Well-combusted duff (<3 cm depth) is generally considered the best seedbed for small-seeded species on upland sites, but we ask here, What is the optimal, postfire residual duff thickness? We hypothesize that a duff thickness equal to (but not greater than) the length of the germinant will offer the best conditions, because at this thickness, the duff layer will not prohibit radicle penetration into the mineral soil, and yet it will serve as a water-conserving mulch. Data from a recent fire in the Rocky mountains of British Columbia were used to show that for three species of Pinus and Picea, (1) duff depths <3 cm were far more clement substrates than thicker duff, and (2) there was a peak in relative survivorship at about 1–2 cm, somewhat shallower than the typical hypocotyl length for these species. Additional data sets from studies previously conducted at boreal and northern cordilleran sites in Alberta, Saskatchewan, Yukon, and Quebec (a combined 21 fires) bolstered these results.