scholarly journals Towards a comprehensive taxonomic revision of the Neotropical dung beetle subgenus Deltochilum (Deltohyboma) Lane, 1946 (Coleoptera: Scarabaeidae: Scarabaeinae): Division into species-groups

PLoS ONE ◽  
2021 ◽  
Vol 16 (1) ◽  
pp. e0244657
Author(s):  
Arturo González-Alvarado ◽  
Fernando Z. Vaz-de-Mello

Deltochilum Eschscholtz, 1822 is perhaps the most speciose genus of the tribe Deltochilini sensu Tarasov & Dimitrov (2016) (Scarabaeidae: Scarabaeinae) and has been traditionally divided into eight subgenera. Among them, the subgenus Deltohyboma Lane, 1946, is the most speciose with 47 species, five of which are described here (D. genieri sp. nov., D. gilli sp. nov., D. susanae sp. nov., D. bolivariensis sp. nov. and D. inesae sp. nov.), and at least 165 species still undescribed. Due to the large number of species, and for practical purposes, the subgenus is here divided into 19 species-groups, with D. inesae sp. nov. left as incertae sedis. This division into species-groups will help in the reliable identification of species and will aid in the completion of the revision of subgenus. This report is the first part of the taxonomic revision of the subgenus Deltohyboma; it is based on the examination of all type specimens and almost 9,800 specimens of which approximately 1,200, mostly males, had their genitalia studied. The 19 species-groups recognized here are based mainly on characters described for the first time for Deltohyboma, namely, the state of a) the anterior margin of the clypeus (between the clypeal teeth), b) the internal margin of hypomera, c) the ventral face of the protibia, d) the posterior margin of the metafemur, and e) several new characters resulting from the first detailed study of the aedeagus and the endophallus of the group. Diagnosis, description, geographic distribution, composition and identifications keys (for males as well as males and females) for the species-groups are provided.

Zootaxa ◽  
2019 ◽  
Vol 4596 (1) ◽  
pp. 1 ◽  
Author(s):  
CONTRERAS-FÉLIX GERARDO A. ◽  
FRANCKE B. OSCAR F.

Within the scorpion genus Vaejovis C.L. Koch, the “mexicanus” group is composed of species distributed in the mountains of México. This group presents taxonomic problems, because its characterization and the species included in the group have varied through the years. In the present work, we redefine this group based on several morphological characters, and we differentiate it from the other two species groups within the genus: “vorhiesi” and “nit dulus+nigrescens”. Additionally, five new species are described: Vaejovis ceboruco sp. nov., Vaejovis nanchititla sp. nov., Vaejovis santibagnezi sp. nov., Vaejovis talpa sp. nov. and Vaejovis tapalpa sp. nov; the males of three species are described for the first time (V. dugesi, V. nigrofemoratus and V. tesselatus); and the updated diagnosis for all species is included. Keys for the identification of males and females of the 30 species included in this group are given. Lastly, notes on the natural history and distribution of some species are provided, with maps of known distribution for all the species.


Zootaxa ◽  
2011 ◽  
Vol 3029 (1) ◽  
pp. 1 ◽  
Author(s):  
S. D. BIJU ◽  
INES VAN BOCXLAER ◽  
STEPHEN MAHONY ◽  
K. P. DINESH ◽  
C. RADHAKRISHNAN ◽  
...  

A taxonomic revision of the genus Nyctibatrachus is presented. As a result, we recognize 27 species, 12 of which are new to science and described herein. Three ‘lost’ species—Nyctibatrachus sanctipalustris, Nyctibatrachus sylvaticus and Nannobatrachus kempholeyensis are rediscovered after their original descriptions and the latter two species are neotypified. Four species are lectotypified (Nannobatrachus beddomii, Nyctibatrachus deccanensis, Nyctibatrachus major and Nyctibatrachus sanctipalustris). The taxonomic revision and description of new species is based on new morphological, behavioural, ecological and molecular data from available type specimens and/or recent collections from throughout the distribution range of Nyctibatrachus. Specimens from the type localities of all previously described species were collected and discussed along with the type specimens. Nyctibatrachus sholai is considered a junior subjective synonym of N. deccanensis. Nyctibatrachus sanctipalustris modestus is resurrected from synonymy, but allocated to the genus Fejervarya. A new gland, the subocular gland is introduced to anuran morphological terminology. Both the courtship behaviour and ovipositional sites of four species are described for the first time. The process of fertilisation was carried out in the absence of physical contact between males and females in all four species. Furthermore, previously unreported parental care behaviour is documented for the genus, including biparental egg attendance (probably clutch hydration) and egg guarding (active predator defense). A brief outline of tadpole development from egg to metamorph is provided for Nyctibatrachus aliciae, with figures of the major developmental stages.


Zootaxa ◽  
2020 ◽  
Vol 4792 (1) ◽  
pp. 1-155
Author(s):  
CAROLYN TRIETSCH ◽  
ISTVÁN MIKÓ ◽  
BRIANA EZRAY ◽  
ANDREW R. DEANS

We revise the species of Conostigmus Dahlbom, 1858 (Hymenoptera: Ceraphronoidea: Megaspilidae) found in North America, north of Mexico. We describe the following 12 new species: Conostigmus dessarti Trietsch & Mikó sp. nov.; C. duncani Trietsch sp. nov.; C. franzinii Trietsch & Mikó sp. nov.; C. johnsoni Trietsch & Mikó sp. nov.; C. lepus Trietsch sp. nov.; C. longiharpes Trietsch sp. nov.; C. michaeli Trietsch sp. nov.; C. minimus Trietsch & Mikó sp. nov.; C. muratorei Trietsch sp. nov.; C. musettiae Trietsch & Mikó sp. nov.; C. rosemaryae Trietsch sp. nov.; and C. washburni Trietsch sp. nov. We also redescribe the following 12 species: Conostigmus abdominalis (Boheman, 1832); C. bipunctatus Kieffer, 1907; C. dimidiatus (Thomson, 1858); C. erythrothorax (Ashmead, 1893); C. laeviceps (Ashmead, 1893); C. muesebecki Dessart & Masner, 1965; C. nigrorufus Dessart, 1997; C. obscurus (Thomson, 1858); C. orcasensis (Brues, 1909); C. pulchellus Whittaker, 1930; C. quadratogenalis Dessart & Cooper, 1975; and C. triangularis (Thomson, 1858). We report specimens of C. abdominalis (Boheman, 1832) and C. bipunctatus Kieffer, 1907 from the Nearctic for the first time, expanding the range from Palearctic to Holarctic for both species. We regard the following 19 species as having uncertain status due to reasons such as missing type specimens: Conostigmus ambiguus (Ashmead, 1893); C. bacilliger (Kieffer, 1906); C. bakeri Kieffer, 1908; C. californicus (Ashmead, 1893); C. canadensis (Ashmead,1888); C. crawfordi (Mann, 1920); C. harringtoni (Ashmead, 1888); C. hyalinipennis (Ashmead, 1887); C. inermis (Kieffer, 1906); C. integriceps (Kieffer, 1906); C. marylandicus (Ashmead, 1893); C. nevadensis (Kieffer, 1906); C. nigripes (Kieffer, 1906); C. ottawensis (Ashmead, 1888); C. pergandei (Ashmead, 1893); C. popenoei (Ashmead, 1893); C. rufoniger (Provancher, 1888); C. schwarzi (Ashmead, 1893); and C. trapezoidus Kieffer, 1908. We transfer Conostigmus arietinus (Provancher, 1887) to Dendrocerus Ratzeburg, 1852, and consider Conostigmus subinermis (Kieffer, 1907) to be absent from the Nearctic and limited to the Palearctic. The Nearctic species C. timberlakei Kamal, 1926 remains incertae sedis. We provide the name Conostigmus fulgidus Mikό and Trietsch to replace the junior homonym Conostigmus lucidus Mikό and Trietsch 2016. We provide a key for the identification of Nearctic Conostigmus species, and provide comments on their natural history. Finally, we infer evolutionary relationships within Megaspilinae using male genitalia and other morphological characters. This work represents the first in-depth study and revision of Conostigmus in North America, and contributes the first annotated identification key to Nearctic Conostigmus species. 


2014 ◽  
Vol 46 (1-2) ◽  
pp. 19 ◽  
Author(s):  
Paolo Audisio ◽  
Simone Sabatelli ◽  
Josef Jelìnek

A taxonomic revision of members of the genus <em>Meligethes</em> Stephens, 1830 is carried out. Taxonomic and distributional notes are provided on fiftythree <em>Meligethes</em> species, mostly known from the Eastern Palearctic and marginally from the Western Palearctic and the Oriental Regions. Among these, twentythree new species of <em>Meligethes</em> are diagnosed, figured and described: <em>M</em>. <em>argentithorax</em> sp. n. (Central China: Shaanxi, Shanxi), <em>M</em>. <em>aurantirugosus</em> sp. n. (Nepal),<em> M. aureolineatus</em> sp. n. (Central China: Sichuan), <em>M. aurifer</em> sp. n. (Central China: Shaanxi, Shanxi), <em>M. brassicogethoides</em> sp. n. (SW China: Yunnan), <em>M. clinei</em> sp. n. (SW China: Yunnan), <em>M. elytralis</em> sp. n. (Central China: Sichuan), <em>M. ferruginoides</em> sp. n. (Central China: Sichuan), <em>M.</em> <em>cinereoargenteus</em> sp. n. (Central China: Sichuan), <em>M. henan</em> sp. n. (Central China: Henan), <em>M. luteoornatus</em> sp. n. (SW China: Yunnan), <em>M. marmota</em> sp. n. (Nepal), <em>M. nivalis</em> sp. n. (SW and central China: Xizang and Chongqing), <em>M. martes</em> sp. n. (Central China: Shaanxi, Shanxi, Sichuan), <em>M. nigroaeneus</em> sp. n. (SW China: Yunnan), <em>M. occultus</em> sp. n. (SW China: Yunnan), <em>M</em>. <em>pseudochinensis</em> sp. n. (Central China: Hubei), <em>M. pseudopectoralis</em> sp. n. (SW China: Yunan),<em> M. schuelkei</em> sp. n. (Central China: Sichuan, Shaanxi), <em>M. simulator</em> sp. n. (Central-N China: Shanxi), <em>M. stenotarsus</em> sp. n. (SW China: Yunnan, Xizang), <em>M. tryznai</em> sp. n. (SW China: Yunnan), and <em>M. volkovichi</em> sp. n. (SW China: Yunnan). Revaluations at specific rank from synonymy are introduced for <em>Meligethes lutra</em> Solsky 1860, and for <em>M. melleus</em> Grouvelle, 1908. Three new synonymies are established: <em>Meligethes brevipilus</em> Kirejtshuk, 1980 = <em>M. auripilis</em> Reitter, 1889 (syn. n.), <em>Meligethes zakharenkoi</em> Kirejtshuk, 2005 = <em>M. shirakii</em> Sadanari Hisamatsu, 1956 (syn. n.), and <em>Meligethes shirozui</em> Sadanari Hisamatsu, 1965 =<em> M. wagneri</em> Rebmann, 1956 (syn. n.). Complete redescriptions are given for <em>Meligethes binotatus</em> Grouvelle, 1894, <em>M. castanescens</em> Grouvelle, 1903, <em>M. ferrugineus</em> Reitter, 1873, and <em>M. melleus</em> Grouvelle, 1908. The male of <em>Meligethes lloydi</em> Easton, 1968, is described and figured for the first time. The female genitalia of <em>Meligethes auricomus</em> Rebmann, 1956, <em>M. cinereus</em> Jelínek, 1978, and <em>M. griseus</em> Jelínek, 1978 are described and figured for the first time. Available information on insect-host-plant relationships and ecology are summarized for each species; probably all are associated as larvae with flowers of Rosaceae, chiefly of members of the closely related genera <em>Rosa</em> L., <em>Rubus</em> L., <em>Prunus</em> L., and <em>Crataegus</em> Tourn. ex L. All treated species are grouped in two here revaluated subgenera (<em>Meligethes</em> s.str. and <em>Odonthogethes</em> Reitter, 1871), and tentatively grouped also in species-groups and (when necessary) species-complexes, based on their morphology.


Zootaxa ◽  
2021 ◽  
Vol 5026 (4) ◽  
pp. 480-506
Author(s):  
LAURA A. MICHIE ◽  
R. S. K. BARNES ◽  
PAUL F. CLARK ◽  
WAYNE A. BENNETT ◽  
SIMON M. CRAGG

Ten species of fiddler crab are reported inhabiting the intertidal zone of a shore on Kaledupa Island, Indonesia. This is one of the highest recorded numbers of fiddler crab species living in sympatry, equating to over two-thirds of those known from the Wallacea biogeographic region and more than half of all those recorded from Indonesia. The descriptions to identify and distinguish these ten species are provided using a suite of characters e.g., carapace, major cheliped, male gonopods, gastric mills, life colouration in males and females, and notes on their ecology and distribution. Specimens were observed and collected in the Wakatobi National Park, near the village of Ambeua on Kaledupa island, Sulawesi Tenggara, Indonesia. Gastric mills are described for the first time for Gelasimus jocelynae, Paraleptuca crassipes, Tubuca coarctata, T. demani and T. dussumieri. A tabulation of anatomical features and colouration for all species in this study is provided as a support for field studies. It identifies features that support the recently proposed taxonomic revision of fiddler crabs by Shih et al. (2016).  


Author(s):  
Fernando Z. Vaz-de-Mello ◽  
Luis Gabriel De Oliveira Albuquerque Nunes ◽  
Vinícius Da Costa-Silva

A new species of Canthon is described, illustrated and the morphological similarities with related species are discussed. The species Canthon cleidecostae was named after Dr. Cleide Costa a great beetles specialist. The subgeneric category of species is considered incertae sedis until the taxonomic revision of the genus Canthon is fully addressed. Also, we provide a key for genera, subgenera and isolated species-groups that can be confused with the genus Canthon.


Zootaxa ◽  
2020 ◽  
Vol 4806 (1) ◽  
pp. 1-144
Author(s):  
STEFFEN BAYER ◽  
HUBERT HÖFER ◽  
HEIKO METZNER

We propose a revision of the spider genus Corythalia C.L. Koch, 1850 (Salticidae: Euophryini) with a revised genus diagnosis based on examination of all species available to us. In this paper we redescribe all previously described species from South America with revised species diagnoses and describe 20 new species from South America (and the nearby islands). For C. latipes, the type species of the genus Corythalia, a neotype is designated. In total, 52 nominal species of the genus are herein treated, 46 species are recognized as valid. The females of C. waleckii Taczanowski, 1871, C. luctuosa Caporiacco, 1954 and C. latipes (C.L. Koch, 1846) are described for the first time. Corythalia sellata Simon, 1901, erroneously considered as nomen nudum in the present version of the World Spider Catalog, is here recognised as a valid species. Corythalia fulgipedia Crane, 1948 is also considered a valid species and is removed from the synonymy of C. tropica (Mello-Leitão, 1939). One name is considered a nomen dubium (Corythalia variegata Caporiacco, 1954), two are nomina nuda (C. major Simon, 1901; C. dimidiata Simon, 1901). Two species are transferred to other genera: C. argyrochrysos (Mello-Leitão, 1946) to Pachomius Peckham & Peckham, 1896 as Pachomius argyrochrysos (Mello-Leitão, 1946), comb. nov. and C. heliophanina (Taczanowski, 1871) to Neonella Gertsch, 1936, as Neonella heliophanina (Taczanowski, 1871), comb. nov. under incertae sedis. One species is synonymised: C. barbipes (Mello-Leitão, 1939) is a junior synonym of C. cincta (Badcock, 1932), syn. nov. The new Corythalia species are: C. conferta sp. nov. (♂♀, Brazil), C. concinna sp. nov. (♀, Brazil), C. drepane sp. nov. (♂♀, Brazil), C. drepanopsis sp. nov. (♀, Brazil), C. antepagmenti sp. nov. (♂♀, Brazil), C. ricti Bayer, sp. nov. (♂, Guyana), C. protensa sp. nov. (♂, Brazil), C. gasnieri sp. nov. (♂, Brazil), C. verhaaghi sp. nov. (♀, Brazil), C. scutellaris Bayer, sp. nov. (♂♀, Ecuador), C. dakryodes Bayer, sp. nov. (♀, Colombia), C. foelixi Bayer, sp. nov. (♂♀, French Guiana), C. longiducta sp. nov. (♀, Brazil), C. latior sp. nov. (♂, Bolivia), C. trochophora Bayer, sp. nov. (♂, Ecuador), C. lineata Bayer, sp. nov. (♂, Guyana), C. hamulifera Bayer, sp. nov. (♂, Ecuador), C. tribulosa sp. nov. (♂, Colombia), C. flagrans sp. nov. (♂, Brazil) and C. fragilis sp. nov. (♂♀, Brazil). Illustrations are provided for all of the new species and for all (primary) type specimens of the species re-described. Hypotheses of possible relationships among the different species of Corythalia are discussed. 


Zootaxa ◽  
2019 ◽  
Vol 4679 (2) ◽  
pp. 318-340
Author(s):  
ROBERTO CALDARA ◽  
IVO TOŠEVSKI

The species of Rhinusa Stephens, 1829 (Curculionidae, Curculioninae, Mecinini) belonging to the R. linariae, R. herbarum, R. melas and R. mauritii groups are revised. The R. linariae group includes three species: R. linariae (Panzer, 1795), R. brisouti (Faust, 1891) and R. kumatschevi (Bajtenov, 1977). The R. herbarum group also includes three species: R. herbarum (H. Brisout de Barneville, 1863), R. mateui (Hoffmann, 1965) and R. vulpeculus (Reitter, 1907). The R. melas group includes two species: R. melas (Boheman, 1838) and R. korotyaevi sp. n. (Israel, Jordan). Lastly, the R. mauritii group is monobasic. The lectotypes of Gymnetron consors Desbrochers des Loges, 1898, Gymnetron herbarum H. Brisout de Barneville, 1863, Gymnetron heydeni Desbrochers des Loges, 1869, and Gymnetron melas Boheman, 1838 are designated. The following new synonymy is proposed: Rhinusa herbarum (H. Brisout de Barneville, 1863) = Gymnetron semirufum Desbrochers des Loges, 1898 (syn. n.). For each species treated, description or redescription, notes on type specimens, synonymies, comparative notes, distribution, bionomics when available, photographs of the habitus and drawings of the rostra, genitalia and other structures useful for taxonomy are provided. A key including all the groups of Rhinusa as currently considered is provided for the first time. 


Zootaxa ◽  
2007 ◽  
Vol 1496 (1) ◽  
pp. 1-30 ◽  
Author(s):  
RICARDO KAWADA ◽  
CELSO O. AZEVEDO

Neotropical species of Decevania Huben, 2003 are revised for the first time. Species in this genus are characterized by having an 8-segmented flagellum, eye relatively reduced, wings frequently large and floppy with reduced venation (M+CU, 1CUa, 1Cub and 2CU), anterior wing with only one cell enclosed by tubular vein (costal), and hind tarsomere 1–3 posteriorly expanded. Thirteen species are recognized, two of which were previously described, D. parva (Enderlein, 1901) and D. striatigena (Kieffer, 1910), and eleven which are described and illustrated as new; D. brevis sp. nov., D. deansi sp. nov., D. destituta sp. nov., D. elongata sp. nov., D. glabra sp. nov., D. hemisphaerica sp. nov., D. nigra sp. nov., D. nuda sp. nov., D. polita sp. nov., D. reticulata sp. nov., D. unidentata sp. nov. Keys for species identification based on males and females are provided.


Zootaxa ◽  
2010 ◽  
Vol 2609 (1) ◽  
pp. 1 ◽  
Author(s):  
WEICHUN LI ◽  
HOUHUN LI ◽  
MATTHIAS NUSS

Scoparia Haworth, 1811 is revised from China with a total of twenty-six species. Sineudonia Leraut, 1986 syn. n. is synonymized with Scoparia and its type species, Scoparia brunnea (Leraut, 1986) comb. n. is transferred from Sineudonia to this genus. Scoparia isochroalis Hampson, 1907 syn. n. is synonymized with S. congestalis Walker, 1859 and Scoparia kiangensis Leraut, 1986 syn. n. is synonymized with S. spinata Inoue, 1982. Six species are described as new: Scoparia spinosa sp. n., S. jiuzhaiensis sp. n., S. uncinata sp. n., S. brevituba sp. n., S. bifaria sp. n. and S. largispinea sp. n. Scoparia ancipitella (La Harpe, 1855), S. afghanorum Leraut, 1985 and S. utsugii Inoue, 1994 are recorded for the first time in China. Scoparia metaleucalis Hampson, 1907, S. caradjai Leraut, 1986 and S. sinensis Leraut, 1986 are redescribed. The previously unknown male of S. caradjai Leraut, 1986 and the female of S. metaleucalis Hampson, 1907 are described for the first time. Images of wing pattern and genitalia of new and redescribed species are provided. The occurrence of S. basistrigalis Knaggs, 1866 and S. molestalis Inoue, 1982 in China is falsified. The occurrence of S. ambigualis (Treitschke, 1829), S. molestalis Inoue, 1982, S. murificalis Walker, 1859, S. vinotinctalis Hampson, 1896 in China could not be verified. The identification of S. stoetzneri Caradja, 1927 and S. submedinella Caradja, 1927 remains unknown as the type specimens are currently untraceable.


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