thus far treated in the genera Boreonectes, Nebrioporus, Oreodytes and Stictotarsus result from the new classification: Clarkhydrus corvinus (Sharp, 1887) n. comb., C. decemsignatus (Clark, 1862) n. comb., C. deceptus (Fall, 1932) n. comb., C. eximius (Motschulsky, 1859) n. comb., C. falli (Nilsson, 2001) n. comb., C. interjectus (Sharp, 1882) n. comb., C. minax (Zimmerman, 1982) n. comb., C. opaculus (Sharp, 1882) n. comb., C. roffii (Clark, 1862) n. comb., C. spectabilis (Zimmerman, 1982) n. comb., Deuteronectes angustior (Hatch, 1928) n. comb., Hornectes quadrimaculatus (Horn, 1883) n. comb., Iberonectes bertrandi (Legros, 1956) n. comb., Larsonectes minipi (Larson, 1991) n. comb., Leconectes striatellus (LeConte, 1852) n. comb., Mystonectes coelamboides (Fall, 1923) n. comb., M. grammicus (Sharp, 1887) n. comb., M. neomexicanus (Zimmerman & Smith, 1975) n. comb., M. panaminti (Fall, 1923) n. comb., M. titulus (Leech, 1945) n. comb., Nectoboreus aequinoctialis (Clark, 1862) n. comb., N. dolerosus (Leech, 1945) n. comb., N. funereus (Crotch, 1873) n. comb., Nectomimus okulovi (Lafer, 1988) n. comb., Nectoporus angelinii (Fery, 2015) n. comb., N. congruus (LeConte, 1878) n. comb., N. crassulus (Fall, 1923) n. comb., N. obesus obesus (LeConte, 1866) n. comb., N. obesus cordillerensis (Larson, 1990) n. comb., N. rhyacophilus (Zimmerman, 1985) n. comb., N. sanmarkii sanmarkii (C.R. Sahlberg, 1826) n. comb., N. sanmarkii alienus (Sharp, 1873) n. comb., N. sierrae (Zimmerman, 1985) n. comb., N. subrotundus (Fall, 1923) n. comb., Zaitzevhydrus formaster formaster (Zaitzev, 1908) n. comb., and Z. formaster ulanulana (C.-K. Yang, 1996) n. comb. 

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