Advanced cladid crinoids from the middle Mississippian of the east-central United States: advanced-grade calyces

1994 ◽  
Vol 68 (2) ◽  
pp. 339-351 ◽  
Author(s):  
Thomas W. Kammer ◽  
William I. Ausich

Species of the late Osagean and early Meramecian advanced cladid crinoid genera (suborder Poteriocrinina) Adinocrinus, Eratocrinus, Lekocrinus n. gen., Linocrinus, Sarocrinus, and Worthenocrinus n. gen. from Illinois, Indiana, Iowa, Kentucky, and Missouri are redescribed and redefined from study of type material and museum collections. These species are considered to have advanced-grade calyces because of the substantial size reduction of the infrabasals and basals relative to the radials. Nomenclatural and systematic changes include the following: 1) Eratocrinus coxanus (Worthen, 1882) is a senior synonym of Zeacrinus keokuk Worthen, 1882; 2) E. commaticus (Miller, 1891a) is a senior synonym of Z. blairi Miller and Gurley, 1895; 3) Lekocrinus n. gen. is described and Scaphiocrinus divaricatus Hall, 1860, and Z. planobrachiatus Meek and Worthen, 1860, are assigned to this new genus; 4) Linocrinus praemorsus (Miller and Gurley, 1890a) is a senior synonym of Poteriocrinus lautus Miller and Gurley, 1896; 5) Sarocrinus kirki n. sp. is described; 6) P. varsoviensis Worthen, 1882, is assigned to Sarocrinus and is a senior synonym of P. tentaculatus Worthen, 1883, P. granilineus Miller and Gurley, 1890a, Z. pocillum Miller, 1891b, S. nitidus Kirk, 1942, and S. plenus Kirk, 1942; 7) P. asperatus Worthen, 1882, is assigned to Sarocrinus; 8) Worthenocrinus n. gen. is described and W. paterus n. sp. is assigned to this new genus; 9) Poteriocrinus iowensis Worthen, 1882, is considered a nomen dubium and placed in incertae sedis.

1993 ◽  
Vol 67 (4) ◽  
pp. 614-639 ◽  
Author(s):  
Thomas W. Kammer ◽  
William I. Ausich

Species of the late Osagean and early Meramecian advanced cladid crinoid genera (suborder Poteriocrinina) Abrotocrinus, Aulocrinus, Cromyocrinus. Decadocrinus, Dinotocrinus, Holcocrinus, Hylodecrinus, Lanecrinus n. gen., Pachylocrinus, Parascytalocrinus n. gen., and Stinocrinus from Illinois, Indiana, Iowa, Kentucky, and Missouri are redescribed and redefined from study of type material and museum collections. Nomenclatural and systematic acts include the following: 1) Abrotocrinus cymosus Miller and Gurley, 1890a, is a valid species and not a synonym of A. coreyi (Meek and Worthen, 1869); 2) A. coreyi is a senior synonym of Poteriocrinus orestes Worthen, 1882; 3) Scaphiocrinus manus Miller and Gurley, 1890a, is assigned to Abrotocrinus; 4) Abrotocrinus granulatus n. sp. is described; 5) S. bellus Miller and Gurley, 1890a, is assigned to Aulocrinus and is a senior synonym of Aulocrinus agassizi Wachsmuth and Springer, 1897; 6) S. orbicularis Hall, 1861a, is assigned to Cromyocrinus; 7) P. latidactylus Worthen, 1882, is assigned to Dinotocrinus; 8) S. gibsoni White, 1878, is assigned to Hylodecrinus with P. nauvooensis Worthen, 1882, questionably placed in synonymy; 9) P. briareus Worthen, 1882, is assigned to Hylodecrinus and is a senior synonym of S. lacunosus Miller and Gurley, 1890a, S. boonvillensis Miller, 1891b, S. constrictus Miller, 1891a, and S. arrosus Miller and Gurley, 1893; 10) S. bonoensis Miller and Gurley, 1890a, is assigned to Hylodecrinus; 11) Hylodecrinus robustus n. sp. is described; 12) P. validus Worthen, 1882, is questionably assigned to Hylodecrinus; 13) Lanecrinus n. gen. is described and the following species are assigned to this new genus: S. halli Hall, 1861 a, S. depressus Meek and Worthen, 1870, P. milleri Wetherby, 1881, P. fountainensis Worthen, 1882, S. repertus Miller and Gurley, 1890a, and Ramulocrinus consectatus Strimple and Watkins, 1969; P. otterensis Worthen, 1882, is questionably assigned to Lanecrinus; 14) Lanecrinus depressus is the senior synonym of P. ulrichi Worthen in Miller, 1889, and S. maniformis Miller, 1892; 15) Lanecrinus repertus is the senior synonym of S. gorbyi Miller, 1891a; 16) P. asper Worthen, 1882, is assigned to Pachylocrinus and is a senior synonym of S. extensus Wachsmuth and Springer, 1886; 17) Parascytalocrinus n. gen. is described and the following species are assigned to this new genus: Poteriocrinus hamiltonensis Worthen, 1882, and Scytalocrinus validus Wachsmuth and Springer, 1897; 18) Poteriocrinites mcadamsi Meek and Worthen, 1873, is questionably placed in synonymy with Parascytalocrinus hamiltonensis; and 19) Poteriocrinus burketi Worthen, 1882, nomen dubium and Zeacrinus dubius Miller and Gurley, 1890a, are placed in Incertae Sedis.Cromyocrinus orbicularis is the oldest species of its genus extending the range of the genus downward from the Upper to the Lower Mississippian.


1996 ◽  
Vol 70 (5) ◽  
pp. 835-866 ◽  
Author(s):  
Thomas W. Kammer ◽  
William I. Ausich

Species of the late Osagean and early Meramecian primitive cladid crinoid generaAtelestocrinus, Barycrinus, Cestocrinus, Costalocrinus, Cyathocrinites, Meniscocrinusn. gen.,Parisocrinus, Pellecrinus, andSaccosomopsisfrom Illinois, Indiana, Iowa, Kentucky, Missouri, and Tennessee are reviewed, redescribed, and redefined from study of type material, museum collections, and field collections. Nomenclatural and systematic acts include the following: 1)Barycrinus spurius(Hall, 1858) is a senior synonym ofCyathocrinus tumidusHall, 1858,C. protuberansHall, 1858, andB. mammatusMeek and Worthen, 1873; 2)B. rhombiferus(Owen and Shumard, 1852a) is a senior synonym ofC. sculptilisHall, 1860,C. thomaeHall, 1860,C. hoveyiHall, 1861a,C. wachsmuthiMeek and Worthen, 1861,B. herculeusMeek and Worthen, 1868,B. pentagonusMeek and Worthen, 1873,B. striatusWorthen, 1875,B. boonvillensisMiller, 1891b,B. formosusMiller and Gurley, 1894,B. washingtonensisMiller and Gurley, 1895, andB. elrodiMiller and Gurley, 1896a; 3)B. magister(Hall, 1858) is a senior synonym ofC. solidusHall, 1861a andB. magnificusMeek and Worthen, 1868; 4)B. stellatus(Hall, 1858) is a senior synonym ofC. bullatusHall, 1858,C. angulatusMeek and Worthen, 1860,C. quinquelobusMeek and Worthen, 1865, andB. astericusVan Sant, 1964; 5)B. crassibrachiatus(Hall, 1860) is a senior synonym ofB. princepsMiller and Gurley, 1890a; 6)B. geometricusMeek and Worthen, 1873, is considered a nomen dubium; 7)B. benedicti(Miller, 1891a) is considered a nomen dubium; 8)Cyathocrinus signatusMiller and Gurley, 1894, is assigned toCestocrinusand is a senior synonym ofCestocrinus striatusKirk, 1940; 9)Cyathocrinites iowensis(Owen and Shumard, 1850) is a senior synonym ofC. malvaceusHall, 1858,C. divaricatusHall, 1858,C. rotundatusHall, 1858,C. viminalisHall, 1861a,C. parvibrachiatusHall, 1861a,C. hamiltonensisWorthen, 1882,C. nodosusWachsmuth and Springer, 1890,C. brevisacculusWachsmuth and Springer, 1890,C. opimusMiller and Gurley, 1890a, andC. gurleyiMiller, 1891a; 10)C. kelloggi(White, 1862) is a senior synonym ofC. subtumidusMeek and Worthen, 1865; 11)C. farleyi(Meek and Worthen, 1866b) is a senior synonym ofC. andersoniMiller and Gurley, 1894,C. granulosusRowley, 1902, andC. snivelyiRowley, 1902; 12)C. harrodi(Wachsmuth and Springer, 1880) is a senior synonym ofC. boonvillensisMiller, 1891b,C. gorbyiMiller, 1892b, andC. astralusKammer, 1984; 13)Meniscocrinusn. gen. is described andM. magnitubusn. sp. is assigned to this new genus; 14)C. labyrinthicusMiller, 1891a, is assigned toParisocrinus; 15)C. intermediusHall, 1858, is assigned toPellecrinus; and 16)C. insperatusLyon, 1869, is assigned toSaccosomopsisand is a senior synonym ofC.?poteriumMeek and Worthen, 1870.


2003 ◽  
Vol 77 (1) ◽  
pp. 121-138 ◽  
Author(s):  
Thomas W. Kammer ◽  
Forest J. Gahn

All 19 known species of the primitive cladid crinoid genera Atelestocrinus, Cyathocrinites, Goniocrinus, Parisocrinus, Pellecrinus, and Zygotocrinus from the early Osagean Burlington Limestone of the North American midcontinent are reviewed and redescribed or, where necessary, redefined. Nine of these species are illustrated for the first time herein. Sixteen are considered valid, including C. deroseari n. sp. Of the remaining three species, one is left in open nomenclature, and two are considered nomen dubia. Pellecrinus is recognized for the first time from the Burlington Limestone, although the specimens can not be identified to the species level and are left in open nomenclature.Cyathocrinites ranges from the Middle Silurian to at least the Middle Mississippian. During the Early Mississippian Cyathocrinites experienced an evolutionary radiation with a maximum diversity of nine species in the Burlington Limestone. Phylogenetic relationships were investigated in a parsimony-based phylogenetic analysis by combining morphologic data from the Burlington species with data from the four other species of Cyathocrinites from the late Osagean and early Meramecian of the east-central United States. The Kinderhookian C. chouteauensis (Miller and Gurley, 1896) served as the outgroup. A phylogenetic analysis of 14 species of Mississippian Cyathocrinites yielded a single most parsimonious tree with a length of 28 steps (C. I. = 0.607, H. I. = 0.392, R. I. = 0.718, R. C. = 0.436). Results of this analysis suggest that at least two major clades existed within Mississippian Cyathocrinites. One clade contains C. sampsoni (Miller, 1891b), C. gilesi (Wachsmuth and Springer, 1878), C. farleyi (Meek and Worthen, 1866), and C. barydactylus (Wachsmuth and Springer, 1878). The second clade contains C. iowensis, C. kelloggi (White, 1862), C. barrisi (Hall, 1861a), C. rigidus, C. deroseari n. sp., C. asperrimus (Springer, 1911), C. lamellosus (White, 1863), and C. harrodi (Wachsmuth and Springer, 1880). Cyathocrinites multibrachiatus forms a polytomy with these two clades. Members of the first clade exhibit a unique overall morphology present only during the Mississippian, suggesting the clade arose during this time. Members of the second clade, plus C. multibrachiatus, exhibit some characters present in Cyathocrinites species as old as the Middle Silurian and, thus, may have its roots among Silurian and Devonian species.


1992 ◽  
Vol 66 (3) ◽  
pp. 461-480 ◽  
Author(s):  
Thomas W. Kammer ◽  
William I. Ausich

Species of the late Osagean and early Meramecian advanced cladid crinoid genera (suborder Poteriocrinina)Armenocrinus, Bollandocrinus, Corythocrinus, Cosmetocrinus, Culmicrinus, Cydrocrinus, Histocrinus, Hypselocrinus, Lebetocrinus, Ophiurocrinus, Poteriocrinites, Scytalocrinus, Springericrinus, andUlrichicrinusfrom Illinois, Indiana, Iowa, and Missouri are redescribed and redefined from study of type material. Nomenclatural and systematic acts include the following: 1)Poteriocrinus tenuidactylusWorthen, 1882, is a senior synonym ofScaphiocrinus obscurusWachsmuth and Springer, 1886, and is assigned toArmenocrinus; 2)P. verusMiller and Gurley, 1890, is assigned toCulmicrinus; 3)P. mammaeformisWortheninMiller, 1889, is questionably assigned toCulmicrinus; 4)Cydrocrinus coxanus(Worthen, 1882) is a junior synonym ofC. concinnus(Meek and Worthen, 1870); 5)Decadocrinus stellatusLane and Howell, 1986, is questionably assigned toHistocrinus; 6)P. crawfordsvillensisMiller and Gurley, 1890, is a junior synonym ofHypselocrinus hoveyi(WortheninWorthen and Meek, 1875); 7)Hypselocrinus boonvillensis(Miller, 1891) andH. neglectus(Miller and Gurley, 1896) are junior synonyms ofH. arcanus(Miller and Gurley, 1890); 8)Hypselocrinus vansantin. sp. is described; 9)Hypselocrinus indianensis(Meek and Worthen, 1865) is a nomem dubium; 10)Poteriocrinus arachnaeformisWorthen, 1882, is a valid species ofPoteriocrinites; 11)P. cantonensisMiller and Gurley, 1890, is assigned toScytalocrinus; 12)Ulrichicrinus coryphaeus(Miller, 1891),P. amoenusMiller, 1891, andP. brittsiMiller, 1891, are junior synonyms ofU. agnatus(Miller, 1891); and 13)P. illinoisensisWorthen, 1882, is questionably assigned toUlrichicrinus.Bollandocrinus? sp. from the Keokuk Limestone might represent the first report of the genus from North America, being previously known only from the lower Carboniferous of Great Britain.Ophiurocrinussp. from the Keokuk Limestone is the first report of this genus from the Osagean, being previously known only from the Chesterian of North America and the upper Carboniferous of Europe.


1985 ◽  
Vol 16 (1) ◽  
pp. 27-67 ◽  
Author(s):  
Henrik Enghoff

AbstractThe family Nemasomatidae is redefined to include onty genera with all sterna secondarily free from pleurotergites. Comments are given on the included genera, viz., Antrokoreana, Basoncopus gen. n. (type-species B. filiformis sp. n.) (Kazakhstan), Dasynemasoma, Thalassisobates, Sinostemmiulus, Nemasoma, and Orinisobates. Isobates coiffaiti Demange, 1961 is synonymized with Thalassisobates littoralis (Silvestri, 1903). Orinisobates is revised and shown to include O. soror sp. n. (Kuril Islands), O. microthylax sp. n. (Kamchatka and Siberia), O. gracilis (Verhoeff, 1933) (NW China), O. sibiricus (Gulicka, 1963) (Altai region, Kazakhstan), O. kasakstanus (Lohmander, 1933) (Kazahkstan), O. nigrior (Chamberlin, 1943) (eastern United States), O. utus (Chamberlin, 1912) (northwestern United States), and O. expressus (Chamberlin, 1941) (northwestern United States and adjacent Canada). Mimolene oregona Chambertin, 1941 and M. sectile Loomis & Schmitt, 1971 are synonymized with O. expressus. A possible case of parthenogenesis in O. microthylax is recorded. Evidence is presented for the following sister-group relationships: Antrokoreana + (Basoncopus + (Dasynemasoma + (Thalassisobates + (Sinostemmiulus + (Orinisobates + Nemasoma))))). The position of Basoncopus is uncertain, and O. soror may belong in a separate genus and constitute the sister-group of Orinisohates + Nemasoma. If soror does belong in Orinisobates, it is the sister-group of all its congeners. The American species of Orinisobates are shown probably to constitute a monophyietic group. The family is suggested to have originated in the eastern Palearctic region, Orinisobates having invaded North America via the Bering Bridge. Doubtful species and species erroneously assoiciated with the Nemasomatidae are listed. The genera Okeanobates and Yosidaiulus are excluded from the family and referred to Okeanobatidae stat. n. in superfamily Blaniuloidea. The genera Trichonemasoma, Telsonemasoma, and Chelojulus are also excluded from the Nemasomatidae and relegated to Julida incertae sedis.


Zootaxa ◽  
2018 ◽  
Vol 4444 (1) ◽  
pp. 1 ◽  
Author(s):  
ERIC SIMON ◽  
BERNARD MOTTEQUIN

Size reduction and development of a simplified brachidial structure occurred several times during the long evolution of the Phylum Brachiopoda. Even Recent forms may be micromorphic and paedomorphic with reduced brachidia or none at all. A revision of the Maastrichtian (Late Cretaceous) Terebratella (Morrisia?) suessi Bosquet, 1859 has allowed us to erect a new genus, Jagtithyris gen. nov., because its singular brachidium development does not match any platidiid structure. Such a brachidium has also been observed in another European Late Cretaceous brachiopod, which indicates that this type was not a unique morphological curiosity. This species is the micromorphic Campanian-Maastrichtian Leptothyrellopsis polonicus Bitner & Pisera, 1979, which has brachidial structures in common with Jagtithyris suessi comb. nov., although a number of differences have been observed. The genera Leptothyrellopsis and Jagtithyris gen. nov., are included in a new family, Jagtithyrididae fam. nov. During an ongoing revision of extant brachiopod faunas we have been led to recognize a link between this family and representatives of the genus Simplicithyris Zezina, 1976. The taxonomic position of this peculiar group is also discussed. 


2014 ◽  
Vol 15 (3) ◽  
pp. 92-96 ◽  
Author(s):  
Guirong Zhang ◽  
Carl A. Bradley

Historically, frogeye leaf spot (FLS, caused by Cercospora sojina) of soybean has been observed more frequently in the southern United States than the north-central United States. However, in recent years, FLS field observations have increased in the north-central United States. To better understand the survival rate of C. sojina in Illinois, a field study was conducted at three locations: Monmouth (west-central Illinois), Urbana (east-central Illinois), and Dixon Springs (southeastern Illinois). At each location, soybean leaves affected by FLS were placed at depths of 0, 10, and 20 cm and retrieved at different durations up to 24 months. To determine the viability of C. sojina in the collected leaves, a greenhouse bioassay was used. Survival of C. sojina declined equally with time at all three locations through 19 months. After 24 months, C. sojina from leaves that had been placed at Monmouth and Urbana were no longer viable, whereas leaves that had been placed at Dixon Springs produced viable inoculum. Depth of leaf placement had no effect on survival of C. sojina at any of the locations. These results suggest that planting a nonhost crop for two years in central Illinois will reduce levels of C. sojina inoculum to a negligible amount; however, soybean farmers in southern Illinois may need a longer rotation for FLS management. Accepted 10 May 2014. Published 23 July 2014.


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