The aim of the study was to compare a patch-mosaic pattern in the old-growth forest stands developed in various climate and soil conditions occurring in different regions of Poland. Based on the assumption, that the patch-mosaic pattern in the forest reflect the dynamic processes taking place in it, and that each type of forest ecosystem is characterized by a specific regime of natural disturbances, the following hypotheses were formulated: (i) the patches with a complex structure in stands composed of latesuccessional, shade-tolerant tree species are more common than those composed of early-successional, light-demanding ones, (ii) the patch-mosaic pattern is more heterogeneous in optimal forest site conditions than in extreme ones, (iii) in similar site conditions differentiation of the stand structure in distinguished patches is determined by the successional status of the tree species forming a given patch, (iv) the successional trends leading to changes of species composition foster diversification of the patch structure, (v) differentiation of the stand structure is negatively related to their local basal area, especially in patches with a high level of its accumulation. Among the best-preserved old-growth forest remaining under strict protection in the Polish national parks, nineteen research plots of around 10 ha each were selected. In each plot, a grid (50 × 50 m) of circular sample subplots (with radius 12,62 m) was established. In the sample subplots, species and diameter at breast height of living trees (dbh ≥ 7 cm) were determined. Subsequently, for each sample subplot, several numerical indices were calculated: local basal area (G), dbh structure differentiation index (STR), climax index (CL) and successional index (MS). Statistical tests of Kruskal- Wallis, Levene and Generalized Additive Models (GAM) were used to verify the hypotheses. All examined forests were characterized by a large diversity of stand structure. A particularly high frequency of highly differentiated patches (STR > 0,6) was recorded in the alder swamp forest. The patch mosaic in the examined plots was different – apart from the stands with a strongly pronounced mosaic character (especially subalpine spruce forests), there were also stands with high spatial homogeneity (mainly fir forests). The stand structure in the distinguished patches was generally poorly related to the other studied features. Consequently, all hypotheses were rejected. These results indicate a very complex, mixed pattern of forest natural dynamics regardless of site conditions. In beech forests and lowland multi-species deciduous forests, small-scale disturbances of the gap dynamics type dominate, which are overlapped with less frequent medium-scale disturbances. In more difficult site conditions, large-scale catastrophic disturbances, which occasionally appear in communities formed under the influence of gap dynamics (mainly spruce forests) or cohort dynamics (mainly pine forests), gain importance.
Forest stand structure and coarse woody debris determine the biodiversity of beetle communities in Mediterranean mountain beech forests