Subspeciation within a punctuated equilibrium evolutionary event: Phylogenetic history of the Lower-Middle Ordovician Paroistodus originalis—P. horridus complex (Conodonta)

2000 ◽  
Vol 74 (3) ◽  
pp. 492-502
Author(s):  
Guillermo L. Albanesi ◽  
Christopher R. Barnes

A microevolutionary event involving the conodont Paroistodus lineage is documented in the Gualcamayo Formation (Middle Ordovician), Argentine Precordillera. A detailed sampling of limestones throughout the upper part of the San Juan Formation and the lower member of the Gualcamayo Formation yielded over 14,000 well-preserved conodont elements. Paroistodus originalis (Sergeeva, 1963) was recorded through the upper 230 m of the San Juan Formation and the lower member (10 m thick) of the Gualcamayo Formation. The derived species Paroistodus horridus (Barnes and Poplawski, 1973) was recorded throughout the middle member of the Gualcamayo Formation (65 m thick). The intermediate linking forms between both species are identified as two new taxa: Paroistodus horridus primus Albanesi, 1998b, and P. h. secundus Albanesi, 1998b. They were recorded in the uppermost 70 cm of the lower member. Apparently, the speciation event occurred under stressed environmental conditions with the drowning of the carbonate platform, i.e., the San Juan Formation, and the beginning of a deeper and restricted environment represented by the Gualcamayo black shales. The demise of the carbonate production was caused by a sea level rise and a significant influx of volcanic ashes. The punctuated speciation event occurred within an allopatric setting while the Precordillera occupied an isolated (Iapetus) oceanic position in its overall drift from Laurentia to Gondwana.

2020 ◽  
Vol 47 (1) ◽  
pp. 125 ◽  
Author(s):  
Fernanda Serra ◽  
Nicolás A. Feltes ◽  
Matías Mango ◽  
Miles A. Henderson ◽  
Guillermo L. Albanesi ◽  
...  

The Ordovician System is extensively represented in the Precordillera of San Juan Province, Argentina. At the Cerro La Chilca in the Jáchal area, the limestone of the San Juan Formation is paraconformably overlain by interbedded limestone and shale of the Gualcamayo Formation. The present contribution reports new data on the conodont fauna and biostratigraphy of these darriwilian units, revising local and regional chronostratigraphic relationships. New information on the composition of conodont and graptolite associations through the stratigraphic sequence is presented. The presence of Paroistodus horridus horridus, Yangtzeplacognathus crassus, and Histiodella sinuosa constrain the uppermost strata of the San Juan Formation to the lower part of the Y. crassus Zone, according to the Baltoscandian scheme, and to the H. sinuosa Subzone of the Periodon macrodentatus Zone of the North American scheme. In the overlying Gualcamayo Formation the co-occurrence of Y. crassus with Histiodella holodentata enable the recognition of the Y. crassus Zone and the H. holodentata Subzone of the P. macrodentatus Zone. The identification of these zones allows for precise global and regional correlation. A graptolite assemblage that belongs to the epipelagic and deep-water biotopes with some components restricted to low paleolatitudes is recognized. This diverse assemblage is characteristic of the pelagic biofacies. The important diversity of graptolites in this section suggests a favorable environment for their development. Local changes in the taxonomic composition are recognized through the Gualcamayo Formation. When comparing this fauna with that of different study localities from the Central Precordillera (Cerro Potrerillo, Oculta Creek, Cerro Viejo de Huaco and Las Aguaditas Creek) slight differences in the generic composition are observed. Taxonomic differences support the preference of certain associations for particular environments; though, graptolites are more diverse in black shales facies, which represent deeper environments (the Los Azules Formation), in relation to the calcareous-shale facies of the Gualcamayo Formation from Cerro La Chilca and correlative unit at Las Aguaditas Creek.


1990 ◽  
Vol 27 (1) ◽  
pp. 14-26 ◽  
Author(s):  
Sheila R. Stenzel ◽  
Ian Knight ◽  
Noel P. James

Carbonates of the Table Head Group and associated strata were deposited along the western margin of a foreland basin during initial stages of the Taconian Orogeny and record collapse and cannibalization of a long-lived carbonate platform. The stratigraphy of Klappa, Opalinski, and James is here revised to reflect better understanding of lithologic units within this complex tectono-stratigraphic assemblage. The Table Head Group now contains only three formations: the Table Point and Table Cove formations, as originally defined, and the Cape Cormorant Formation, redefined and restricted to western Port au Port Peninsula. Black Cove Formation shales are removed from the Table Head. Distinctive conglomerates once placed in the Cape Cormorant are now recognized as separate units within the overlying flysch and called the Daniel's Harbour Member.The foreland basin developed in three stages: (1) fragmentation, uplift, and erosion of the platform and subsequent deposition of shallow-water limestones (Table Point) on a tectonically unstable shelf; (2) foundering of platform blocks and deposition of deep-water-slope carbonates (Table Cove), basinal black shales (Black Cove), or conglomerates of older shelf carbonates shed from submarine cliffs (Cape Cormorant); and (3) siliciclastic sedimentation interrupted by sediment gravity flows of Table Head clasts shed from submarine escarpments (Daniel's Harbour).


2021 ◽  
Vol 15 (1) ◽  
pp. 37
Author(s):  
Matilde Sylvia Beresi ◽  
Susana Emma Heredia

Sponge spicule assemblages are described fom residues of conodont samples from Ordovician strata in the Sierra Pintada, southern Mendoza Province, Argentina. Spicules have been recovered from the Arenigian allochthonous megaconglomerates and from autochthonous limestones and carbonates sandstones of the Ponón Trehue Formation. This formation is a elastic-carbonate sequence representing olistostromic and turbidite facies. Conodonts in this formation are Llandeillan in age. The spicules are calcified and moderately preserved. The material shows a low diversity. Poriferan taxa found in this formation include heteractinid spicules as well as hexactinellid hexactines and non-lithistid demospongiid triaene and oxeas with some doubt. Associations of exclusively heteractinid spicules are restricted to allochthonous blocks of the shallow carbonate platform of the San Juan Formation (Arenig). In the outer platform and slope, autochthonous calcarenites and dark limestones contain hexactine spicules. These spicules evidence the existence of sponges in the Ordovician of the Ponón Trehue area, as a part of the Precordillera terrane. 


1980 ◽  
Vol 17 (8) ◽  
pp. 1007-1019 ◽  
Author(s):  
Colin F. Klappa ◽  
Paul R. Opalinski ◽  
Noel P. James

Lithostratigraphic nomenclature of early Middle Ordovician strata from western Newfound land is formally revised. The present Table Head Formation is raised to group status and extended to include overlying interbedded terrigenoclastic-rich calcarenites and shales with lime megabreccias. Four new formation names are proposed: Table Point Formation (previously lower Table Head); Table Cove Formation (previously middle Table Head); Black Cove Formation (previously upper Table Head); and Cape Cormorant Formation (previously Caribou Brook formation). The Table Point Formation comprises bioturbated, fossiliferous grey, hackly limestones and minor dolostones; the Table Cove Formation comprises interbedded lime mudstones and grey–black calcareous shales; the Black Cove Formation comprises black graptolitic shales; and the Cape Cormorant Formation comprises interbedded terrigenoclastic and calcareous sandstones, siltstones, and shales, punctuated by massive or thick-bedded lime megabreccias. The newly defined Table Head Group rests conformably or disconformably on dolostones of the Lower Ordovician St. George Group (an upward-migrating diagenetic dolomitization front commonly obscures the contact) and is overlain concordantly by easterly-derived flysch deposits. Upward-varying lithologic characteristics within the Table Head Group result from fragmentation and subsidence of the Cambro-Ordovician carbonate platform and margin during closure of a proto-Atlantic (Iapetus) Ocean.


2018 ◽  
Author(s):  
David Patrick Gold ◽  
James P. G. Fenton ◽  
Manuel Casas-Gallego ◽  
Vibor Novak ◽  
Irene Pérez-Rodríguez ◽  
...  

The island of Jamaica forms the northern extent of the Nicaraguan Rise, an elongate linear tectonic feature stretching as far as Honduras and Nicaragua to the south. Uplift and subaerial exposure of Jamaica during the Neogene has made the island rare within the Caribbean region, as it is the only area where rocks of the Nicaraguan Rise are exposed on land. Biostratigraphic dating and palaeoenvironmental interpretations using larger benthic foraminifera, supplemented by planktonic foraminifera, nannopalaeontology and palynology of outcrop, well and corehole samples has enabled the creation of a regional relative sea-level curve through identification of several depositional sequences. This study recognises ten unconformity-bounded transgressive-regressive sequences which record a complete cycle of relative sea level rise and fall. Sequences are recognised in the Early to ‘Middle’ Cretaceous (EKTR1), Coniacian-Santonian (STR1), Campanian (CTR1), Maastrichtian (MTR1-2), Paleocene-Early Eocene (PETR1), Eocene (YTR1-3) and Late Eocene-Oligocene (WTR1). These transgressive-regressive cycles represent second to fourth order sequences, although most tie with globally recognised third order sequences. Comparisons of the Jamaican relative sea-level curve with other published global mean sea-level curves show that local tectonics exerts a strong control on the deposition of sedimentary sequences in Jamaica. Large unconformities (duration >1 Ma) are related to significant regional tectonic events, with minor overprint of a global eustatic signal, while smaller unconformities (duration <1 Ma) are produced by global eustatic trends. The relatively low rates of relative sea-level rise calculated from the regional relative sea-level curve indicate that carbonate production rates were able to keep pace with the rate of relative sea-level rise accounting for the thick successions of Maastrichtian carbonates and those of the Yellow and White Limestone Groups. Carbonate platform drowning within the White Limestone Group during the Oligocene to Miocene is attributed to environmental deterioration given the low rates of relative sea-level rise.


2006 ◽  
pp. 13-17 ◽  
Author(s):  
Platon Tchoumatchenco ◽  
Dragoman Rabrenovic ◽  
Barbara Radulovic ◽  
Vladan Radulovic

In the region across the Serbian/Bulgarian state border, there are individualized 5 Jurassic paleogeographic units (from West to East): (1) the Thracian Massif Unit without Jurassic sediments; (2) the Luznica-Koniavo Unit - partially with Liassic in Grsten facies and with deep water Middle Callovian-Kimmeridgian (p. p) sediments of the type "ammonitico rosso", and Upper Kimmeridgian-Tithonian siliciclastics flysch; (3) The Getic Unit subdivided into two subunits - the Western Getic Sub-Uni - without Lower Jurassic sediments and the Eastern Getic Sub-Unit with Lower Jurassic continental and marine sediments, which are followed in both sub-units by carbonate platform limestones (type Stramberk); (4) the Infra (Sub)-Getic Unit - with relatively deep water Liassic and Dogger sediments (the Dogger of type "black shales with Bossitra alpine") and Middle Callovian-Tithonian of type "ammonitico rosso"; (5) the Danubian Unit - with shallow water Liassic, Dogger and Malm (Miroc-Vrska Cuka Zone, deep water Dogger and Malm (Donjomilanovacko-Novokoritska Zone).


2006 ◽  
pp. 19-33 ◽  
Author(s):  
Platon Tchoumatchenco ◽  
Dragoman Rabrenovic ◽  
Barbara Radulovic ◽  
Vladan Radulovic

The Infra-Getic Unit is a palaeogeographic unit, predestined by palaeotectonics. From the point of view of geological heritage, it represents a geosites framework. For the purpose of the correlation, the Serbian sections of Lukanja, Bogorodica Monastery, Rosomac and Senokos, as well as the Bulgarian sections of Komshtitsa, Gintsi, and Stanyantsi were used. The Jurassic sediments of the Infra-Getic Unit crop out on the southern slops of the Stara Planina Mountain in east Serbia and west Bulgaria. The Lower Jurassic started with continental and continental-marine sediments (clays and sandstones) (Lukanja clastics and Lukanja coal beds in Serbia and the Tuden Formation in Bulgaria) and continue with Lukanja quartz sandstones (Serbia) and the Kostina Formation (Bulgaria). These sediments are covered by Lukanja brachiopod beds and Lukanja limestones (Serbia) and the Romanov Dol, Ravna and Dolni Loukovit Members of the Ozirovo Formation (Bulgaria) predominantly consist of bioclastic limestones. The sedimentations follow with Lukanja belemnites-gryphaea beds (marls and clayey limestones), which in Bulgaria correspond to the Bukorovtsi Member (also marls and clayey limestones) of the Ozirovo Formation. The Middle Jurassic sedimentation started with black shales with Bossitra alpine. These sediments are individualized in Serbia as Senokos aleurolites and clays and in Bulgaria they are known as the Etropole Formation. In Serbia the section continues with sandstones called Vodenicki sandstones of Bajocian age, known in Bulgaria as the Dobrogled Member of the Polaten Formation. However, in Bulgaria, the age is Upper Bajocian-Lower Bathonian, and it cover the marls of the lower member (Gornobelotintsi Member) of the Bov Formation and is covered by the upper member - alternation of marls and clayey limestones - the Verenitsa Member of the Bov Formation. The Vodenicki sandstones-Dobrogled Member which ended their distribution in the section of Komshtitsa, to the east (in the Gintsi section), they are not represented - build a body of sandstones, a prodelta coming from the west to the east. The Bov Formation corresponds to the Senokos ammonite beds in east Serbia. The upper boundary of the Senokos ammonite beds and of the Bov Formation is sharp. It is covered by grey limestones of the Yavorec Formation in Bulgaria and by the Kamenica limestones in eastern Serbia. They are covered by grey or red nodular/lithoclastic limestones ("ammonitico rosso" type) of the Gintsi Formation in Bulgaria and the Pokrovenik ammonitic (acanthicum) limestones in Serbia. The Jurassic section in the Infra-Getic ended with grey micritic and lithoclastic limestones, which belong to the Rosomac and Rsovci limestones in east Serbia and to the Glozhene Formation in Bulgaria.


1990 ◽  
Vol 27 (6) ◽  
pp. 731-741 ◽  
Author(s):  
Rudolf Bertrand

Carbonate platform sequences of Anticosti Island and the Mingan Archipelago are Early Ordovician to Early Silurian in age. With the exception of the Macasty Formation, the sequences are impoverished in dispersed organic matter, which is chiefly composed of zooclasts. Zooclast reflectances suggest that the Upper Ordovician and Silurian sequences outcropping on Anticosti Island are entirely in the oil window but that the Lower to Middle Ordovician beds of the Mingan Archipelago and their stratigraphic equivalents in the subsurface of most of Anticosti Island belong to the condensate zone. Only the deeper sequences of the southwestern sector of Anticosti Island are in the diagenetic dry-gas zone. The maximum depth of burial of sequences below now-eroded Silurian to Devonian strata increases from 2.3 km on southwestern Anticosti Island to 4.5 km in the Mingan Archipelago. A late upwarp of the Precambrian basement likely allowed deeper erosion of the Paleozoic strata in the vicinity of the Mingan Archipelago than on Anticosti Island. Differential erosion resulted in a southwestern tilting of equal maturation surfaces. The Macasty Formation, the only source rock of the basin (total organic carbon generally > 3.5%, shows a wide range of thermal maturation levels (potential oil window to diagenetic dry gas). It can be inferred from the burial history of Anticosti Island sequences that oil generation began later but continued for a longer period of geologic time in the northeastern part than in the southeastern part of the island. Oil generation was entirely pre-Acadian in the southern and western parts of Anticosti Island, but pre- and post-Acadian in the northern and eastern parts.


2019 ◽  
Vol 93 (2) ◽  
pp. 244-259
Author(s):  
Silviu O. Martha ◽  
Paul D. Taylor ◽  
William L. Rader

AbstractThe Glen Rose and Walnut formations of southcentral and northcentral Texas comprise shallow-water carbonates deposited during the late Aptian to middle Albian on a carbonate platform. The formations are famous for their rich fossil faunas. Although bryozoans are absent in late Aptian sediments, they are frequently found encrusting bivalve shells from the early to middle Albian parts of these formations. Here, we describe the cyclostome bryozoan fauna, which includes six species;Stomatoporasp.,Oncousoecia khirarn. sp.,Reptomultisparsa mclemoreaen. sp.,Hyporosopora keeran. sp.,Mesonopora bernardwalterin. sp., and ?Unicaveasp. Most cyclostomes are found encrusting rudist shells from Unit 2 of the Lower Member of the Glen Rose Formation and units 3 and 6 of the Upper Member of the Glen Rose Formation.UUID:http://zoobank.org/4380dcb5-63b2-4aa9-959c-09eb6b03831f


1984 ◽  
Vol 74 (4) ◽  
pp. 183-191 ◽  
Author(s):  
Lorraine E. Craig

ABSTRACTSediments, mainly sandstones, conglomerates and shales, accumulated in small turbidite fans along the northern arc–trench margin of the Iapetus Ocean from middle Ordovician to Silurian time. These fans, together with the underlying pelagic facies and part of the oceanic crust, were sliced and accreted northward resulting in the Lower Palaeozoic accretionary prism which forms the Scottish Southern Uplands and the Longford-Down inlier in Ireland. North Down is the continuation of the Northern belt of the Southern Uplands of Scotland into Ireland, bounded to the S by the Orlock Bridge fault. Lithological and petrographical comparison with the rest of the Northern belt indicates closer affinities with the Southern Uplands of Scotland than with the western end of the Longford-Down inlier. Major ENE—WSW-trending Caledonian strike faults define five blocks, in which new formations of Caradoc and ? Ashgill age are defined. Pillowed spilitic rock, interpreted as a fragment of the ocean-floor, is only recognised in the Ballygrot block. Pelagic and hemipelagic black shales and cherts are overlain by arenaceous sediments in all blocks.


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