Harvesting at the Cohort and Population Levels
This chapter explores dynamical behaviors that go beyond globally stable outcomes to include alternate stable states, and non-equilibrium behaviors. The possibility of multiple equilibria emerges quite readily in models with non-linear harvesting functions. In practice, most fisheries management protocols at least implicitly assume that harvested populations have well-behaved stable equilibrium properties. If this is not the case, then sudden changes (including collapse) can occur and be totally unanticipated. This chapter describes the spectrum of single-species harvesting models from biomass dynamics models that do not include age or size structure, to delay–difference models with a simple demographic structure, to full age-structured models. Dynamic-pool models combine yield per recruit and egg-per-recruit with a stock-recruitment model to obtain an equilibrium yield curve. These single-species models are used to estimate biological reference points with which to assess stock status.