Protohemiphlebiidae fam. nov., a stem hemiphlebioid damselfly from Cretaceous amber in Kachin

2021 ◽  
pp. SP521-2020-249
Author(s):  
Daran Zheng ◽  
Edmund A. Jarzembowski ◽  
De Zhuo ◽  
André Nel

AbstractHemiphlebiidae are the most basal lestomorphan family following the latest phylogenetic analysis of the Zygoptera: this unique damselfly family today contains one relict species found in the wetlands of Australia. It was, however, very diverse and widespread during the Mesozoic. Nevertheless, very few species were known obscuring the origination and early evolution of the family. Here we propose a new stem hemiphlebioid taxon (Protohemiphlebiidae Zheng, Jarzembowski & Nel, fam. nov.) based on a new genus and two species: Protohemiphlebia zhangi Zheng, Jarzembowski & Nel, sp. nov. and Protohemiphlebia meiyingae Zheng, Jarzembowski & Nel, sp. nov. The new family shares the characters of both Hemiphlebiidae and Coenagrionoidea, but it is more closely related to Hemiphlebiidae in having the pterostigma with a ‘star-shaped’ microsculpture, and AA originating from the wing base slightly distal of Ax0. Protohemiphlebia Zheng, Jarzembowski & Nel, gen. nov. is further considered to belong to the stem group of Hemiphlebioidea, instead of belonging to the Hemiphlebiidae, in possessing pretibial combs and a weakly kinked RP1 below the Pt-brace. The new damselflies will help to calibrate the origin of Hemiphlebiidae, which could be earlier than their current oldest records in the Kimmeridgean (Late Jurassic).

1988 ◽  
Vol 62 (2) ◽  
pp. 218-233 ◽  
Author(s):  
John Mark Malinky

Concepts of the family Hyolithidae Nicholson fide Fisher and the genera Hyolithes Eichwald and Orthotheca Novak have been expanded through time to encompass a variety of morphologically dissimilar shells. The Hyolithidae is here considered to include only those hyolithid species which have a rounded (convex) dorsum; slopes on the dorsum are inflated, and the venter may be flat or slightly inflated. Hyolithes encompasses species which possess a low dorsum and a prominent longitudinal sulcus along each edge of the dorsum; the ligula is short and the apertural rim is flared. The emended concept of Orthotheca includes only those species of orthothecid hyoliths which have a subtriangular transverse outline and longitudinal lirae covering the shell on both dorsum and venter.Eighteen species of Hyolithes and one species of Orthotheca from the Appalachian region and Western Interior were reexamined in light of more modern taxonomic concepts and standards of quality for type material. Reexamination of type specimens of H. similis Walcott from the Lower Cambrian of Newfoundland, H. whitei Resser from the Lower Cambrian of Nevada, H. billingsi Walcott from the Lower Cambrian of Nevada, H. gallatinensis Resser from the Upper Cambrian of Wyoming, and H. partitus Resser from the Middle Cambrian of Alabama indicates that none of these species represents Hyolithes. Hyolithes similis is here included under the new genus Similotheca, in the new family Similothecidae. Hyolithes whitei is designated as the type species of the new genus Nevadotheca, to which H. billingsi may also belong. Hyolithes gallatinensis is referred to Burithes Missarzhevsky with question, and H. partitus may represent Joachimilites Marek. The type or types of H. attenuatus Walcott, H. cecrops Walcott, H. comptus Howell, H. cowanensis Resser, H. curticei Resser, H. idahoensis Resser, H. prolixus Resser, H. resseri Howell, H. shaleri Walcott, H. terranovicus Walcott, and H. wanneri Resser and Howell lack shells and/or other taxonomically important features such as a complete aperture, rendering the diagnoses of these species incomplete. Their names should only be used for the type specimens until better preserved topotypes become available for study. Morphology of the types of H.? corrugatus Walcott and “Orthotheca” sola Resser does not support placement in the Hyolitha; the affinities of these species are uncertain.


1992 ◽  
Vol 70 (8) ◽  
pp. 1617-1658 ◽  
Author(s):  
R. A. Shoemaker ◽  
C. E. Babcock
Keyword(s):  

Six genera of the order Pleosporales are revised taxonomically. Although five genera have unusual applanate ascospores, they are members of three different families. Clathrospora (10 species, 2 new), Comoclathris (21 species, 7 new), and Macrospora (3 species) are placed in a new family Diademaceae in which the ascoma opening is a characteristic flat circular lid. Diademosa, a new genus with one species with terete ascospores is placed in Diademaceae. Graphyllium (3 species) is placed in the family Hysteriaceae characterized by hysterothecia with a slit-like opening. Platysporoides n.gen. (11 species, 1 new) is maintained in the Pleosporaceae because of the terete pored beak of the ascomata. Key words: Pleosporales, applanate ascospores, Clathrospora, Comoclathris, Macrospora, Diademosa, Graphyllium, Platysporoides.


Zootaxa ◽  
2021 ◽  
Vol 4964 (2) ◽  
pp. 330-344
Author(s):  
YEHUDA BENAYAHU ◽  
LEEN P. VAN OFWEGEN ◽  
J. P. RUIZ ALLAIS ◽  
CATHERINE S. MCFADDEN

Because of the problematical identity and status of the type of the xeniid soft coral genus Cespitularia Milne-Edwards & Haime, 1850, the species C. stolonifera Gohar, 1938 is revised. Examination of the type colonies has led to the establishment of the new genus Unomia gen. n. which is described and depicted. This genus features a stalk, commonly divided into branches featuring a diffuse polypiferous part consisting of distal clustered polyps and proximal individual ones on the stalk or the basal membranous part of the colonies. The sclerites are ellipsoid platelets composed of dendritic calcite rods whose tips are distinct on the surface of the platelets. Freshly collected material from Venezuelan reefs where the species is invasive was subjected to molecular phylogenetic analysis, the results of which substantiate the taxonomic assignment of the new genus under U. stolonifera comb. n. A new species, U. complanatis, from Japan and Green Island (Taiwan) is described and further illustrates the extent of the interspecific morphological variation within the genus. The results reveal that the biogeographic distribution of Unomia gen. n. includes Pacific Ocean reefs in addition to the previously reported invaded Caribbean reefs. 


2020 ◽  
Vol 94 (4) ◽  
pp. 748-757
Author(s):  
Jobst Wendt

AbstractIn contrast to almost all other invertebrate phyla that constructed biomineralized skeletons during the “Cambrian explosion” and maintained them during the entire fossil record, ascidian tunicates evolved this protective and stabilizing advantage only during the Permian, although soft-bodied representatives of this subphylum made their first appearance already in the early Cambrian. It remains enigmatic why these compound calcareous skeletons persisted only until the Late Triassic, subsequently followed by less-rigid internal skeletons from the Lower Jurassic onwards, which consist of scattered isolated spicules only. In addition to recently described aragonitic ascidian exoskeletons from the Permian and Triassic, new discoveries of similar, but colonial ascidian compound endoskeletons in the lower Carnian exhibit a short-living branch of this group, which moreover contain the first indubitable calcareous spicules. The latter are embedded in the solid endoskeleton, which is composed of polygonal aragonitic plates with smooth outer and zigzag lined inner boundaries. They consist of irregular, parallel (orthogonal), or fan-shaped (clinogonal) arrangements of acicular aragonite crystals. The following taxa are described as new: order Cassianomorpha new order with the family Cassianosomidae new family and the genus Toscanisoma new genus with the species T. multipartitum new species and T. triplicatum new species.UUID: http://zoobank.org/03555353-cdab-42e8-8e99-9bfce15fa249


2010 ◽  
Vol 60 (2) ◽  
pp. 460-468 ◽  
Author(s):  
Miao Miao ◽  
Yangang Wang ◽  
Weibo Song ◽  
John C. Clamp ◽  
Khaled A. S. Al-Rasheid

Recently, an undescribed marine ciliate was isolated from China. Investigation of its morphology and infraciliature revealed it as an undescribed species representing a new genus, Eurystomatella n. gen., the type of the new family Eurystomatellidae n. fam. The new family is defined by close-set, apically positioned oral membranelles and a dominant buccal field that is surrounded by an almost completely circular paroral membrane. The new genus is defined by having a small oral membranelle 1 (M1), bipartite M2 and well-developed M3, a body surface faintly sculptured with a silverline system in a quadrangular, reticulate pattern and a cytostome located at the anterior third of a large buccal field. The type species of the new genus, Eurystomatella sinica n. sp., is a morphologically unique form that is defined mainly by the combination of a conspicuously flattened body, several caudal cilia, extremely long cilia associated with the buccal apparatus and a contractile vacuole located subcaudally. According to phylogenetic analyses of small-subunit (SSU) rRNA gene sequences, Eurystomatella clusters with the genus Cyclidium, as a sister group to the family Pleuronematidae. The great divergence in both buccal and somatic ciliature between Eurystomatella and all other known scuticociliates supports the establishment of a new family for Eurystomatella.


Zootaxa ◽  
2010 ◽  
Vol 2416 (1) ◽  
pp. 61 ◽  
Author(s):  
ELIZABETH DERRYBERRY ◽  
SANTIAGO CLARAMUNT ◽  
KELLY E. O’QUIN ◽  
ALEXANDRE ALEIXO ◽  
R. TERRY CHESSER ◽  
...  

Phylogenetic analysis of the family Furnariidae (Aves: Passeriformes) indicates that the genus Asthenes is polyphyletic, consisting of two groups that are not sister taxa. Pseudasthenes, a new genus of ovenbird, is described for one of these groups. The four species included in the new genus, formerly placed in Asthenes, are P. humicola, P. patagonica, P. steinbachi, and P. cactorum.


2019 ◽  
Vol 33 (6) ◽  
pp. 892
Author(s):  
Adriano B. Kury ◽  
Abel Pérez-González ◽  
Daniel N. Proud

The laniatorean family Phalangodidae has been largely reduced to a core of Holarctic species; however, many taxa were never formally transferred to other families. We examined a group of harvestmen related to Beloniscus Thorell, 1891, from South-east Asia and determined the nature of the relationships of the ‘Beloniscus-like harvestmen’, herein described as the new family Beloniscidae, fam. nov., in the broad context of Laniatores. Based on a molecular phylogenetic analysis of a broad representative laniatorean Sanger-sequences dataset we found support for our taxonomic hypotheses that (1) inclusion of Beloniscidae, fam. nov. in Phalangodidae would render the family non-monophyletic, and (2) Beloniscidae, fam. nov. represents a lineage that is morphologically and genetically distinct from all other known laniatorean families. The new family Beloniscidae is endemic to South-east Asia and comprises 37 species in two new subfamilies: Beloniscinae, subfam. nov. and Buparinae, subfam. nov. Beloniscinae includes the genera Beloniscellus Roewer, 1931, Beloniscops Roewer, 1949, Belonisculus Roewer, 1923, Beloniscus (type genus) and Kendengus Roewer, 1949. Buparinae includes the genera Buparellus Roewer, 1949, Bupares Thorell, 1889 (type genus) and Buparomma Roewer, 1949. Members of Beloniscidae are relatively homogeneous in their external morphology, but are recognised by well marked genitalic features. Male genitalia are described for exemplar species, and the morphology is compared with that of Epedanidae. Names are given to two new genitalic structures: sella curulis and pseudocalyx. http://zoobank.org/urn:lsid:zoobank.org:act:540BCE49-6F2E-4372-BFD4-D3C6068F045D http://zoobank.org/urn:lsid:zoobank.org:act:FE8B011B-AA6D-4E6C-A321-123542A0563F http://zoobank.org/urn:lsid:zoobank.org:act:E4A559CD-170A-40F2-924A-541C22974344


1971 ◽  
Vol 49 (6) ◽  
pp. 869-880 ◽  
Author(s):  
David Malloch ◽  
R. F. Cain

The family Sordariaceae is briefly defined and a list of cleistothecial genera given. Two species of Anixiella Saito & Minoura ex Cain, A. endodonta sp. nov., and A. monospora sp. nov. are described and a key to the four species of the genus included. Apodus deciduus gen. et sp. nov. is described. Two new combinations in Echinopodospora Robison, E. spinosa (Cailleux) Malloch & Cain and E. verruculosa (Cailleux) Malloch & Cain, are proposed to accept transfers from Tripterospora Cain and are followed by a key to species of Echinopodospora. The concept of Zopfiella Winter is broadened to include Z. pleuropora sp. nov. and three new combinations, Z. inermis (Cailleux) Malloch & Cain, Z. latipes (Lundqvist) Malloch & Cain, and Z. leucotricha (Speg.) Malloch & Cain, transferred from Tripterospora. A key to the six species of Zopfiella is presented. The new family Coniochaetaceae is proposed to accommodate the new genus Coniochaetidium and Coniochaeta (Sacc.) Massee. Two species of Coniochaetidium are described, C. ostreum sp. nov. and C. savoryi (Booth) Malloch & Cain, comb, nov., based on Thielavia savoryi Booth.


Author(s):  
Agnieszka Soszyńska-Maj ◽  
Wiesław Krzemiński ◽  
Katarzyna Kopeć ◽  
Robert A. Coram

ABSTRACTA new family, Worcestobiidae fam. nov., is established for two Triassic fossil species of Mecoptera removed from the family Orthophlebiidae: Orthophlebia gigantea Tillyard, 1933 and O. haradai Ueda, 1991. A new genus, Worcestobia gen. nov., is erected and both species are transferred to this genus. The new taxa were established as a result of ongoing taxonomic work on Orthophlebiidae, one of the most problematic families within fossil Mecoptera, considered to be a paraphyletic group of species.


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