The Douglas-fir forests of Banff and Jasper National Parks, Canada
Twenty stands of uniform, mature, undisturbed Pseudotsuga menziesii var. glauca forest were selected in Banff and Jasper National Parks, using a combination of airphoto and ground surveys. Quantitative sampling methods were used to describe their floristic composition, vegetation structure, and physical habitats.Scattered trees of Pinus contorta, Picea glauca, and Populus tremuloides occurred in the Pseudotsuga forests. The poorly developed shrub strata contained Shepherdia canadensis, Rosa acicularis, Spiraea lucida, Juniperus communis. Major herb, dwarf shrub, bryophyte, and lichen species were Elymus innovatus, Calamagrostis rubescens, Fragaria virginiana, Astragalus decumbens, Aster conspicuus; Arctostaphylos uva-ursi, Linnaea borealis; Hylocomium splendens, Abietinella abietina, Tortula ruralis; Peltigera canina. A cyclical pattern of vegetation change is related to the death, replacement, and maturation of Pseudotsuga trees.Relations between vegetation and physical habitat were analyzed by simple correlation and multiple regression. Available potassium in soil, stand age, and slope exposure were most influential in regression equations for tree stratum development. Understory strata were primarily correlated with tree stratum development.A two-dimensional stand ordination based on the vegetational similarity of subordinate vascular strata showed a pronounced separation of Banff and Jasper Pseudotsuga forests, and prompted recognition of two new climax associations (sensu Daubenmire): Pseudotsuga / Elymus innovatus in Jasper; Pseudotsuga / Calamagrostis rubescens – Elymus innovatus in Banff. Habitat differences between the two associations include macroclimate, parent materials, and soil moisture.