Stem-group stick insects (Phasmatodea) in the early Eocene at McAbee, British Columbia, Canada, and Republic, Washington, United States of America

2015 ◽  
Vol 147 (6) ◽  
pp. 744-753 ◽  
Author(s):  
S. Bruce Archibald ◽  
Sven Bradler
Keyword(s):  
United States ◽  
British Columbia ◽  
New Species ◽  
United States Of America ◽  
New Genus ◽  
Early Eocene ◽  
New Genus And Species ◽  
Stem Group ◽  
Stick Insects ◽  
The Republic

AbstractStem-group Phasmatodea, known as the Susumanioidea, are previously established from the Jurassic through the Paleocene. Here, we extend this record to the early Eocene with five new fossils: two forewings from the Klondike Mountain Formation exposures at Republic, Washington, United States of America, and three partially complete specimens from the McAbee locality in southern British Columbia, Canada. We assign both of the Republic specimens to the new genus and species Eoprephasma hichensinew genus, new species. Two of the McAbee fossils appear to represent two further new species, which we refer to as Susumanioidea species A and B for lack of clearly preserved diagnostic species-level character states. The third might belong to one of these two species, but this is unclear. In all three, the mesothorax and metathorax are not notably extended, the forewings are not shortened, the foreleg femur is straight, and species A possesses an extended, external ovipositor with an operculum (unknown in the other specimens). These conditions are rare and never found in combination in Euphasmatodea. All other stem-group Phasmatodea younger than the Early Cretaceous of China are only known from isolated wings.

New early Eocene Siricomorpha (Hymenoptera: Symphyta: Pamphiliidae, Siricidae, Cephidae) from the Okanagan Highlands, western North America

2015 ◽  
Vol 148 (2) ◽  
pp. 209-228 ◽  
Author(s):  
S. Bruce Archibald ◽  
Alexandr P. Rasnitsyn
Keyword(s):  
United States ◽  
British Columbia ◽  
New Species ◽  
United States Of America ◽  
Host Plants ◽  
New Genus ◽  
Early Eocene ◽  
New Genera ◽  
Western North America ◽  
Okanagan Highlands

AbstractWe describe three new genera and four new species (three named) of siricomorph sawflies (Hymenoptera: Symphyta) from the Ypresian (early Eocene) Okanagan Highlands: Pamphiliidae,Ulteramus republicensisnew genus, new speciesfrom Republic, Washington, United States of America; Siricidae,Ypresiosirex orthosemosnew genus, new speciesfrom McAbee, British Columbia, Canada; and Cephidae,Cuspilongus cachecreekensisnew genus, new speciesfrom McAbee and another cephid treated as Cephinae species A from Horsefly River, British Columbia, Canada. These are the only currently established occurrences of any siricomorph family in the Ypresian. We treat the undescribed new siricoid from the Cretaceous Crato Formation of Brazil as belonging to the Pseudosiricidae, not Siricidae, and agree with various authors that the YpresianMegapterites mirabilisCockerell is an ant (Hymenoptera: Formicidae). The Miocene speciesCephites oeningensisHeer andC. fragilisHeer, assigned to the Cephidae over a century and a half ago, are also ants. Many of the host plants that siricomporphs feed upon today first appeared in the Eocene, a number of these in the Okanagan Highlands in particular. The Okanagan Highlands sites where these wasps were found also had upper microthermal mean annual temperatures as are overwhelmingly preferred by most modern siricomorphs, but were uncommon in the globally warm Ypresian, only found then in higher elevations and highest latitudes.

Early Eocene big headed flies (Diptera: Pipunculidae) from the Okanagan Highlands, western North America

2014 ◽  
Vol 146 (4) ◽  
pp. 429-443 ◽  
Author(s):  
S. Bruce Archibald ◽  
Christian Kehlmaier ◽  
Rolf W. Mathewes
Keyword(s):  
United States ◽  
British Columbia ◽  
New Species ◽  
United States Of America ◽  
New Genus ◽  
Late Eocene ◽  
Character State ◽  
Early Eocene ◽  
The Family ◽  
Okanagan Highlands

AbstractThree new species of Pipunculidae (Diptera) are described (one named), from the early Eocene (Ypresian) Okanagan Highlands of British Columbia, Canada and Washington State, United States of America: Metanephrocerus belgardeaenew species from Republic, Washington; and Pipunculidae species A and Pipunculinae species A from Quilchena, British Columbia. We re-describe the late Eocene (Priabonian) species Protonephrocerus florissantius Carpenter and Hull from Florissant, Colorado, United States of America, and assign it to a new genus proposed here, Priabonanew genus. Pipunculinae species A is the oldest known member of the family whose wing lacks a separated M2 vein; previously this had been known in species only as old as Miocene Dominican amber. This is a presumably derived character state that is predominant in modern species. Molecular analysis indicates an origin of the Pipunculidae in the Maastrichtian; the morphological and taxonomic diversity seen here in the Ypresian is consistent with an early radiation of the family. This is concordant with the radiation of Auchenorrhyncha, upon which they mostly prey, which is in turn associated with the early Paleogene diversification of angiosperm-dominated forests recovering from the K-Pg extinction event.

scholarly journals The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America

Zootaxa ◽  
2021 ◽  
Vol 4934 (1) ◽  
pp. 1-133
Author(s):  
S. BRUCE ARCHIBALD ◽  
ROBERT A. CANNINGS ◽  
ROBERT J. ERICKSON ◽  
SETH M. BYBEE ◽  
ROLF W. MATHEWES
Keyword(s):  
New Species ◽  
New Taxa ◽  
New Genus ◽  
Early Eocene ◽  
Western North America ◽  
New Genus And Species ◽  
Genus Level ◽  
New Family ◽  
Okanagan Highlands ◽  
Head Morphology

We describe the Cephalozygoptera, a new, extinct suborder of Odonata, composed of the families Dysagrionidae and Sieblosiidae, previously assigned to the Zygoptera, and possibly the Whetwhetaksidae n. fam. The Cephalozygoptera is close to the Zygoptera, but differs most notably by distinctive head morphology. It includes 59 to 64 species in at least 19 genera and one genus-level parataxon. One species is known from the Early Cretaceous (Congqingia rhora Zhang), possibly three from the Paleocene, and the rest from the early Eocene through late Miocene. We describe new taxa from the Ypresian Okanagan Highlands of British Columbia, Canada and Washington, United States of America: 16 new species of Dysagrionidae of the existing genus Dysagrion (D. pruettae); the new genera Okanagrion (O. threadgillae, O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum, O. worleyae, all new species); Okanopteryx (O. jeppesenorum, O. fraseri, O. macabeensis, all new species); Stenodiafanus (S. westersidei, new species); the new genus-level parataxon Dysagrionites (D. delinei new species, D. sp. A, D. sp. B, both new); and one new genus and species of the new family Whetwhetaksidae (Whetwhetaksa millerae). 

Early Eocene snakeflies (Raphidioptera) of western North America from the Okanagan Highlands and Green River Formation

Zootaxa ◽  
2021 ◽  
Vol 4951 (1) ◽  
pp. 41-79
Author(s):  
S. BRUCE ARCHIBALD ◽  
VLADIMIR N. MAKARKIN
Keyword(s):  
United States ◽  
British Columbia ◽  
North America ◽  
United States Of America ◽  
Green River Formation ◽  
Early Eocene ◽  
Western North America ◽  
Green River ◽  
The Family ◽  
Okanagan Highlands

Four new species of Raphidiidae are described from the early Eocene of western North America: Megaraphidia antiquissima sp. nov. from McAbee, M. ootsa sp. nov. from Driftwood Canyon, M. hopkinsi sp. nov. from the Allenby Formation (all from British Columbia, Canada), M. klondika sp. nov. from Republic (Washington, United States of America). Archiinocellia Handlirsch, 1910, Archiinocellia oligoneura Handlirsch, 1910 from Horsefly River (British Columbia, Canada), and A. protomaculata (Engel, 2011), comb. nov., from the Green River Formation (Colorado, United States of America) are redescribed. Archiinocellia is assigned to Raphidiidae, sit. nov. The apparent absence of sclerotized gonocoxites 9 in the Archiinocellia protomaculata male is probably plesiomorphic at the family level. As some modern snakeflies do not require a cold interval to complete their development and Eocene Inocelliidae and Raphidiidae lived in regions of warm winters (especially A. protomaculata), adaptation to cold winters in many modern snakeflies is a post-Eocene phenomenon. Eocene Raphidiidae of Europe (Priabonian) differ greatly from those of North America (Ypresian and Priabonian). This pattern might reflect dispersal in either direction or ranges established prior to continental separation. Eocene Inocelliidae of Europe (Priabonian), however, are more similar to those of North America (Ypresian and Lutetian). 

scholarly journals A new genus and three new species of noctuid moths from western United States of America and Mexico (Lepidoptera, Noctuidae, Noctuinae, Eriopygini)

ZooKeys ◽  
2018 ◽  
Vol 788 ◽  
pp. 183-199 ◽  
Author(s):  
Lars G. Crabo
Keyword(s):  
United States ◽  
New Species ◽  
North America ◽  
United States Of America ◽  
New Genus ◽  
The United States ◽  
Western United States ◽  
Distribution Maps ◽  
Three New Species ◽  
Lepidoptera Noctuidae

The genus Rhabdorthodesgen. n. is described for three previously unnamed noctuid moths from the mountains of south-western United States and Mexico. It is assigned to subfamily Noctuinae, tribe Eriopygini. Rhabdorthodespattersonisp. n. from the United States and Rhabdorthodesdurangosp. n. and Rhabdorthodespetersonisp. n. from Mexico are described. These moths are small, dull gray brown, and lack highly diagnostic wing markings, but are distinctive structurally. The adults and genitalia of both sexes are illustrated and distribution maps are presented. Two species eponyms honor persons who have facilitated the study and enjoyment of moths in North America by creating moth-specific websites.

Taxonomic status of Macrocerides species (Diptera: Phoridae)

2011 ◽  
Vol 143 (6) ◽  
pp. 697-705
Author(s):  
Brian V. Brown
Keyword(s):  
United States ◽  
New Species ◽  
United States Of America ◽  
New Genus ◽  
Taxonomic Status ◽  
New Combination ◽  
New Combinations ◽  
New Subgenus ◽  
Two New Species ◽  
Northern South America

AbstractThe taxonomic status of the nine species of the mostly Neotropical genus Macrocerides Borgmeier is reviewed after the holotypes of seven species are examined. Macrocerides anacleti Borgmeier, M. brevicornis Borgmeier, and M. luteus Borgmeier are transferred to Myriophora Brown (new combinations). Macrocerides neivai Borgmeier is transferred to Apocephalus (new combination), creating a homonym with Apocephalus neivai Borgmeier for which the replacement name A. elizaldae is proposed. The African Macrocerides leydheckeri Schmitz is placed in a new genus, Macrocerophora. Tauricornus, a new subgenus of Macrocerides, is proposed for a monophyletic group that includes M. (T.) abaristalis Borgmeier, M. (T). attophilus Disney, and two new species, M. (T.) taurocephalus from southern Texas, United States of America, and M. (T.) borkenti from Costa Rica to northern South America.

A new genus, four new species and taxonomic and geographic notes in Mexican Cerambycidae

Zootaxa ◽  
2018 ◽  
Vol 4420 (2) ◽  
pp. 189
Author(s):  
ANTONIO SANTOS-SILVA ◽  
JUAN PABLO BOTERO ◽  
STÉPHANE LE TIRANT
Keyword(s):  
United States ◽  
New Species ◽  
Geographical Distribution ◽  
United States Of America ◽  
New Genus ◽  
The United States ◽  
New Combination ◽  
New Mexican ◽  
First Time ◽  
Mexican Species

Four new Mexican species are described: Ironeus curoei from Guerrero state, Psyrassa garciai from Tamaulipas state, Psyrassa belangeri from Guerrero state (Elaphidiini, Cerambycinae); and Cirrhicera bankoi from Chiapas state (Hemilophini, Lamiinae). A new Hesperophanini genus, Makromastax, and a new combination, Makromastax mandibularis are proposed. The current key to the Mexican genera of Hesperophanini is updated to include the new genus. Haplidus nitidus Chemsak and Linsley, 1963 is proposed as synonyms of Haplidus mandibularis Chemsak and Linsley, 1963. The male of Eburia (E.) girouxae is illustrated by the first time and Anelaphus hirtus is redescribed and figured. The geographical distribution of 9 species is expanded, the type locality of Osmidus guttatus is corrected, and Neocompsa intrincata is excluded from the fauna of the United States of America. 

Arenaeocoris enervatus (Hemiptera: Heteroptera: Reduviidae: Stenopodainae), a new genus and species from the Southeastern United States

Zootaxa ◽  
2012 ◽  
Vol 3478 (1) ◽  
pp. 105-110
Author(s):  
ROBERT L. BLINN
Keyword(s):  
United States ◽  
New Species ◽  
New Genus ◽  
Southeastern United States ◽  
The United States ◽  
New Genus And Species ◽  
A New Species

Arenaeocoris gen. nov., and a new species, A. enervatus sp. nov., is described from southeastern United States. Diagnosticcharacters to distinguish the new genus from related taxa in the United States, illustrations of the species, and a key to the genera for the United States incorporating the new genus, are provided.

A new Isectolophid Tapiromorph (Perissodactyla, Mammalia) from the Early Eocene of Pakistan

2001 ◽  
Vol 75 (2) ◽  
pp. 407-417 ◽  
Author(s):  
M. C. Maas ◽  
S. T. Hussain ◽  
J. J. M. Leinders ◽  
J. G. M. Thewissen
Keyword(s):  
New Species ◽  
North American ◽  
New Genus ◽  
Middle Eocene ◽  
Early Eocene ◽  
New Genus And Species ◽  
Northwest India

A new genus and species of tapiromorph, Karagalax mamikhelensis, is described from the Eocene Mami Khel Formation of northwest Pakistan. The new species is known from adult and juvenile dentitions, juvenile skulls, and partial postcrania. It is the most primitive perissodactyl yet reported from Indo-Pakistan. The morphology of its lophodont molars indicates that Karagalax is a tapiromorph, and it is here included in the primitive family Isectolophidae. Karagalax is more derived (more lophodont) than North American isectolophids Systemodon and Cardiolophus or the Asian early Eocene Orientolophus and Homogalax wutuensis, and more primitive (less lophodont) than North American Homogalax and Isectolophus. It is distinct from the poorly known and enigmatic Indian isectolophid Sastrilophus. Karagalax lacks any derived features of the Deperetellidae, Helaletidae or Lophialetidae, including Kalakotia, a primitive lophialetid from the middle Eocene of northwest India. The partial postcrania of Karagalax, which include fragmentary humeri, femora, ulnae, tibiae and metapodials, show a combination of primitive and derived features and suggest that it was more cursorial than other basal tapiromorphs for which postcrania are known.A provisional analysis of the phylogenetic positions of Karagalax and Kalakotia supports the hypothesis that primitive perissodactyls dispersed to Indo-Pakistan, most probably by way of continental Asia. The evolutionary position of Karagalax is consistent with an early Eocene age for H-GSP Locality 300, as argued previously on the basis of other mammals.

scholarly journals A new genus and new species of Baetidae (Ephemeroptera) from lakes and reservoirs in eastern North America

Zootaxa ◽  
2010 ◽  
Vol 2481 (1) ◽  
Author(s):  
MATTHEW A. HILL ◽  
JOHN PFEIFFER ◽  
LUKE M. JACOBUS
Keyword(s):  
United States ◽  
New Species ◽  
North America ◽  
United States Of America ◽  
New Genus ◽  
Eastern North America ◽  
Eastern United States ◽  
Lakes And Reservoirs

Waynokiops dentatogriphus, new genus and new species (Ephemeroptera: Baetidae), is described from nymphs collected from seven lakes and reservoirs in the eastern United States of America, specifically from the states of Virginia, Kentucky, Ohio, Indiana and Arkansas (type locale), during sampling for the USEPA National Lakes Assessment. The lateral abdominal expansions and the dorsal abdominal armature of the new genus are unique among Cloeon complex genera. Abdominal tergal projections are unusual for a small minnow mayfly from lentic habitats.

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