The Triassic isopod Protamphisopus wianamattensis (Chilton) and comparison by extant taxa (Crustacea, Phreatoicidea)

2003 ◽  
Vol 77 (3) ◽  
pp. 454-470
Author(s):  
George D. F. Wilson ◽  
Gregory D. Edgecombe
Keyword(s):  
Middle Triassic ◽  
Cladistic Analysis ◽  
Late Paleozoic ◽  
Late Permian ◽  
Habitat Shift ◽  
Crown Group ◽  
Early Mesozoic ◽  
The Family ◽  
Minimum Age ◽  
Character Set

Protamphisopus wianamattensis (Chilton, 1918) from the Middle Triassic (Anisian) Ashfield Shale in the Sydney Basin, Australia, is the earliest-known freshwater representative of the basal isopod suborder Phreatoicidea. In contrast, the late Paleozoic Palaeophreatoicidae species, which are morphologically distinct from extant families, are found in marine or estuarine facies. Comparison of Protamphisopus wianamattensis with living Phreatoicidea permits the external morphology of the fossils to be reconstructed and the species to be coded for cladistic analysis using a revised and expanded character set developed for living phreatoicideans. In resulting parsimonious trees as well as immediately suboptimal trees, Protamphisopus is nested within clades related to the family Amphisopodidae. Although not included in the analysis, the Late Permian Protamphisopus reichelti Malzahn (in Glaessner and Malzahn, 1962) appears to be a member of the Palaeophreatoicidae, rather than among the crown group of the Phreatoicidea. Therefore, a minimum age of Middle Triassic can be assigned to the basal branches within the phreatoicid crown group. The minimum age for the colonisation of fresh water by the suborder is also established although, given the advanced position of Protamphisopus wianamattensis in the cladograms, the habitat shift may have occurred earlier. The biogeographic distribution of extant Phreatoicidea on fragments of Gondwana is consistent with early Mesozoic origins for the major clades of this isopod suborder.

scholarly journals Cladistic analysis of the Paleozoic bryozoan families Monticuliporidae and Mesotrypidae

2013 ◽  
Vol 87 (4) ◽  
pp. 635-649
Author(s):  
Amber D. Adamczyk ◽  
Joseph F. Pachut
Keyword(s):  
Type Species ◽  
Group Membership ◽  
Cladistic Analysis ◽  
Morphological Similarity ◽  
Single Family ◽  
Crown Group ◽  
Stem Group ◽  
The Family ◽  
Stratigraphic Ranges ◽  
Significant Match

A set of 127 binary and multistate characters, weighted by the number of derived character states, degree of covariation, and level of homoplasy, was used in a cladistic analysis of type species representing 12 genera previously assigned to families Monticuliporidae and Mesotrypidae. The most parsimonious tree consisted of a 10-genus monophyletic crown group with the remaining two genera forming a basal paraphyletic stem group. The composition of the monticuliporid crown group is broadly similar to two earlier classifications while stem group membership matches the family Mesotrypidae. Phenetic groupings, based on overall morphological similarity, have memberships that are similar to those of clades but provide no means of determining the polarity of evolutionary relationships either within or between them. Finally, only the observed stratigraphic ranges of the type species of genera provide a statistically significant match with cladistic branching sequence, perhaps because current composite generic ranges reflect the mixing of species belonging to different genera. Based on cladogram topology, we propose the placement of all 12 genera into a single family Monticuliporidae.

The limits of the family Evaniidae (Insecta: Hymenoptera) and a new genus from Lebanese Amberamber

2002 ◽  
Vol 33 (1) ◽  
pp. 23-34 ◽  
Author(s):  
Hasan H. Basibuyuk ◽  
Mike G. Fitton ◽  
Alexandr P. Rasnitsyn ◽  
Donald L.J. Quicke
Keyword(s):  
Fore Wing ◽  
New Genus ◽  
Cladistic Analysis ◽  
Wing Venation ◽  
New Genus And Species ◽  
Ground Plan ◽  
The Family ◽  
Definition Of

AbstractThe definition of the family Evaniidae is revised and Cretevaniidae are synonymised with Evaniidae based on evidence derived from recently described Mesozoic taxa and a new genus and species, Lebanevania azari, described here from Lebanese amber. A fore leg with a long trochanter and a 12-segmented antenna are autapomorphies of the new genus. A large, high and wide head and a high and short mesosoma are derived characters shared with other Evaniidae. The new genus also has complete fore wing venation and lacks a tubular petiole, which are ground plan features of the Evanioidea. A cladistic analysis of fossil and extant members of the superfamily Evanioidea and notes on fossil taxa are presented.

The halloporid trepostome bryozoans from the Ordovician Simpson Group of Oklahoma

1991 ◽  
Vol 65 (2) ◽  
pp. 200-212 ◽  
Author(s):  
Marcus M. Key
Keyword(s):  
Discriminant Analysis ◽  
North America ◽  
Cladistic Analysis ◽  
Middle Ordovician ◽  
Bryozoan Species ◽  
The Family ◽  
Distinct Lineage

The Bromide Formation of the Middle Ordovician Simpson Group of Oklahoma contains one of the oldest diverse bryozoan faunas in North America. The early divergence of many trepostome clades is revealed in these rocks. Three trepostome bryozoan species belonging to family Halloporidae are described from this fauna. Discriminant analysis is used to define the following halloporid species: Diplotrypa schindeli n. sp., Tarphophragma karklinsi n. sp., and Tarphophragma macrostoma (Loeblich). Preliminary cladistic analysis indicates that the family Halloporidae was already a distinct lineage by the Middle Ordovician. This suggests that by this time, many of the major trepostome clades were already established.

scholarly journals A New Late Permian Burnetiamorph From Zambia Confirms Exceptional Levels of Endemism in Burnetiamorpha (Therapsida: Biarmosuchia) and an Updated Paleoenvironmental Interpretation of the Upper Madumabisa Mudstone Formation

2021 ◽  
Vol 9 ◽  
Author(s):  
Christian A. Sidor ◽  
Neil J. Tabor ◽  
Roger M. H. Smith
Keyword(s):  
South Africa ◽  
Fossil Record ◽  
Cladistic Analysis ◽  
Dorsal Margin ◽  
Rift Basin ◽  
Late Permian ◽  
Paleoenvironmental Interpretation ◽  
Assemblage Zone ◽  
Trigonal Pyramid ◽  
Ceratopsian Dinosaurs

A new burnetiamorph therapsid, Isengops luangwensis, gen. et sp. nov., is described on the basis of a partial skull from the upper Madumabisa Mudstone Formation of the Luangwa Basin of northeastern Zambia. Isengops is diagnosed by reduced palatal dentition, a ridge-like palatine-pterygoid boss, a palatal exposure of the jugal that extends far anteriorly, a tall trigonal pyramid-shaped supraorbital boss, and a recess along the dorsal margin of the lateral temporal fenestra. The upper Madumabisa Mudstone Formation was deposited in a rift basin with lithofacies characterized by unchannelized flow, periods of subaerial desiccation and non-deposition, and pedogenesis, and can be biostratigraphically tied to the upper Cistecephalus Assemblage Zone of South Africa, suggesting a Wuchiapingian age. Isengops is the second burnetiamorph recognized from Zambia and is part of a tetrapod assemblage remarkably similar to others across southern Pangea during the Wuchiapingian. A revised cladistic analysis of Biarmosuchia yielded over 500 most parsimonious trees that generally reaffirm the results of previous analyses for burnetiamorphs: Lemurosaurus is basal, Lobalopex and Isengops are proximate burnetiid outgroups, and Bullacephalus, Burnetia, Mobaceras, Niuksenitia, and Pachydectes are burnetiines. Furthermore, Russian biarmosuchians are scattered throughout the tree and do not form sister taxon relationships with each other. Burnetiamorphs display a wide disparity of cranial adornments and are relatively speciose (13 species), especially when compared to the number of specimens discovered to date (∼16 specimens). As has been suggested in some other tetrapod clades (e.g., ceratopsian dinosaurs), the burnetiamorph fossil record supports an inferred macroevolutionary relationship between cranial adornment and increased speciation rate.

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