scholarly journals PINE FORESTS OF THE CLASS ERICO–PINETEA HORVAT 1959 FROM DAGESTAN (NORTHERN CAUCASUS)

Ekosistemy ◽  
2020 ◽  
pp. 27-42
Author(s):  
N. B. Ermakov ◽  
Z. A. Abdurakhmanova ◽  
Yu. V. Plugatar

The paper represents the results of grass pine forests classification in Dagestan by the Brown-Blanquet method. A number of 68 releves of pine forests (taken from the paper by Abdurakhmanova et al., 2018) were included in two higher categories of vegetation – the boreal forests of the class Vaccinio–Piceetea Br.-Bl. in Br.-Bl. et al. 1939 and the sub-Mediterranean thermophilous forests of the class Erico–Pinetea Horvat 1959 based on the quantitative classification. Comparative syntaxonomic analysis of the Caucasian communities of the Erico–Pinetea class revealed their high level of floristic originality in comparison with similar forests of southern Europe, Western Asia and Crimea. This corresponds to the unique bioclimatic conditions of Dagestan located on the territory with a clear effect of «rain shadow» from the high mountain ranges of the Greater Caucasus stretching on the way of the western Atlantic moist air masses that result in the high climate continentality. These essential ecological and floristic peculiarities of the grass pine forests of the Eastern Caucasus provided a basis for the description of a new order — Alchemillo sericaceae–Pinetalia sylvestris ord. nova hoc loco representing the extreme eastern part of the Erico–Pinetea class range. The diagnostic species group of the order includes dominantly Eastern-Euxinian, Caucasian endemics and Eurasian xerophilous species: Juniperus oblonga, Alchemilla sericata, Galium valantioides, Peucedanum ruthenicum, Thalictrum foetidum, Rosa elasmacantha, Filipendula vulgaris, Koeleria cristata, as well as diagnostic species of subordinated alliances. The order includes two alliances demonstrating the ecological and floristic differences of Dagestan pine forests due to the thermic factor. The alliance Bupleuro polyphylli–Pinion sylvestris all. nova hoc loco (holotypus of the order) includes grass pine forests with predominance of thermophilous floristic elements. Differential species of the alliance are Astrantia major, Valeriana alliariifolia, Polygonatum verticillatum, Pyrethrum coccineum, Psephellus daghestanicus, Galium rubioides, Calamagrostis arundinacea, Rubus saxatilis, Brachypodium pinnatum, Fragaria vesca, Fragaria viridis, Primula macrocalyx, Anthriscus sylvestris, Amoria ambigua, Bupleurum polyphyllum, Hylocomium splendens, Pleurozium schreberi, Dicranum scoparium, Rhytidiadelphus triquetrus, Rhodobryum roseum, Climacium dendroides, Abietinella abietina, Plagiomnium cuspidatum, Betula litwinowii, B. pendula. Three associations and two non-rank communities were included in the alliance (Carici albae–Pinetum sylvestris Ermakov, Abdurakhmanova, Potapenko 2019 — holotypus of the alliance, Ranunculo caucasicae–Pinetum sylvestris ass. nova hoc loco, Viburno lanatae–Pinetum sylvestris Ermakov, Abdurakhmanova, Potapenko 2019, community Stachys atherocalyx–Pinus sylvestris, community Centaurea phrygia–Pinus sylvestris). The alliance Onobrichido cornutae–Pinion sylvestris all. nova hoc loco includes unique cryophilous pine forests with predominance of the cushion plants in the gound layer and numerous constant species of Caucasian cryophilous and xerophilous endemics. Diagnostic species of the alliance are Onobrychis cornuta, Festuca woronowii, Gypsophila tenuifolia, Salvia canescens, Rosa elasmacantha, Androsace koso-poljanskii, Galium brachyphyllum, Thymus daghestanicus, Asperula alpina, Anthemis dumetorum, Scutellaria oreophila, Satureja subdentata, Potentilla recta, Teucrium polium, Euphorbia virgate, Scabiosa gumbetica, Astragalus alexandri, A. fissuralis, Seseli alexeenkoi, Pseudomuscari pallens, Rhamnus pallasii, Vincetoxicum funebre, Jurinea ruprechtii, Helianthemum dagestanicum, Stipa caucasica, Centaurea edmondii, Selaginella helvetica, Scorzonera filifolia, Dracocephalum austriacum, Viola somchetica, Oxytropis lanata. At present the alliance is represented by a single association – the Onobrichido cornutae–Pinetum sylvestris ass. nova hoc loco. The results of the classification have demonstrated a significant level of originality of the Caucasian grass pine forests and a high floristic status of the Caucasus as a one of the important centers of modern phytodiversity.

2003 ◽  
pp. 28-40 ◽  
Author(s):  
L. L. Zanokha

The 2 new associations, Carici stantis—Salicetum reptantis and Salico-Polemonietum acutiflori, are described within the all. Caricionstantis of the class Scheuchzerio-Caricetea fuscae R. Tx. 1937 for the northern belt of the typical tundra subzone of Taymyr. The diagnostic species group of the first syntaxon includes plants typical of sites with excessive watering. The ass. Carici stantis—Salicetum reptantis is restricted to bottoms or lower parts of the watershed and moun­tain terrace slopes. The ecological regime of these sites is subject to slight variations reflected in presence/ absence of certain species. Due to floristic differences, the association is subdivided into several subassociations which form a topographic-ecological series along the humidity gradient: epilobietosum palustris→ptilidietosum ciliaris→typicum→petasitetosum frigidi. The ass. Carici stantis—Salicetum reptantis is referred to the all. Caricionstantis which also includes the associations Meesiotriquetris—Caricetumstantis and Pooarcticae—Dupontietumfisheri (also described from the Ragozinka R. basin); the latter occur in the wet depres­sions between sloping hills and the flowing valleys, respectively. If compared to these two, the ass. Carici stantis—Salicetum reptantis is shown to hold an intermediate position between them. The ass. Salico-Polemonietum acutiflori with the 2 variants, Salix lanata and S. reptans, represents various variants of mire vegetation which have under­gone zoogenic transformation (by lemmings). The stands size 1—1.5 m2 in space. The diagnostic species group comprises plants common of the wet mossy stream banks. The association is referred to the all. Caricion stantis, although many diagnostic species of the alliance are not present in its composition. When the new data are available, the association is probable to be separated into an independent alliance.


1892 ◽  
Vol 9 (2) ◽  
pp. 54-64
Author(s):  
Henry H. Howorth

In some papers which you have done me the favour to print in the GEOLOGICAL MAGAZINE, I have endeavoured to apply a new touchstone to test the age of high mountain chains and of land of high elevation, namely, the presence or absence of distinct and prominent traces of former glaciation, and I have argued that where such traces are not forthcoming in a very unmistakable manner, we are justified in concluding that these highlands have been elevated since the so-called Glacial Period. I have endeavoured to apply the touchstone in question to the Ural and Altai Mountains, to the Thian Shan and Himalaya ranges in Asia, and to the great Cordillera which binds together the two continents forming the New World. I wish to make my survey more complete by an examination of an interesting area comprising Eastern Europe and South Western Asia.


Author(s):  
Nina F. Kuznetsova ◽  
◽  
Elena S. Klushevskaya ◽  
Elena Yu. Amineva

Forest steppe of the Central Chernozem Region (CCR) of Russia belongs to the zone of highly productive pine forests. In 2015, for the first time a partial destabilization of Scots pine (Pinus sylvestris L.) was recorded within the territory of the CCR. It affected the population, organism and cellular levels of Scots pine (Pinus sylvestris L.). The destabilization was caused by the 8-year heatwave of 2007–2014 followed by a sharp drop in the water table and four severe droughts (2007, 2010, 2012, and 2014). The analysis was carried out on two sites of pine forest plantations growing in the environmentally sound region: the Stupino test site (Voronezh region, typical plantation for the CCR) and the Usman site (Lipetsk region, lands with elevated groundwater level). The results of morphological, cytogenetic and biochemical studies of model trees of the Stupino test site during the following periods are presented: 4 optimal years in terms of weather conditions, 2014 drought year and 2015 destabilization year. It was found that prolonged hydrothermal stress resulted in the transition of pine from the basic equilibrium state to a slightly nonequilibrium state. The trigger mechanism for changing their vital state was a severe autumn soil drought in 2014, after which the plants became weakened right before winter. A decrease in cone bioproductivity by the traits of seed fullness and the total number of seeds per cone, a change in population sampling structure, an increase in the number of mitosis pathologies, and an increase in proline content in needles were observed despite optimal weather conditions in 2015. The recovery of species was studied for three subsequent optimal years on the example of the Stupino and Usman populations. Experimental data indicate that the processes of vital state normalization involve profound changes in metabolism and require certain energy expenditures. It took the Stupino population longer to return to the regional norm, which indicates a different depth of destabilization of the tree genetic material of the studied populations. For citation: Kuznetsova N.F., Klushevskaya E.S, Amineva E.Yu. Highly Productive Pine Forests in a Changing Climate. Lesnoy Zhurnal [Russian Forestry Journal], 2021, no. 6, pp. 9–23. DOI: 10.37482/0536-1036-2021-6-9-23


Zootaxa ◽  
2007 ◽  
Vol 1602 (1) ◽  
pp. 1-89 ◽  
Author(s):  
RUBÉN RÍOS ◽  
J. EMMETT DUFFY

We review the taxonomy of sponge-dwelling shrimp in the “Gambarelloides species group” within the genus Synalpheus Bate 1888, an informal but widely recognized group that is largely endemic to the western Atlantic and contains the majority of Synalpheus species in that region. The validity of most species described from the western Atlantic is reevaluated based on extensive new material from Carrie Bow Cay, Belize, and on examination of types of most described species. Twenty-eight species, including all those historically considered as part of the Gambarelloides group, are herein removed from Synalpheus and transferred to Zuzalpheus, n. gen., which is diagnosed by two synapomorphies: the dense brush of curved setae on the minor first pereopod, and the mesial lamellae on the coxae of the 3 rd pereopods. Six new species are described (Zuzalpheus dardeaui, Z. elizabethae, Z. idios, Z. kensleyi, Z. ul, Z. yano) and Z. osburni (Schmitt 1933) n. comb. is removed from synonymy with Synalpheus goodei. An identification key to all 34 species of West Atlantic Zuzalpheus is presented, as are known host associations, and color plates of most species.


Zootaxa ◽  
2018 ◽  
Vol 4403 (3) ◽  
pp. 540 ◽  
Author(s):  
GABRIEL E. RAMOS-TAFUR

A new species of deep water alpheid shrimp, Alpheus luiszapatai sp. nov., from Arusí, Chocó, Pacific coast of Colombia is described. The single female known was collected between the discarded bycatch of deep water shrimp trawls dedicated to the commercial fisheries of the “coliflor” shrimp Solenocera spp. This new species is placed putatively in the Alpheus brevirostris (Olivier, 1811) species group, and share some external morphological characters with Alpheus hephaestus Bracken-Grissom & Felder, 2014. It can be differentiated by the shape and ornamentation of major and minor chelipeds, the propodi and dactyli of third to fifth pereopods, the diaresis of uropodal exopod, the length of the rostral carina, color in life and bathymetric distribution. Additional comparison with another congeners pertaining to this species group complex from the eastern Pacific, western Atlantic and other oceanographic regions is discussed. A key for Alpheus brevirostris species group from the eastern Pacific is presented. 


Lazaroa ◽  
2013 ◽  
Vol 34 (1) ◽  
pp. 191-208 ◽  
Author(s):  
José Antonio López-Sáez ◽  
Daniel Sánchez-Mata ◽  
Francisca Alba-Sánchez ◽  
Daniel Abel-Schaad ◽  
Rosario G. Gavilán ◽  
...  

2001 ◽  
Vol 79 (6) ◽  
pp. 739-746 ◽  
Author(s):  
José M Gómez ◽  
José A Hódar ◽  
Regino Zamora ◽  
Jorge Castro ◽  
Daniel García

The spatial structure of plant communities as well as the quality and abundance of neighbours can strongly influence the intensity of herbivory suffered by a plant. In this paper, we study the effect of the association with shrubs on the ungulate herbivory suffered by Scots pine (Pinus sylvestris var. nevadensis Christ., Pinaceae) saplings in two isolated, fragmented populations in southeastern Spain. For this, we monitored herbivory on saplings with regard to the microhabitat in which they grew. We distinguished pines growing in open interspaces, on the edge of shrubs and within the canopy of shrubs, and also we distinguished four functional types of shrubs: thorny shrubs, nonthorny shrubs, thorny scrubs, and nonthorny scrubs. Our results show that association with shrubs increases the damage inflicted on Scots pine saplings. In fact, saplings growing in the open patches, far from the shrubs, escaped from herbivory more frequently and incurred less damage than did saplings growing close to shrubs. However, herbivory was also reduced when pine saplings were completely surrounded by shrubs, since then they served as a mechanical barrier. The type of neighbouring shrub did not affect the overall damage suffered by pines, despite the fact that the ungulates damaged the nonthorny scrubs more intensely than the other kinds of shrubs. Consequently, saplings have an advantage when growing within the canopy of shrubs; these constitute key microsites for pine recruitment in these Mediterranean forests.Key words: associational resistance, associational susceptibility, mammalian herbivory, Mediterranean high mountain, neighbouring effects, Pinus sylvestris var. nevadensis.


2003 ◽  
Vol 33 (7) ◽  
pp. 1211-1226 ◽  
Author(s):  
Mikhail Yatskov ◽  
Mark E Harmon ◽  
Olga N Krankina

Coarse woody debris (CWD), represented by logs and snags >10 cm in diameter and >1 m in length, was sampled at eight sites in Russian boreal forests to determine the specific density of decay classes and decomposition rates. Tree species sampled included Abies siberica Ledeb., Betula pendula Roth., Betula costata Trautv., Larix siberica Ledeb., Larix dahurica Turcz., Picea abies (L.) Karst., Picea obovata Ledeb., Picea ajanensis Fisch., Pinus koraiensis Sieb. et Zucc., Pinus siberica Ledeb., Pinus sylvestris L., and Populus tremula L. The mean densities for decay clas ses 1 through 5 ranged from 0.516 to 0.084 g·cm–3, respectively. Annual decomposition rates varied among the species, and for logs, decomposition rates ranged from 4.2 to 7.8% for B. pendula, 2.6 to 4.9% for Picea spp., 2.7 to 4.4% for Pinus sylvestris, 1.5 to 3.1% for Larix spp., and 1.5 to 1.9% for Pinus koraiensis and Pinus siberica. Logs decomposed faster than snags. Among the sites examined, temperature and precipitation did not correlate with decomposition rates, which is consistent with other studies in the boreal region. Globally, a positive correlation between decomposition and mean annual temperatures was found, with decay-resistant trees less responsive than those with low decay resistance.


2013 ◽  
Vol 43 (10) ◽  
pp. 929-938 ◽  
Author(s):  
Anna Lintunen ◽  
Pekka Kaitaniemi ◽  
Jari Perttunen ◽  
Risto Sievänen

This is a first attempt to analyse species-specific light attenuation in mixed boreal forests created by shoot-level 3D tree models. The models are configurations of real individual Scots pine (Pinus sylvestris L.) and silver birch (Betula pendula Roth.) trees grown in mixed-forest stands. We study species-specific differences in radiation transmission by using the ray-casting method. Light transmission was found to be higher in dense birch-dominated stands compared with dense pine-dominated stands because of the higher total foliage area and the higher location of foliage in the pine canopy. Transmission of light per leaf area index (LAI) was nevertheless higher in the pine canopy compared with the birch canopy because of foliage clumping. Especially in clumped canopy, species-specific shoot-level light simulations enable a more realistic estimation of light transmission compared with simpler calculations based on LAI and Beer–Lambert’s law. The observed differences in light-transmission characteristics of the studied species may influence the development of target crowns in the neighbourhood. Light attenuation caused by the dense foliage zone in the upper pine canopy could be avoided in dense stands by mixing species with different vertical foliage distributions.


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