A happy family: systematic revision of the endemic Theridion spiders (Araneae, Theridiidae) of the Hawaiian Islands

The Body ◽

Junior Synonym ◽

Happy Face ◽

Phylogenetic Structure ◽

Systematic Revision ◽

Happy Family ◽

Since the description in 1900 of the iconic Happy Face spider, Theridion grallator, Simon, along with nine relatives, the Theridion fauna of the Hawaiian Islands has remained unstudied. Here, we present a systematic revision of the Hawaiian Theridion, which includes the examination of abundant material collected during the last 50 years, with scanning of the genitalia of several species using SEM techniques, and a cladistic analysis based on 22 morphological characters, to provide a first hypothesis of the phylogenetic structure of the group. We describe eight new species, namely T. ariel, sp. nov., T. caliban, sp. nov., T. ceres, sp. nov., T. ferdinand, sp. nov., T. juno, sp. nov., T. miranda, sp. nov., T. prospero, sp. nov. and T. sycorax, sp. nov. Additionally, we provide new diagnoses for former species and illustrate and describe for the first time the male of T. kauaiense Simon, 1900 and the female of T. praetextum Simon, 1900. We further propose that T. campestratum Simon, 1900 is a junior synonym of T. melinum Simon, 1900 and T. praetextum concolor Simon, 1900 is a junior synonym of T. praetextum. Finally, we provide updated information on the distribution of the species. Most species are easily diagnosed based on the male and female genitalia, but we also reveal the existence of somatic characters that differ among species, such as the body size and the shape and size of the chelicerae, which may have played a role in the diversification and coexistence of some of the species. The preferred cladogram from the cladistic analysis, although compatible with a progression rule, also suggests a complex pattern of multiple back and forward colonisations, albeit most of the clades are poorly supported.

Revision of the southern Andean genus Sadocus Sørensen, 1886 (Opiliones, Gonyleptidae, Pachylinae)

ZooKeys ◽

10.3897/zookeys.1025.57806 ◽

2021 ◽

Vol 1025 ◽

pp. 91-137

Author(s):

Marília Pessoa-Silva ◽

Marcos Ryotaro Hara ◽

Ricardo Pinto-da-Rocha

Keyword(s):

Cladistic Analysis ◽

Junior Synonym ◽

Valid Species ◽

Nominal Species ◽

New Classification ◽

As Species ◽

First Time ◽

Species Inquirendae

Species of the genus Sadocus Sørensen, 1886 are conspicuous gonyleptids that occur in Chile and Argentina. Here, the genus is revised for the first time and the cladistic analysis based on morphological characters does not corroborate its monophyly unless a phylogenetically unrelated species is excluded (explained further on). A new classification is proposed for the seven species left in the genus and considered valid, of the 13 nominal species previously recognized. Two out of the seven valid species are considered as species inquirendae: Sadocus allermayeri (Mello-Leitão, 1945) [= Carampangue allermayeri Mello-Leitão, 1945] and Sadocus nigronotatus (Mello-Leitão, 1943) [= Carampangue nigronotatum Mello-Leitão, 1943]. The following synonymies are proposed: Sadocus bicornis (Gervais, 1849) [original combination = Gonyleptes bicornis Gervais, 1849] is a junior synonym of Sadocus asperatus (Gervais, 1847) [= Gonyleptes asperatus Gervais, 1847]; Sadocus conspicillatus Roewer, 1913, Sadocus exceptionalis (Mello-Leitão, 1946) [= Araucanoleptes exceptionalis Mello-Leitão, 1946] and Sadocus guttatus Sørensen, 1902 are junior synonyms of the valid name Sadocus polyacanthus (Gervais, 1847) [= Gonyleptes polyacanthus Gervais, 1847]; and Sadocus calcar (Roewer, 1913) [= Lycomedes calcar Roewer, 1913] is a junior synonym of the valid name Gonyleptes horridus Kirby, 1819. Sadocus brasiliensis Soares & Soares, 1949 is not congeneric with Argentinean/Chilean species of the genus according to the cladistic analysis and is here synonymized with Discocyrtus catharinensis (Mello-Leitão, 1923 [= Sadocus catharinensis Mello-Leitão, 1923]).

The Problematic Genus Sclerocardius (Coleoptera: Curculionidae: Molytinae: Ithyporini)

Diversity ◽

10.3390/d10030074 ◽

2018 ◽

Vol 10 (3) ◽

pp. 74

Author(s):

Christopher Lyal

Keyword(s):

Junior Synonym ◽

Key To Species ◽

The genus Sclerocardius is revised, using morphological characters. Four species are recognized, including S. africanus (Boheman), S. bohemani Schoenherr stat.rev., S. indicus Hartmann and S. kuscheli sp.nov. The species Sclerocardius madecassus Ferragu is synonymized with S. bohemani syn.nov., and Charactocnemus hintzi Hartmann is treated as a junior synonym of S. bohemani, not S. africanus. A key to species is given. Lectotypes are designated for Heteramphus africanus Boheman and Sclerocardius africanus Schoenherr. A female elytro-tergal stridulatory system involving the modification of the wing-binding patch of the seventh tergite is reported for the Sclerocardiina for the first time and supports the inclusion of the subtribe within the Ithyporini.

A new species of Zoniopoda Stål (Orthoptera: Acridoidea: Romaleidae) from Argentina and its phylogenetic position within the genus

Zootaxa ◽

10.11646/zootaxa.2913.1.3 ◽

2011 ◽

Vol 2913 (1) ◽

pp. 27 ◽

Cited By ~ 1

Author(s):

MARTINA E. POCCO ◽

GONZALO D. RUBIO ◽

M. MARTA CIGLIANO

Keyword(s):

New Species ◽

Cladistic Analysis ◽

Type Specimen ◽

Color Pattern ◽

Species Group ◽

The Body ◽

Phylogenetic Position ◽

Median Carina ◽

A New Species

A new species of the romaleid grasshopper genus Zoniopoda Stål (Romaleidae: Romaleini) is described and illustrated from the Sierras Chicas of Córdoba Province, central Argentina. A cladistic analysis based on morphological characters indicates that the genus Zoniopoda constitutes a monophyletic group and that Zoniopoda serrana n. sp. must be assigned to Iheringi species group based on synapomorphies of the pronotum and body color. The new species is similar to Z. similis Bruner from Paraguay, Bolivia and Brazil, from which it can be distinguished by the color pattern of the body, shape of the pronotal dorso-median carina and characters of the male terminalia and epiphallus. This paper has been formatted with embedded links to images of the type specimen, maps based on georeferenced specimen data for the genus and an updated key to the species of Zoniopoda available on the Orthoptera Species File (OSF) online (http://orthoptera.speciesfile.org).

Revision of Hypoxys Amyot & Serville, 1843 stat. rest. (Heteroptera Pentatomidae)

Zootaxa ◽

10.11646/zootaxa.4742.3.1 ◽

2020 ◽

Vol 4742 (3) ◽

pp. 401-441

Author(s):

BENEDITO MENDES NUNES ◽

LOURIVAL DIAS CAMPOS ◽

MARIA THAYANE DA SILVA MENDONCA ◽

EDUARDO VICTOR DE PAIVA CUNHA ◽

JOSE ANTONIO MARIN FERNANDES

Keyword(s):

Type Species ◽

Proximal Part ◽

Junior Synonym ◽

Similar Species ◽

Costal Margin ◽

Distribution Maps ◽

Species Groups ◽

Heteroptera Pentatomidae ◽

Hypoxys is one of four subgenera of Edessa (Fabricius, 1787) together with Aceratodes (De Geer, 1773), Dorypleura (Lepeletier & Serville, 1825) and Edessa (Fabricius, 1787). This taxon has a very confusing taxonomy due to incorrect use of the names available, imprecise descriptions, and large number of very similar species. To illustrate the confusion, Hypoxys potentially includes a large number of species of Edessa, but actually contains only the type species, according to the most recent catalogue. The name E. quadridens Fabricius, 1803, type species of Hypoxys, was used for more than a century as a label to identify a large group of similar species. In this work we continue revising Edessa elevating Hypoxys to genus based on the following morphological characters: proximal part of costal margin black; evaporatorium with a deep notch on lateral margin; anterior arms of metasternal process acuminate; pygophore longer than wide and subrectangular or barrel-shaped in dorsal view; dorsal rim of pygophore well-projected, almost reaching posterolateral angles and finishing in a small lobe; and posterolateral angles of pygophore not developed. Hypoxys is being divided in four species groups to include 17 species transferred from Edessa. H. quadridens group comprises: Hypoxys quadridens (Fabricius, 1803) (type species), H. boerneri (Breddin, 1904) comb. nov., H. claricolor (Breddin, 1907) comb. nov., H. dolosus (Breddin, 1907) comb. nov., H. eburatulus (Breddin, 1907) comb. nov., H. necopinatus (Breddin, 1907) comb. nov., H. offuscatus (Breddin, 1907) comb. nov. and H. trabeculus (Breddin, 1907) comb. nov. H. triangularis group comprises: H. triangularis (Dallas, 1851) comb. nov. (type species), H. capito (Breddin, 1904) comb. nov. and H. subrastratus (Bergroth, 1891) comb. nov. H. oxyacanthus group comprises: H. oxyacanthus (Breddin, 1904) comb. nov. (type species), H. brachyacanthus (Breddin, 1904) comb. nov., H. infulatus (Breddin, 1904) comb. nov., H. leptacanthus (Breddin, 1904) comb. nov. and H. tragelaphus (Breddin, 1903) comb. nov. H. balteatus group comprises only H. balteatus (Walker, 1868) comb. nov. These groups of species will be useful to organize the new species of Hypoxys that will be described in upcoming papers. Descriptions, measurements, photos of genitalia of both sexes; photos in dorsal and ventral views of the species; and distribution maps are provided. Males of H. capito, H. claricolor, H. eburatulus, H. infulatus, and H. subrastratus are described for the first time. Female of H. necopinatus is described for the first time. Edessa jugalis is considered a junior synonym of H. quadridens; E. rimata a junior synonym of H. offuscatus; E. scabriventris and E. leprosula junior synonyms of H. triangularis; E. pachyacantha a junior synonym of H. tragelaphus; E. orba a junior synonym of E. oxyacanthus. A key to the species of Hypoxys is also provided.

Taxonomic revision and preliminary phylogenetic analysis of the subgenus Megalostomis (Scaphigenia) Lacordaire (Coleoptera: Chrysomelidae)

Insect Systematics & Evolution ◽

10.1163/187631207788754402 ◽

2007 ◽

Vol 38 (3) ◽

pp. 335-359 ◽

Cited By ~ 2

Author(s):

Sergio Roig-Juñent ◽

Martha Domínguez ◽

Federico Agrain

Keyword(s):

Phylogenetic Analysis ◽

Arid Regions ◽

Cladistic Analysis ◽

Taxonomic Revision ◽

Distribution Data ◽

Species Status ◽

Systematic Revision ◽

Male And Female ◽

Preliminary Phylogenetic Analysis

AbstractThe subgenus M. (Scaphigenia) Lacordaire includes six species distributed in arid regions of South America. A systematic revision of this subgenus is presented, including re-descriptions and an update of distribution data of the six species. A preliminary cladistic analysis is performed to test the relationships among the species of the subgenus and if the morphological characters used are suitable. A key is presented to separate the seven subgenera of Megalostomis Chevrolat as well as a key for the species of the subgenus M. (Scaphigenia). Male aedeagus internal sac of the nine studied taxa are described and illustrated. This constitutes the first internal sac descriptions for members of this subtribe and could help clarify the homology of such structures within Cryptocephalinae. M. (S) consimilis Achard is reassigned to the species status on the base of genitalic characters. The cladistic analysis was done using forty-one characters from adult external morphology and male and female genitalia. Two species of two different subgenera of Megalostomis: M. (Megalostomis), and M. (Heterostomis) Lacordaire, and one representative of the genus Themesia Lacordaire were selected as out groups. Results were obtained by implicit enumeration using parsimony software. Three equally parsimonious trees were obtained of 45 steps, Ri=0.952, and Ci= 0.941. Support of each group was evaluated by Jacknifing, Bootstrap and Bremer values. Relationships between species are discussed.

Systematic revision and phylogeny of the Australian myrmecomorphic seed bug genus Daerlac Signoret (Insecta:Heteroptera:Rhyparochromidae:Udeocorini)

Invertebrate Systematics ◽

10.1071/is11030 ◽

2012 ◽

Vol 26 (1) ◽

pp. 41 ◽

Cited By ~ 2

Author(s):

Gerasimos Cassis ◽

Celia Symonds

Keyword(s):

Leaf Litter ◽

Cladistic Analysis ◽

Key To Species ◽

Systematic Revision ◽

Seed Predators ◽

Taxonomic Key ◽

Open Forest ◽

High Degree ◽

Degree Of Overlap

The Australian endemic ant-mimetic seed bug genus Daerlac is revised. This paper provides a redescription of the genus Daerlac and four species: D. apicalis, D. cephalotes, D. nigricans and D. picturatus. Daerlac tricolor is synonomised with D. cephalotes. A taxonomic key to species is provided. Known distributions of D. apicalis, D. nigricans and D. picturatus are each extended beyond previously known ranges. Daerlac species are found predominantly in temperate open forest and woodlands in association with ants and eucalypts. All species are broadly distributed and there is a high degree of overlap in distributions. They are seed predators found on the ground, in leaf litter, under bark or on trunks of eucalypts, and putatively forage on post-dispersed seeds. Cladistic analysis of morphological characters finds that the four species of Daerlac form two well-supported sister-groups (D. apicalis + D. picturatus, and D. cephalotes + D. nigricans). A discussion of the distribution, biology and myrmecomorphy of the genus is provided, and the tribal placement of Daerlac and its relationship to Laryngodus are discussed.

Patagonian Eocene Archaeopithecidae Ameghino, 1897 (Notoungulata): systematic revision, phylogeny and biostratigraphy

Journal of Paleontology ◽

10.1017/jpa.2017.53 ◽

2017 ◽

Vol 91 (6) ◽

pp. 1272-1295 ◽

Cited By ~ 4

Author(s):

Bárbara Vera

Keyword(s):

Middle Eocene ◽

Mass Range ◽

Late Eocene ◽

The Body ◽

South American ◽

Type Specimens ◽

Systematic Revision ◽

Size And Morphology ◽

Land Mammal ◽

AbstractThe Archaeopithecidae is a very poorly known group of native ungulates from the Eocene of Patagonia (Argentina), whose alpha taxonomy has remained obscure since Ameghino’s times. It is traditionally considered as a family representative of the Casamayoran (middle Eocene) South American Land Mammal Age, and is thought to be morphologically close to the notopithecids. After studying >200 specimens from several institutions, including all the type specimens, a taxonomic overestimation is established. Out of the six species considered originally as archaeopithecids, Archaeopithecus rogeri Ameghino, 1897 is here recognized as the only valid name and species; subsequent synonymies are proposed and previous taxonomic hypotheses discarded. This exhaustive revision has permitted improving the knowledge of A. rogeri and, for the first time, it has revealed many craniodental characters, which allow amending its diagnosis and differentiating this taxon from other Eocene notoungulates. Archaeopithecus rogeri is a small-sized taxon characterized by its complete and rooted dentition, which is relatively higher than that of other contemporaneous short-crowned notoungulates and shows ontogenetic variation in size and morphology. The body mass range of A. rogeri (1.4–2.5 kg) is comparable to those of notopithecids and some small hegetotheriids. The phylogenetic analysis shows A. rogeri is not directly related to any family within Notoungulata, appearing into a polytomy, as a basal taxon of typotherians. The biochronological range of A. rogeri is adjusted to Vacan (middle Eocene) through Barrancan subages (late middle Eocene); older (Riochican, late early Eocene) and younger (Mustersan, late Eocene) records remain to be confirmed.

Description of Ossinissa, a new pholcid genus from the Canary Islands (Araneae: Pholcidae)

Zootaxa ◽

10.11646/zootaxa.982.1.1 ◽

2005 ◽

Vol 982 (1) ◽

pp. 1 ◽

Cited By ~ 3

Author(s):

DIMITAR DIMITROV ◽

CARLES RIBERA

Keyword(s):

Canary Islands ◽

Type Species ◽

New Genus ◽

Cladistic Analysis ◽

New Combination ◽

Ossinissa new genus (Araneae, Pholcidae) is described to place a Canarian pholcid species formerly considered belonging to Spermophorides. The male of the type species, Ossinissa justoi (Wunderlich) new combination, is described for the first time and the female is re-described. This new genus is supported by a revision of the morphological characters of the female, the newly discovered male, and a cladistic analysis.

Taxonomic notes on owlflies from Pakistan (Neuroptera: Myrmeleontidae: Ascalaphinae)

Zootaxa ◽

10.11646/zootaxa.4970.3.1 ◽

2021 ◽

Vol 4970 (3) ◽

pp. 401-452

Author(s):

MUHAMMAD ASGHAR HASSAN ◽

XINGYUE LIU

Keyword(s):

Geographical Distribution ◽

Male Genitalia ◽

New Records ◽

Junior Synonym ◽

Identification Key ◽

Distribution Maps ◽

The Family ◽

Taxonomic notes are presented on the former Ascalaphidae (owlflies), now subfamily Ascalaphinae of the family Myrmeleontidae from Pakistan. An updated checklist of new records is provided that accounts for all known 15 genera and 22 species from Pakistan, excluding the species of the tribe Palparini. Geographical distribution maps and an updated identification key to all known genera and species from Pakistan are also given. We synonymized two monotypic genera, Horischema Mészáros & Ábrahám, 2003 and Perissoschema Mészáros & Ábrahám, 2003 both as junior synonyms of Ogcogaster Westwood, 1847 based on very similar male genitalia and external morphological characters. Moreover, Perissoschema evae Mészáros & Ábrahám, 2003 is treated as a junior synonym of Ogcogaster segmentator Westwood, 1847. We recorded three genera for the first time from Pakistan, i.e., Acheron Lefèbvre, 1842, Glyptobasis McLachlan, 1873, and Suphalomitus van der Weele, 1909. To more completely resolve the generic or specific status of those poorly known species from Pakistan, broader sampling throughout the country is needed.

Phylogeny of the Limnophilinae (Limoniidae) and early evolution of the Tipulomorpha (Diptera)

Invertebrate Systematics ◽

10.1071/is08017 ◽

2008 ◽

Vol 22 (6) ◽

pp. 627 ◽

Cited By ~ 24

Author(s):

Guilherme Cunha Ribeiro

Keyword(s):

Adult Male ◽

Male Genitalia ◽

Cladistic Analysis ◽

Sister Group ◽

Systematic Position ◽

Sensu Stricto ◽

Crane Flies ◽

Male Morphology ◽

Tipulomorpha (crane flies) comprise one of the largest subgroups of Diptera, but its phylogeny at different levels has been poorly explored. This study presents the most comprehensive cladistic analysis of the group ever made, with emphasis on the genera and subgenera of the subfamily Limnophilinae (Limoniidae), assumed to include some of the earliest lineages of Tipulomorpha sensu stricto and therefore important for the understanding of the early patterns in the evolution of the crane flies. Eighty-eight characters of the male imago were scored for 104 exemplar species. The most parsimonious trees were searched using implied weighting, in the framework of a sensitivity analysis with different values of k (2 to 6). The dataset based on the characters of adult male morphology showed high levels of homoplasy and yielded very incongruent and unstable phylogenetic results, which are very sensitive to changes in analytical parameters. In the preferred and most parsimonious phylogenetic hypothesis, the Pediciidae is the sister-group of all other Tipulomorpha sensu stricto. The results indicate the paraphyly of the Limoniidae with respect to the Cylindrotomidae and Tipulidae, which are considered sister-groups. The Limoniidae subfamilies Limnophilinae, Limoniinae and Chioneinae are considered non-monophyletic. The study allowed a reconstruction of the possible ground plan condition of selected features of the adult male morphology of crane flies. The genera/subgenera Epiphragma (Epiphragma), Acantholimnophila, Shannonomyia, Limnophila (Arctolimnophila), Eloeophila, Conosia, Polymera, Polymera (Polymerodes), Prionolabis, Eutonia, Phylidorea (Phylidorea), Metalimnophila, Gynoplistia (Cerozodia), Gynoplistia (Dirhipis), Nothophila, Pseudolimnophila (Pseudolimnophila), Pilaria and Ulomorpha are considered monophyletic, but in general are defined by combinations of very homoplastic character states. Two Temperate Gondwanan clades, (Tonnoirella + (Edwardsomyia + (Tinemyia + (Rhamphophila + (Nothophila))))) and ((Notholimnophila + Bergrothomyia) + (Mesolimnophila + (Chilelimnophila + Ctenolimnophila))) are recovered. The genera Limnophila, Neolimnomyia, Gynoplistia (sensu lato) and Hexatoma (sensu lato) are considered non-monophyletic. The systematic position and some morphological characters of ‘problematic’ taxa, such as Dactylolabis, Elephantomyia, Helius and Atarba are discussed on the light of the proposed phylogeny and the analysis of the characters. Character states are richly illustrated. A detailed study of the morphology of the male genitalia is made, and several genera and species have the morphology of the male genitalia illustrated for the first time.