In both Europe and North America, modern herpetological families and genera became established quite early in the Cenozoic, and modern species occurred as early as the Miocene. Because of deteriorating climates that began late in the Eocene, a marked decrease in herpetological diversity occurred in the Oligocene in both continents. However, both areas became herpetologically enriched in the Miocene. In post-Miocene times Europe was isolated from Africa and warm areas in the east by the Mediterranean Sea and eastern mountain ranges, and a depauperate herpetofauna developed there that continued into recent times. In North America, however, with its vast, accessible southern land mass, the richness of the Miocene herpetofauna (with the exception of several archaic colubrid genera [Parmley and Holman, 1995] that became extinct in the the Miocene) persisted into modern times. The following discussion of changes in the European herpetofauna in the Cenozoic era has been synthesized from Auge (1986), Ballón (1991a), Bailon ct al. (1988), Barbadillo et al. (1997), Crochet et al. (1981), Estes (1981, 1982, 1983), Fritz (1995), Holman (1995c), Milncr (1986), Milner et al. (1982), Mlynarski (1976), Rage (1984a, 1984c, 1986, 1993), Rage and Auge (1993), Rage and Ford (1980), Roček (1994), Sanchiz. (1977b, in press), Sanchiz and Mlynarski (1979), Sanchiz and Roček (1996), Spinar (1972), Szyndlar (1984, 1991b, 1991c), and Szyndlar and Bohme (1993). Because of the high probability that herpetological fossils have been identified correctly at the family level, herpetological families arc used here to reflect the taxonomic diversity of the European herpetofauna from the Paleocene through the Pliocene. In a following section, the earliest appearance of herpetological genera and species in the European Tertiary arc discussed. Extinct families are prefixed with an asterisk (*). Families that became extinct in Europe in the Cenozoic but presently occur elsewhere are prefixed with a number sign (#). Two primitive, extinct, presumably permanently aquatic salamander families, me *Albanerpetontidae and *Batrachosauroididae (the latter also known from the Tertiary of North America) made limited appearances in the Cenozoic of Europe. The *Albanerpetontidae occurred only in the Middle Miocene (having reappeared from the Cretaceous), and the *Batrachosauroididae occurred from the Upper Paleocene to the Lower Eocene.
A fossil water measurer (Insecta, Hemiptera, Hydrometridae) from the Paleocene/Eocene of Denmark and its phylogenetic relationships
