Anatomical, morphometric, and stratigraphic analyses of theropod biodiversity in the Upper Cretaceous (Campanian) Dinosaur Park Formation

2021 ◽  
pp. 1-15
Author(s):  
Thomas M. Cullen ◽  
Lindsay Zanno ◽  
Derek W. Larson ◽  
Erinn Todd ◽  
Philip J. Currie ◽  
...  
Keyword(s):  
High Resolution ◽  
North America ◽  
Fossil Record ◽  
Upper Cretaceous ◽  
Environmental Changes ◽  
Morphometric Analyses ◽  
Variable Distribution ◽  
Taxic Diversity ◽  
Dinosaur Park Formation ◽  
Broad Scale

The Dinosaur Park Formation (DPF) of Alberta, Canada, has produced one of the most diverse dinosaur faunas, with the record favouring large-bodied taxa, in terms of number and completeness of skeletons. Although small theropods are well documented in the assemblage, taxonomic assessments are frequently based on isolated, fragmentary skeletal elements. Here we reassess DPF theropod biodiversity using morphological comparisons, high-resolution biostratigraphy, and morphometric analyses, with a focus on specimens/taxa originally described from isolated material. In addition to clarifying taxic diversity, we test whether DPF theropods preserve faunal zonation/turnover patterns similar to those previously documented for megaherbivores. Frontal bones referred to a therizinosaur (cf. Erlikosaurus), representing among the only skeletal record of the group from the Campanian–Maastrichtian (83–66 Ma) fossil record of North America, plot most closely to troodontids in morphospace, distinct from non-DPF therizinosaurs, a placement supported by a suite of troodontid anatomical frontal characters. Postcranial material referred to cf. Erlikosaurus in North America is also reviewed and found most similar in morphology to caenagnathids, rather than therizinosaurs. Among troodontids, we document considerable morphospace and biostratigraphic overlap between Stenonychosaurus and the recently described Latenivenatrix, as well as a variable distribution of putatively autapomorphic characters, calling the validity of the latter taxon into question. Biostratigraphically, there are no broad-scale patterns of faunal zonation similar to those previously documented in ornithischians from the DPF, with many theropods ranging throughout much of the formation and overlapping extensively, possibly reflecting a lack of sensitivity to environmental changes, or other cryptic ecological or evolutionary factors.

The first record of spiral coprolites from the Dinosaur Park Formation (Judith River Group, Upper Cretaceous) southern Alberta, Canada

1995 ◽  
Vol 69 (6) ◽  
pp. 1191-1194 ◽  
Author(s):  
Clive E. Coy
Keyword(s):  
North America ◽  
Upper Cretaceous ◽  
First Record ◽  
Niobrara Formation ◽  
Southern Alberta ◽  
Dinosaur Park Formation

Spiral coprolites from the Upper Cretaceous of North America are poorly known. Enterospirae (fossilized intestines) reported from the Upper Cretaceous Niobrara Formation of western Kansas (Stewart, 1978) were disputed by McAllister (1985), who felt they represented spiral coprolites similar to those described from the Permian by Neumayer (1904). Previously described coprolites from the Upper Cretaceous of Alberta are small, unstructured, ellipsoidal forms thought to derive from a crocodilian or terrestrial, carnivorous reptile of necrophagic or piscivorous habits (Waldman, 1970; Waldman and Hopkins, 1970).

scholarly journals A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada

2017 ◽  
Vol 54 (8) ◽  
pp. 813-826 ◽  
Author(s):  
David C. Evans ◽  
Thomas M. Cullen ◽  
Derek W. Larson ◽  
Adam Rego
Keyword(s):  
New Species ◽  
North America ◽  
Alpha Diversity ◽  
Width Ratio ◽  
New Taxon ◽  
Lateral Region ◽  
Morphometric Analyses ◽  
Dinosaur Park Formation ◽  
Horseshoe Canyon Formation ◽  
A New Species

Troodontid material from the Maastrichtian of North America is extremely rare, beyond isolated teeth from microvertebrate sites. Here we describe troodontid frontals from the early Maastrichtian Horseshoe Canyon Formation (Horsethief Member). The most complete specimen, TMP 1993.105.0001, is notably foreshortened and robust when compared with numerous specimens referred to Troodon from the Dinosaur Park Formation, and exhibits several characteristics that distinguish it from other Late Cretaceous troodontids. Morphometric analyses reinforce shape differences between TMP 1993.105.0001 and other North American troodontids, and show that proportional differences are independent of size. We therefore erect a new taxon, Albertavenator curriei gen. et sp. nov., which is diagnosed by the following autapomorphies: (1) primary supraciliary foramen is truncated anteriorly by the lacrimal contact; (2) superficial (ectocranial) surface of the frontal proportionally shorter than all known troodontids, with a length to width ratio under 1.3; and (3) frontoparietal contact in which an enlarged lappet of the frontal extends medially to extensively overlap the lateral region of the anteromedial process of the parietal. Interestingly, tooth and jaw morphology from the single relatively complete dentary recovered from the Horseshoe Canyon cannot be distinguished from dentaries and teeth from the Dinosaur Park Formation. If the dentary and teeth from the Horsethief Member of the Horseshoe Canyon Formation prove to belong to A. curriei, extensive overlap in tooth morphology between the Dinosaur Park and Horseshoe Canyon formations reinforces the notion that tooth morphotypes do not exhibit strong correspondence to species alpha diversity, and may encompass multiple closely related taxa.

scholarly journals A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada

2018 ◽  
Author(s):  
David Evans ◽  
Thomas Cullen ◽  
Derek Larson ◽  
Adam Rego
Keyword(s):  
New Species ◽  
North America ◽  
Alpha Diversity ◽  
Width Ratio ◽  
New Taxon ◽  
Lateral Region ◽  
Morphometric Analyses ◽  
Dinosaur Park Formation ◽  
Horseshoe Canyon Formation ◽  
A New Species

Troodontid material from the Maastrichtian of North America is extremely rare, beyond isolated teeth from microvertebrate sites. Here we describe troodontid frontals from the early Maastrichtian Horseshoe Canyon Formation (Horsethief Member). The most complete specimen, TMP 1993.105.0001, is notably foreshortened and robust when compared with numerous specimens referred to Troodon from the Dinosaur Park Formation, and exhibits several characteristics that distinguish it from other Late Cretaceous troodontids. Morphometric analyses reinforce shape differences between TMP 1993.105.0001 and other North American troodontids, and show that proportional differences are independent of size. We therefore erect a new taxon, Albertavenator curriei gen. et sp. nov., which is diagnosed by the following autapomorphies: (1) primary supraciliary foramen is truncated anteriorly by the lacrimal contact; (2) superficial (ectocranial) surface of the frontal proportionally shorter than all known troodontids, with a length to width ratio under 1.3; and (3) frontoparietal contact in which an enlarged lappet of the frontal extends medially to extensively overlap the lateral region of the anteromedial process of the parietal. Interestingly, tooth and jaw morphology from the single relatively complete dentary recovered from the Horseshoe Canyon cannot be distinguished from dentaries and teeth from the Dinosaur Park Formation. If the dentary and teeth from the Horsethief Member of the Horseshoe Canyon Formation prove to belong to A. curriei, extensive overlap in tooth morphology between the Dinosaur Park and Horseshoe Canyon formations reinforces the notion that tooth morphotypes do not exhibit strong correspondence to species alpha diversity, and may encompass multiple closely related taxa.

First report of protoceratopsians (Neoceratopsia) from the Late Cretaceous Judith River Group, Alberta, Canada

1998 ◽  
Vol 35 (7) ◽  
pp. 820-826 ◽  
Author(s):  
Michael J Ryan ◽  
Philip J Currie
Keyword(s):  
North America ◽  
Late Cretaceous ◽  
Fossil Record ◽  
Coastal Environments ◽  
First Report ◽  
Skeletal Material ◽  
First Occurrence ◽  
Dinosaur Park Formation ◽  
Milk River Formation

Protoceratopsians are best known in North America from associated skeletal material of Montanoceratops from the early Maastrichtian of Montana and Campanian of Alberta and Leptoceratops from the late Maastrichtian of Alberta and Wyoming. We report here the first occurrence of protoceratopsian elements from the middle Campanian (Dinosaur Park Formation) of Alberta. The specimens consist of a fragmentary right dentary and an almost complete left dentary which can be referred to Leptoceratops sp. Recent examination of Albertan microvertebrate material has identified cf. protoceratopsians teeth from the latest Santonian (Milk River Formation), extending the record of Albertan protoceratopsians back almost 20 million years. The rarity of these small ornithischians in the fossil record of Alberta may have been due to ecological exclusion from the wet, coastal environments that were preferred by the larger, more abundant ceratopsids.

Scolosaurus cutleri (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada

2013 ◽  
Vol 50 (2) ◽  
pp. 171-182 ◽  
Author(s):  
Paul Penkalski ◽  
William T. Blows
Keyword(s):  
North America ◽  
Upper Cretaceous ◽  
Shape Recognition ◽  
Pelvic Morphology ◽  
Valid Taxon ◽  
Dinosaur Park Formation

The synonymy of the ankylosaurid dinosaur Scolosaurus with Euoplocephalus has been widely accepted since the 1970s. However, Scolosaurus cutleri exhibits differences which separate it from Euoplocephalus tutus and Dyoplosaurus acutosquameus. Although the holotype of Euoplocephalus is fragmentary, several other specimens can be reliably referred to this taxon and thus used for comparison. Scolosaurus differs from Euoplocephalus in cervical half-ring, osteoderm, and forelimb morphology. Scolosaurus differs from Dyoplosaurus primarily in pelvic morphology and osteoderm shape. Recognition of Scolosaurus as a valid taxon adds to the growing concept that the Upper Cretaceous ankylosaurid fauna of North America was more diverse than previously thought.

scholarly journals A small caseid synapsid, Arisierpeton simplex gen. et sp. nov., from the early Permian of Oklahoma, with a discussion of synapsid diversity at the classic Richards Spur locality

PeerJ ◽  
2019 ◽  
Vol 7 ◽  
pp. e6615 ◽  
Author(s):  
Robert R. Reisz
Keyword(s):  
North America ◽  
Fossil Record ◽  
Middle Permian ◽  
Lower Permian ◽  
Early Permian ◽  
Faunal Assemblage ◽  
Lower Jaw ◽  
Continental Deposits ◽  
Faunal List ◽  
Taxic Diversity

The fossil record of caseids, a clade of faunivorous to large herbivorous Permian synapsids, is unusual in having a poorly documented history. Although Kungurian caseids are common in the well-known continental deposits of North America, and the fossil record of the group extends into the middle Permian (Guadalupian), with the presence of the large caseid Ennatosaurus in the Mezen Basin faunal assemblage, only two other occurrences are known in older Permian age sediments. One is an undescribed caseid from the Bromacker Quarry in Germany, and the second is Oromycter from the lower Permian of Richards Spur, Oklahoma. The former is known from several articulated skeletons, but the latter is known only from a handful of skeletal elements, including elements of the snout and lower jaw, some phalanges, and a few vertebrae. Here the fragmentary tooth bearing elements and dorsal vertebrae of another small caseid from Richards Spur are described, with a discussion of its significance in the context of caseid evolution, and the continuously expanding faunal list and taxic diversity at this locality.

scholarly journals A small azhdarchoid pterosaur from the latest Cretaceous, the age of flying giants

2016 ◽  
Vol 3 (8) ◽  
pp. 160333 ◽  
Author(s):  
Elizabeth Martin-Silverstone ◽  
Mark P. Witton ◽  
Victoria M. Arbour ◽  
Philip J. Currie
Keyword(s):  
North America ◽  
Late Cretaceous ◽  
Fossil Record ◽  
Upper Cretaceous ◽  
Large Species ◽  
Time Of Death ◽  
The West ◽  
New Material ◽  
The Individual ◽  
Continental Interior

Pterosaur fossils from the Campanian–Maastrichtian of North America have been reported from the continental interior, but few have been described from the west coast. The first pterosaur from the Campanian Northumberland Formation (Nanaimo Group) of Hornby Island, British Columbia, is represented here by a humerus, dorsal vertebrae (including three fused notarial vertebrae), and other fragments. The elements have features typical of Azhdarchoidea, an identification consistent with dominance of this group in the latest Cretaceous. The new material is significant for its size and ontogenetic stage: the humerus and vertebrae indicate a wingspan of ca 1.5 m, but histological sections and bone fusions indicate the individual was approaching maturity at time of death. Pterosaurs of this size are exceedingly rare in Upper Cretaceous strata, a phenomenon commonly attributed to smaller pterosaurs becoming extinct in the Late Cretaceous as part of a reduction in pterosaur diversity and disparity. The absence of small juveniles of large species—which must have existed—in the fossil record is evidence of a preservational bias against small pterosaurs in the Late Cretaceous, and caution should be applied to any interpretation of latest Cretaceous pterosaur diversity and success.

HIGH-RESOLUTION SEDIMENTARY [HG] RECORDS ACROSS THE PETM FROM A TERRESTRIAL LATITUDINAL TRANSECT IN NORTH AMERICA

2020 ◽  
Author(s):  
Natalie Sorrem ◽  
◽  
Clara L. Meier ◽  
Gabriel J. Bowen ◽  
Brady Z. Foreman ◽  
...  
Keyword(s):  
High Resolution ◽  
North America ◽  
Latitudinal Transect

Neritoptyx hogansoninew species (Gastropoda, Mollusca) from the Upper Cretaceous Fox Hills Formation on the Dakota Isthmus, western United States

2018 ◽  
Vol 93 (2) ◽  
pp. 278-290
Author(s):  
J. Mark Erickson
Keyword(s):  
Heat Transfer ◽  
United States ◽  
New Species ◽  
North America ◽  
Upper Cretaceous ◽  
Gulf Coast ◽  
Western United States ◽  
Western Interior Seaway ◽  
Decapod Crustaceans ◽  
Upper Maastrichtian

AbstractIn midcontinent North America, the Fox Hills Formation (Upper Cretaceous, upper Maastrichtian) preserves the last marine faunas in the central Western Interior Seaway (WIS).Neritoptyx hogansoninew species, a small littoral snail, exhibited allometric change from smooth to corded ornament and rounded to shouldered shape during growth. Specimens preserve a zig-zag pigment pattern that changes to an axial pattern during growth.Neritoptyx hogansoninew species was preyed on by decapod crustaceans, and spent shells were occupied by pagurid crabs. Dead mollusk shells, particularly those ofCrassostrea subtrigonalis(Evans and Shumard, 1857), provided a hard substrate to which they adhered on the Fox Hills tidal flats. This new neritimorph gastropod establishes a paleogeographic and chronostratigraphic proxy for intertidal conditions on the Dakota Isthmus during the late Maastrichtian. Presence of a neritid extends the marine tropical/temperate boundary in the WIS northward to ~44° late Maastrichtian paleolatitude. Late Maastrichtian closure of the isthmus subsequently altered marine heat transfer by interrupting northward flow of tropical currents from the Gulf Coast by as much as 1 to 1.5 million years before the Cretaceous ended.UUID:http://zoobank.org/3ba56c07-fcca-4925-a2f0-df663fc3a06b

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