Total-evidence phylogeny of the Rhinomirini, taxonomic review of its subgroupings (Insecta: Heteroptera: Miridae: Cylapinae) and description of new Australian taxa

2019 ◽  
Vol 187 (4) ◽  
pp. 1196-1252 ◽  
Author(s):  
Anna A Namyatova ◽  
Gerasimos Cassis

Abstract Rhinomirini (Insecta: Heteroptera: Miridae: Cylapinae) is a heterogeneous tribe of uncertain systematic position. We provide a phylogenetic analysis of this tribe based on morphological characters and molecular markers. Monophyly of the tribe was not supported by our findings. The Rhinocylapus complex and Rhinomiriella complex are transferred to Fulviini. Rhinomirini are restricted to the Rhinomiris complex. The record of Rhinomirini in Australia is considered dubious. The genera Punctifulvius and Yamatofulvus are transferred to the Rhinocylapus complex. The Rhinocylapus complex is recorded from Australia for the first time, with four new species: Mycetocylapus alexeyi sp. nov., Punctifulvius austellus sp. nov., Punctifulvius aquilonius sp. nov. and Rhinocylapoides valentinae sp. nov. The Philippine species Mycetocylapus major is synonymized with Mycetocylapus minor. The Australian species Rhinomiriella tuberculata is redescribed, and females of this species are described for the first time. Rhinomiriella aidani sp. nov. is described as new to science. The female genitalia of Proamblia elongata, Rhinocylapidius velocipidoides and Rhinomiris conspersus and the male genitalia of Rhinomiris conspersus and Rhinomiridius aethiopicus are illustrated and discussed. Sexual dimorphism in the Rhinocylapus complex and wing modifications in Cylapinae are discussed.

Zootaxa ◽  
2012 ◽  
Vol 3413 (1) ◽  
pp. 44 ◽  
Author(s):  
TOMOHIDE YASUNAGA ◽  
RAM KESHARI DUWAL ◽  
MICHAEL D. SCHWARTZ

The mirine plant bug genus Neolygus Knight is reported from Nepal and North India for the first time, thus representing the first confident distributional record of this genus from the Himalayas. A new species, Neolygus machanensis, is described from Nepal. Two known members, N. bui Lu & Zheng and N. keltoni (Lu & Zheng), newly recognized as occurring in Nepal and/or North India, are diagnosed. Habitus images of live individuals and illustrations of the male genitalia are provided for all treated species. A checklist of the Old World species, and discussion of the systematic position and zoogeography of Neolygus are included. Neolygus partitus (Walker, 1873) [Capsus], N. mjohjangsanicus (Josifov, 1992) [Lygocoris], and Neolygus zebei (Günther, 1997) [Lygocoris] are proposed as new combinations.


1981 ◽  
Vol 29 (2) ◽  
pp. 233 ◽  
Author(s):  
M Horak ◽  
W Sauter

The male genitalia of the genus Metaselena Diak. are described and illustrated for the first time, and as a result of that the genus is transferred to the Olethreutinae. Eight new species of Metaselena - lepta, rhabdota, symphylos, pemphigodes, pithana, platyptera, diakonoffi and allophlebodes - are described and illustrated. The systematic position of Metaselena, so far known only from Australia and New Guinea, is briefly discussed and the close association of the genus with Eucosmomorpha Obr. is pointed out.


Zootaxa ◽  
2010 ◽  
Vol 2516 (1) ◽  
pp. 1 ◽  
Author(s):  
ANTOINE MANTILLERI

The genera Hoplopisthius and Carcinopisthius are reviewed. All the species are redescribed and illustrated and an updated identification key and distributional maps are provided. One new species from New Guinea is described (Hoplopisthius maximus n. sp.) and two new synonymies are proposed: Hoplopisthius celebensis Kolbe, 1892 = H. trichemerus Senna, 1892, n. syn. and Carcinopisthius lamingtoni Damoiseau, 1987 = C. forcipitiger Damoiseau, 1987, n. syn. Phylogenetic analysis using PAUP (maximum parsimony) was performed using 25 morphological characters of adults. This analysis shows the group Hoplopisthius + Carcinopisthius is monophyletic, but Carcinopisthius alone is paraphyletic. Nomenclatural changes at the generic level are made to reconcile nomenclature and phylogeny: Hoplopisthius is preserved; Carcinopisthius is downgraded to the rank of subgenus for the two oriental species H. oberthueri and H. fruhstorferi; and Pseudotaphroderes is resurrected as a third subgenus and includes all New-Guinean and Australian species.


Zootaxa ◽  
2008 ◽  
Vol 1692 (1) ◽  
pp. 43 ◽  
Author(s):  
MURRAY J. FLETCHER ◽  
JAMES N. ZAHNISER

Deltocephalus centralis Evans is transfered from Arawa Knight (Tribe Athysanini) to Goniagnathus Fieber (Tribe Goniagnathini) as Goniagnathus centralis (Evans) comb. nov. and the male genitalia described for the first time. Four new Australian species of Goniagnathus are described: G. osmelaki Fletcher, G. venenensis Fletcher, G. crocodontis Fletcher and G. pectinatus Fletcher. This represents the first record of the tribe Goniagnathini in Australia. A key for the identification of the Australian species is provided.


Zootaxa ◽  
2020 ◽  
Vol 4766 (2) ◽  
pp. 261-305
Author(s):  
DUNIESKY RÍOS-TAMAYO ◽  
ROBIN LYLE

After more than a century, the genus Lepthercus Purcell, 1902 is revised. Lepthercus dregei Purcell, 1902 and L. rattrayi Hewitt, 1917 are redescribed; with the female of L. dregei described for the first time. Nine new species of Lepthercus are described. A phylogenetic analysis with morphological characters using implied weights and parsimony as optimality criteria, suggests the separation of the genus in two clades. The first clade is formed by L. dippenaarae sp. nov., L. engelbrechti sp. nov., L. haddadi sp. nov., L. rattrayi Hewitt, 1917 and L. sofiae sp. nov., here denominated “Group haddadi”. The species L. confusus sp. nov., Lepthercus dregei Purcell, 1902, Lepthercus filmeri sp. nov., Lepthercus kwazuluensis sp. nov., Lepthercus lawrencei sp. nov. and Lepthercus mandelai sp. nov., form the second clade, here denominated “Group dregei”. The “Group haddadi” is characterized by males with a curved metatarsus I, and a swollen tibia I. The “Group dregei” is supported by the presence of small maxillary cuspules in males. A new diagnosis is provided for Lepthercus as well as an identification key for all species of the genus. New distribution maps for the genus in the country are also presented. 


Zootaxa ◽  
2012 ◽  
Vol 3185 (1) ◽  
pp. 1
Author(s):  
ROMAN LOHAJ ◽  
BORISLAV GUÉORGUIEV ◽  
GÉRARD DUBAULT ◽  
BERNARD LASSALLE

The species belonging to the subgenus Molopsis Schatzmayr, 1943 of genus Tapinopterus Schaum, 1858 are revised. Thestudy is based on 372 specimens and includes, for each taxon, diagnosis, description (only for the new species), references,new distributional records and illustrations. Morphological characters of the male and female genitalia are widely used todelimit the separate species, the significance of some non-gonapophyseal genital structures of the females for the system-atics of Molopsis is explicitly emphasized. For the first time, the male genitalia of T. molopiformis, T. molopinus, T. wie-demanni, and T. rebellis, including also newly described species, as well as female genitalia of Molopsis are described andillustrated. All available type material is listed and represented by habitus photographs.As a result, 9 monotypic species of Molopsis are recognized. Tapinopterus rebellis (Chaudoir, 1868) and T. rebelliskumanensis (Reitter, 1884), formerly assigned to Molopsis are recognized as outgroup-taxa. The following new speciesare described: T. (Molopsis) aenigmaticus sp.nov. (Asian Turkey, „Armenia“ imprecise locality), T. (Molopsis) chaudoirisp.nov. (Asian Turkey, Bursa Villayet, Uludağ Mt., Sakarya (Adapazari) Villayet, Gökdağ Mt.), T. (Molopsis) oyukluensissp.nov. (Asian Turkey, Konya Villayet, Fasihan Pass), and T. (Molopsis) relegatus sp.nov. (Asian Turkey, Bolu Villayet,Mengen env., Dorukhan Pass). The following nomenclatural acts are also proposed: Tapinopterus (Molopsis) molopinus (Chaudoir, 1868) = Tapinopterus (Molopsis) dipojranus brussanus Straneo, 1984, syn.nov.; Tapinopterus (Molopsis)


2008 ◽  
Vol 22 (6) ◽  
pp. 627 ◽  
Author(s):  
Guilherme Cunha Ribeiro

Tipulomorpha (crane flies) comprise one of the largest subgroups of Diptera, but its phylogeny at different levels has been poorly explored. This study presents the most comprehensive cladistic analysis of the group ever made, with emphasis on the genera and subgenera of the subfamily Limnophilinae (Limoniidae), assumed to include some of the earliest lineages of Tipulomorpha sensu stricto and therefore important for the understanding of the early patterns in the evolution of the crane flies. Eighty-eight characters of the male imago were scored for 104 exemplar species. The most parsimonious trees were searched using implied weighting, in the framework of a sensitivity analysis with different values of k (2 to 6). The dataset based on the characters of adult male morphology showed high levels of homoplasy and yielded very incongruent and unstable phylogenetic results, which are very sensitive to changes in analytical parameters. In the preferred and most parsimonious phylogenetic hypothesis, the Pediciidae is the sister-group of all other Tipulomorpha sensu stricto. The results indicate the paraphyly of the Limoniidae with respect to the Cylindrotomidae and Tipulidae, which are considered sister-groups. The Limoniidae subfamilies Limnophilinae, Limoniinae and Chioneinae are considered non-monophyletic. The study allowed a reconstruction of the possible ground plan condition of selected features of the adult male morphology of crane flies. The genera/subgenera Epiphragma (Epiphragma), Acantholimnophila, Shannonomyia, Limnophila (Arctolimnophila), Eloeophila, Conosia, Polymera, Polymera (Polymerodes), Prionolabis, Eutonia, Phylidorea (Phylidorea), Metalimnophila, Gynoplistia (Cerozodia), Gynoplistia (Dirhipis), Nothophila, Pseudolimnophila (Pseudolimnophila), Pilaria and Ulomorpha are considered monophyletic, but in general are defined by combinations of very homoplastic character states. Two Temperate Gondwanan clades, (Tonnoirella + (Edwardsomyia + (Tinemyia + (Rhamphophila + (Nothophila))))) and ((Notholimnophila + Bergrothomyia) + (Mesolimnophila + (Chilelimnophila + Ctenolimnophila))) are recovered. The genera Limnophila, Neolimnomyia, Gynoplistia (sensu lato) and Hexatoma (sensu lato) are considered non-monophyletic. The systematic position and some morphological characters of ‘problematic’ taxa, such as Dactylolabis, Elephantomyia, Helius and Atarba are discussed on the light of the proposed phylogeny and the analysis of the characters. Character states are richly illustrated. A detailed study of the morphology of the male genitalia is made, and several genera and species have the morphology of the male genitalia illustrated for the first time.


Zootaxa ◽  
2012 ◽  
Vol 3281 (1) ◽  
pp. 56
Author(s):  
HAILIN YANG ◽  
LIANGMING CAO ◽  
WAIZHI CAI

The genus Paralenaeus Reuter, 1881, is recorded for the first time from China. The second species of the genus, Parale-naeus schaeferi sp. n., is described from Yunnan. Dorsal habitus, male genitalia, and other diagnostic morphological characters of the new species are illustrated.


Insects ◽  
2019 ◽  
Vol 10 (8) ◽  
pp. 231 ◽  
Author(s):  
Robin Kundrata ◽  
Alexander S. Prosvirov ◽  
Dominik Vondracek ◽  
Eliska Sormova

Senodoniini is a small lineage of click beetles currently comprising 21 species in two genera, distributed in the Himalayas and East and Southeast Asia. The definition and limits of this group have changed considerably during its history. Recent authors treat Senodoniini as a tribe within Dendrometrinae, usually close to Dimini, but this placement has never been rigorously tested. Here, we shed new light on the systematic position and limits of Senodoniini by performing a combined phylogenetic analysis of two nuclear and two mitochondrial molecular markers. Our results recovered Senodoniini not monophyletic, and placed them into the Lissominae complex, where they formed a clade with Austrelater Calder & Lawrence (Protelaterini). Molecular phylogeny is in agreement with the adult morphology. Additionally, we examined the morphology of a monotypic genus Rostricephalus Fleutiaux from Southeast Asia, which was previously classified in various Elateridae groups including Senodoniini, and its position was always uncertain. This genus shares morphological characters with Protelaterini. We provide morphological redescriptions as well as the figures of main diagnostic characters for Senodonia Laporte, Sossor Candèze, and Rostricephalus. Based on our results, we place these genera to Lissominae: Protelaterini, and hence synonymize Senodoniini Schenkling with Protelaterini Schwarz.


Author(s):  
Shingo Hosoishi ◽  
Kazuo Ogata

Two distinct new species of the ant genus Crematogaster, C. khmerensis sp. nov. and C. pfeifferi sp. nov., are described from Cambodia and Malaysia, respectively. The two species are unique among Asian Crematogaster in that they have vertically directed propodeal spines, but their systematic positions have not been determined based on morphological characters alone. Molecular phylogenetic analysis of 89 Crematogaster taxon matrices previously published plus C. khmerensis sp. nov., using nuclear genes, reveals that C. khmerensis sp. nov. is nested within the Australo-Asian Crematogaster clade. Morphological assignment of the developed pronotal shoulders implies a close relationship between C. khmerensis sp. nov. and the C. tetracantha-group. Based on molecular and morphological evidence, we erect a new species group, C. khmerensis-group, to contain C. khmerensis sp. nov. and C. pfeifferi sp. nov. Divergence time estimates using MCMCTree shows that the root node of the C. khmerensis sp. nov. terminal is estimated to be of middle Miocene age at 15 million years old. The position of the C. khmerensis-group well supports the Oriental- to Australian-region dispersal history that has been proposed for the Australo-Asian Crematogaster clade.


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