A revision of Leydigia Kurz, 1875 (Anomopoda, Cladocera, Branchiopoda), and subgeneric differentiation within the genus

Zootaxa ◽  
2009 ◽  
Vol 2082 (1) ◽  
pp. 1-84 ◽  
Author(s):  
ALEXEY A. KOTOV
Keyword(s):  
Majority Rule ◽  
Type Species ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Species Group ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
Bootstrap Simulation ◽  
Group A ◽  
Exact Identification

A revision of the genus Leydigia Kurz, 1875 (Anomopoda, Cladocera, Branchiopoda) is presented. The list of all species-group nominal taxa consists of 34 published and 3 unpublished names. Of these, 12 species are accepted as valid: (1) Leydigia (Leydigia) leydigi (Schödler, 1863); (2) L. (L.) louisi Jenkin, 1934 with two subspecies L. louisi louisi Jenkin, 1934 and L. louisi mexicana Kotov, Elías-Gutiérrez et Nieto, 2003; (3) Leydigia (Neoleydigia) propinqua Sars, 1903; (4) L. (N.) australis Sars, 1885; (5) L. (N.) microps Sars, 1916; (6) L. (N.) sp. nov. from 'L. acanthocercoides' in Alonso, 1996; (7) L. (N.). macrodonta Sars, 1916; (8) L. (N.) acanthocercoides (Fischer, 1854); (9) L. (N.). laevis Gurney, 1927; (10) L. (N.) cf. ipojucae Brehm, 1939; (11) L. (N.) ciliata Gauthier, 1939; (12) L. (N.) cf. striata Birabén, 1939. Lectotypes are selected for 3, 5, 7, and 9. Exact identification of 10 and 12 is not possible without examination of material from type localities and neotype selection. As confirmed by examination of authors' type material, some taxa (Leydigia africana Gurney, 1904 and Leydigia ankammaraoi Prasad, Santa Kumari et Bose, 1985) prove to be junior synonyms of previously described species; species 8-12 form a compact acanthocercoides-group, with fine differences among members. A cladistic analysis for 13 studied taxa and 18 morphological characters resulted in four most-parsimonious trees (TL = 32; CI = 0.78), which differ only in the grouping of members of the L. acanthocercoides-group. A slightly polytomic strict consensus tree (the 50% majority rule bootstrap simulation led to a tree of similar topology to the contree), as well as some 'orthodox' ideas on the evolution of the genus (not contradicting each other), are used to subdivide the genus into two subgenera, Leydigia (Leydigia) Kurz, 1875 and Leydigia (Neoleydigia) subgen. nov. L. (N.) acanthocercoides is the type species of the latter. A key for the identification of well-known species of Leydigia is provided. The level of description of representatives of the genus Leydigia in different continents is estimated, and perspectives for further investigations are outlined.

A cladistic analysis of the Old World species of Lotus L. (Fabaceae: Loteae)

2000 ◽  
Vol 78 (3) ◽  
pp. 351-360 ◽  
Author(s):  
Ana M Arambarri
Keyword(s):  
Cladistic Analysis ◽  
Morphological Characters ◽  
Comparison Method ◽  
Data Matrix ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
World Species ◽  
Old World ◽  
Species Phylogeny ◽  
The World

The diagnostic characters of the genus Lotus L. are a claw with a thickened infolded margin, diadelphous stamens, and a style hardened from the base. This genus contains about 100 species that are distributed throughout the world. To investigate the phylogeny of the Old World species of Lotus, subgenus Edentolotus, sections Krokeria, Xantholotus, and Erythrolotus, a cladistic analysis was performed using 31 morphological characters. To test the phylogenetic relationships among species of Lotus-Edentolotus and Dorycnium, Pedrosia, and Tetragonolobus, these taxa were included as part of the ingroup. The polarity of the characters was based on the outgroup comparison method, using Anthyllis as one outgroup and Tripodion as another. The analysis with Anthyllis as outgroup yielded eight equally parsimonious trees (with all characters equally weighted), each with 62 steps, a consistency index of 0.53, and a retention index of 0.75. All trees (including the strict consensus tree from the eight initial trees) showed that genus Lotus, subgenus Edentolotus, and sections Xantholotus and Erythrolotus are polyphyletic, with only section Krokeria appearing as monophyletic. On the other hand, the groups of species Lotus angustissimus, Lotus corniculatus, Lotus creticus, and Lotus peregrinus are monophyletic. Identical results were derived from the data matrix using Tripodion as the outgroup. Results are compared with previous cytogenetic and biochemical evidence.Key words: cladistic analysis, Fabaceae, Loteae, Lotus, Old World species, phylogeny.

scholarly journals Phylogenetic relationships within the Pylochelidae (Decapoda: Anomura: Paguroidea): A cladistic analysis based on morphological characters

Zootaxa ◽  
2009 ◽  
Vol 2022 (1) ◽  
pp. 1-14 ◽  
Author(s):  
RAFAEL LEMAITRE ◽  
PATSY A. MCLAUGHLIN ◽  
ULF SORHANNUS
Keyword(s):  
Hermit Crab ◽  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Data Matrix ◽  
Parsimony Analysis ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
The Family

Phylogenetic relationships within the “symmetrical” hermit crab family Pylochelidae were analyzed for 41 of the 45 species and subspecies currently considered valid. In the analyses, 78 morphological characters comprised the data matrix and the outgroup consisted of Thalassina anomala, a member of the Thalassinidae, and Munida quadrispina, a member of the Galatheidae. A poorly resolved strict consensus tree was obtained from a heuristic parsimony analysis of unweighted and unordered characters, which showed the family Pylochelidae and the subfamilies Pylochelinae and Pomatochelinae to be monophyletic taxa – the latter two groups had the highest Bremer support values. Additionally, while the subgenus Pylocheles (Pylocheles) was strongly supported, the subgenera Xylocheles, and Bathycheles were not. More fully resolved trees were obtained when using implied weighting, which recognized the monotypic subfamilies Parapylochelinae, Cancellochelinae and Mixtopagurinae. The subfamily Trizochelinae was found to have four distinct clades and several ambiguously placed taxa.

scholarly journals A phylogenetic analysis of the Gruiformes (Aves) based on morphological characters, with an emphasis on the rails (Rallidae)

1998 ◽  
Vol 353 (1378) ◽  
pp. 2077-2151 ◽  
Author(s):  
B. C. Livezey
Keyword(s):  
Cladistic Analysis ◽  
Morphological Characters ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
Sister Relationship ◽  
Species Complexes ◽  
Basal Group ◽  
Close Relationship ◽  
The Matrix ◽  
Porphyrio Porphyrio

The order Gruiformes, for which even familial composition remains controversial, is perhaps the least well understood avian order from a phylogenetic perspective. The history of the systematics of the order is presented, and the ecological and biogeographic characteristics of its members are summarized. Using cladistic techniques, phylogenetic relationships among fossil and modern genera of the Gruiformes were estimated based on 381 primarily osteological characters; relationships among modern species of Grues (Psophiidae, Aramidae, Gruidae, Heliornithidae and Rallidae) were assessed based on these characters augmented by 189 characters of the definitive integument. A strict consensus tree for 20,000 shortest trees compiled for the matrix of gruiform genera (length = 967, CI = 0.517) revealed a number of nodes common to the solution set, many of which were robust to bootstrapping and had substantial support (Bremer) indices. Robust nodes included those supporting: a sister relationship between the Pedionomidae and Turnicidae; monophyly of the Gruiformes exclusive of the Pedionomidae and Turnicidae; a sister relationship between the Cariamidae and Phorusrhacoidea; a sister relationship between a clade comprising Eurypyga and Messelornis and one comprising Rhynochetos and Aptornis ; monophyly of the Grues (Psophiidae, Aramidae, Gruidae, Heliornithidae and Rallidae); monophyly of a clade (Gruoidea) comprising (in order of increasingly close relationship) Psophia , Aramus , Balearica and other Gruidae, with monophyly of each member in this series confirmed; a sister relationship between the Heliornithidae and Rallidae; and monophyly of the Rallidae exclusive of Himantornis . Autapomorphic divergence was comparatively high for Pedionomus , Eurypyga , Psophia , Himantornis and Fulica ; extreme autapomorphy, much of which is unique for the order, characterized the extinct, flightless Aptornis . In the species–level analysis of modern Grues, special efforts were made to limit the analytical impacts of homoplasy related to flightlessness in a number of rallid lineages. A strict consensus tree of 20,000 shortest trees compiled (length = 1232, CI = 0.463) confirmed the interfamilial relationships resolved in the ordinal analysis and established a number of other, variably supported groups within the Rallidae. Groupings within the Rallidae included: monophyly of Rallidae exclusive of Himantornis and a clade comprising Porphyrio (including Notornis ) and Porphyrula ; a poorly resolved, basal group of genera including Gymnocrex , Habroptila , Eulabeornis , Aramides , Canirallus and Mentocrex ; an intermediate grade comprising Anurolimnas , Amaurolimnas , and Rougetius ; monophyly of two major subdivisions of remaining rallids, one comprising Rallina (paraphyletic), Rallicula , and Sarothrura , and the other comprising the apparently paraphyletic ‘long–billed’ rails (e.g. Pardirallus , Cyanolimnas , Rallus , Gallirallus and Cabalus and a variably resolved clade comprising ‘crakes’ (e.g. Atlantisia , Laterallus and Porzana , waterhens ( Amaurornis ), moorhens ( Gallinula and allied genera) and coots ( Fulica ). Relationships among ‘crakes’ remain poorly resolved; Laterallus may be paraphyletic, and Porzana is evidently polyphyletic and poses substantial challenges for reconciliation with current taxonomy. Relationships among the species of waterhens, moorhens and coots, however, were comparatively well resolved, and exhaustive, fine–scale analyses of several genera ( Grus , Porphyrio , Aramides , Rallus , Laterallus and Fulica ) and species complexes ( Porphyrio porphyrio –group, Gallirallus philippensis –group and Fulica americana –group) revealed additional topological likelihoods. Many nodes shared by a majority of the shortest trees under equal weighting were common to all shortest trees found following one or two iterations of successive weighting of characters. Provisional placements of selected subfossil rallids (e.g. Diaphorapteryx , Aphanapteryx and Capellirallus ) were based on separate heuristic searches using the strict consensus tree for modern rallids as a backbone constraint. These analyses were considered with respect to assessments of robustness, homoplasy related to flightlessness, challenges and importance of fossils in cladistic analysis, previously published studies and biogeography, and an annotated phylogenetic classification of the Gruiformes is proposed.

Phylogeny of the family Spongicolidae (Crustacea: Stenopodidea): evolutionary trend from shallow-water free-living to deep-water sponge-associated habitat

2003 ◽  
Vol 83 (1) ◽  
pp. 119-131 ◽  
Author(s):  
Tomomi Saito ◽  
Masatsune Takeda
Keyword(s):  
Deep Water ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Evolutionary Trend ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
Free Living ◽  
The Family ◽  
Outgroup Species ◽  
Hexactinellid Sponges

The phylogenetic relationships within the family Spongicolidae were examined based on a cladistic analysis of 38 adult morphological characters with reference to two outgroup species of the family Stenopodidae. The strict consensus tree identified Microprosthema as the most basal genus, followed by Paraspongicola, Spongicola and the remaining genera. The Spongicoloides represents the most derived genus among spongicolids. The genera Spongicola, Spongicoloides and Spongiocaris should be redefined, because they formed paraphyletic clades. The cladogram indicates that symbiosis with the hexactinellid sponges is coincident with the loss of gills, exopods on maxillipeds 2 and 3, and with the loss of spination on carapace, pereopods, abdomen, tail fan etc. These losses in the spongicolids are thought to be secondarily derived in relation to their sponge-associated habitat.

A cladistic analysis and reclassification of the tribe Bembicini (Hymenoptera: Crabronidae: Bembicinae)

Zootaxa ◽  
2011 ◽  
Vol 2801 (1) ◽  
pp. 27 ◽  
Author(s):  
PAVEL G. NEMKOV ◽  
MICHAEL OHL
Keyword(s):  
Cladistic Analysis ◽  
Morphological Characters ◽  
Identification Key ◽  
Data Matrix ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
Digger Wasp ◽  
Genus Level

A cladistic analysis of the digger wasp tribe Bembicini based on morphological characters is presented. The underlying data matrix comprises 64 terminal taxa (coded on genus-level) and 54 morphological characters. The resulting strict consensus tree was used as the basis for a revised subtribal classification of the Bembicini. Based on a previously published classification, we herewith propose a number of changes. The subtribe Spheciina Nemkov and Ohl, subtrib. nov. (comprising Ammatomus A. Costa 1859, Kohlia Handlirsch 1895, Sphecius Dahlbom 1843, and Tanyoprymnus Cameron 1905) is removed from Handlirschiina Nemkov and Lelej 1996. The subtribe Stictiellina Bohart and Horning 1971, stat. resurr. (composed of Chilostictia Gillaspy 1983, Glenostictia Gillaspy in Gillaspy, Evans, and Lin 1962, Microstictia Gillaspy 1963, Steniolia Say 1837, Stictiella J. Parker 1917, and Xerostictia Gillaspy 1963) is separated from Bembicina Latreille 1802. The subtribe Argogorytina Nemkov and Lelej 1996 (Argogorytes Ashmead 1899, Neogorytes Bohart in Bohart and Menke 1976, Paraphilanthus Vardy 1995) is synonymized with Exeirina Dalla Torre 1897, syn. nov. Finally, the subtribe Trichogorytina Nemkov and Pulawski 2009 (genus Trichogorytes Rohwer 1912 only) is synonymized with Gorytina Lepeletier de Saint Fargeau 1845, syn. nov. An updated identification key to the subtribes of the Bembicini is provided.

Phylogenetic relationships among families of the order Anomopoda (Crustacea, Branchiopoda, Cladocera)

Zootaxa ◽  
2004 ◽  
Vol 760 (1) ◽  
pp. 1 ◽  
Author(s):  
LOURDES M.A. ELMOOR-LOUREIRO
Keyword(s):  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
Trunk Limbs ◽  
Trunk Limb

The phylogenetic relationships among families of the Order Anomopoda (Crustacea, Branchiopoda, Cladocera) were investigated through a cladistic analysis including 93 characters and 37 terminal taxa (2 as outgroups). The strict consensus tree supported the monophyly of the Anomopoda and its families, and indicated the existence of two main clades: (Moinidae+Daphniidae) and (Dumontidae (Ilyocryptidae+Bosminidae+Radopoda)). The later clade was supported by trunk limb characters, probably related to life associated with the bottom or with macrophytes (lifestyle lost in Bosminidae, but still visible in some of its trunk limbs). Within the Radopoda, the Eurycercoidea was monophyletic, but the monophyly of the Macrothricoidea was not supported.

A new species of Zoniopoda Stål (Orthoptera: Acridoidea: Romaleidae) from Argentina and its phylogenetic position within the genus

Zootaxa ◽  
2011 ◽  
Vol 2913 (1) ◽  
pp. 27 ◽  
Author(s):  
MARTINA E. POCCO ◽  
GONZALO D. RUBIO ◽  
M. MARTA CIGLIANO
Keyword(s):  
New Species ◽  
Cladistic Analysis ◽  
Type Specimen ◽  
Morphological Characters ◽  
Color Pattern ◽  
Species Group ◽  
The Body ◽  
Phylogenetic Position ◽  
Median Carina ◽  
A New Species

A new species of the romaleid grasshopper genus Zoniopoda Stål (Romaleidae: Romaleini) is described and illustrated from the Sierras Chicas of Córdoba Province, central Argentina. A cladistic analysis based on morphological characters indicates that the genus Zoniopoda constitutes a monophyletic group and that Zoniopoda serrana n. sp. must be assigned to Iheringi species group based on synapomorphies of the pronotum and body color. The new species is similar to Z. similis Bruner from Paraguay, Bolivia and Brazil, from which it can be distinguished by the color pattern of the body, shape of the pronotal dorso-median carina and characters of the male terminalia and epiphallus. This paper has been formatted with embedded links to images of the type specimen, maps based on georeferenced specimen data for the genus and an updated key to the species of Zoniopoda available on the Orthoptera Species File (OSF) online (http://orthoptera.speciesfile.org).

scholarly journals Northern Apennines as centre of speciation: a new Verdanus species group (Hemiptera, Cicadomorpha, Cicadellidae) from Italy and its phylogenetic relationships with V. bensoni and the V. limbatellus group

Zootaxa ◽  
2009 ◽  
Vol 2264 (1) ◽  
pp. 1-22 ◽  
Author(s):  
ADALGISA GUGLIELMINO ◽  
CHRISTOPH BÜCKLE
Keyword(s):  
New Taxa ◽  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Species Group ◽  
Northern Apennines ◽  
Dispersal Ability ◽  
Group V ◽  
Central Eurasia ◽  
Limited Dispersal

A small sector of Northern Apennines the Tuscan-Emilian Apennines constitutes an interesting diversity centre of a new Verdanus species group closely related to V. bensoni and the V. limbatellus group. It consists of three species: V. tyrannus sp. nov., V. saurosus sp. nov. and V. rosaurus sp. nov., the latter with two subspecies, V. rosaurus rosaurus ssp. nov. and V. rosaurus rex ssp. nov., which doubtless form a monophyletic group (V. rosaurus group). Data on their distribution, ecology and life cycle are added to their original descriptions. The new taxa live allopatrically in a very restricted area and thus occupy a distribution gap of another species group of Verdanus, the V. abdominalis group, present in Italy in the mountain regions of the Alps and Central and Southern Apennines. A hypothesis of the origin of the new taxa is presented based on the ecological conditions in the Tuscan-Emilian Apennines during the last Postglacial period and on the limited dispersal ability of these normally brachypterous insects. Possible synapomorphic characters and phylogenetic relationships of the new taxa with each other and with V. bensoni (China) and the V. limbatellus group (V. limbatellus (Zetterstedt), V. kyrilli (Emeljanov), V. sichotanus (Anufriev), V. kaszabi (Dlabola)) are discussed and a cladistic analysis is conducted. Comparing V. bensoni and the V. limbatellus group on the one hand and the V. rosaurus group on the other, some morphological characters appear to change often in parallel on the same paths, independently from the phylogenetic hypothesis. Remarkably, within the same morphological characters the range of variation among species inhabiting the comparatively minute area of the Tuscan-Emilian Apennines is similar to that found among other taxa distributed across vast areas of northern and central Eurasia.

Morphotaxonomic revision of Simulium (Gomphostilbia) (Diptera: Simuliidae) in the Oriental Region

Zootaxa ◽  
2012 ◽  
Vol 3577 (1) ◽  
pp. 1 ◽  
Author(s):  
HIROYUKI TAKAOKA
Keyword(s):  
Geographical Distribution ◽  
Classification Scheme ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Species Group ◽  
Morphological Identification ◽  
Oriental Region ◽  
Preliminary Attempt ◽  
The Third ◽  
Species Groups

Simulium (Gomphostilbia) Enderlein, the third largest (206 named species included) in the genus Simulium Latreille s. l., is one of the two most abundant and diverse subgenera in the Oriental Region. To provide a classification scheme to facilitate morphological identification of the species within this subgenus, its diagnostic characters are redefined, and nine known species-groups within it are reviewed. Based on putative lineages explored by using certain adult morphological characters, seven more species-groups are proposed: asakoae, darjeelingense, epistum, gombakense, heldsbachense, hemicyclium and palauense species-groups, while the trirugosum species-group is merged in the varicorne species-group. Subgroups are also introduced to represent apparently different lineages within certain species-groups based on certain pupal morphological characters: two in the banauense species-group, seven in the batoense species-group redefined, four in the ceylonicum species-group redefined, four in the epistum species-group, two in the hemicyclium species-group, two in the sherwoodi species-group and four in the varicorne species-group redefined. A new checklist of species of the subgenus Gomphostilbia, and a key to all 15 species-groups within it are provided. The eastward expansion of the geographical distribution of the subgenus Gomphostilbia is inferred on the basis of the more frequent occurrence of apomorphic characters of certain adult and pupal morphological features in insular species-groups than in continental species-groups. A preliminary attempt using a cladistic analysis of morphological characters shows that among 10 subgenera examined, Gomphostilbia has a sister-taxon relationship with the Australasian subgenus Morops Enderlein, and this clade, together with the Central-Western Pacific subgenus Inseliellum Rubtsov, is positioned closest to the most derived clade formed by Daviesellum Takaoka & Adler and Simulium Latreille s. str.

Characterisation of the Australian Nacophorini using adult morphology, and phylogeny of the Geometridae based on morphological characters

Zootaxa ◽  
2008 ◽  
Vol 1736 (1) ◽  
pp. 1 ◽  
Author(s):  
CATHERINE J. YOUNG
Keyword(s):  
Phylogenetic Analysis ◽  
Majority Rule ◽  
Morphological Character ◽  
Morphological Characters ◽  
Data Sources ◽  
Morphological Data ◽  
Bootstrap Support ◽  
Consensus Tree ◽  
Molecular Analyses ◽  
Character Set

The Australian Nacophorini and related taxa are described using a matrix of 116 adult morphological characters. Adults of 72 species are illustrated using photographs and electron micrographs. Subsets of the characters are used to conduct a phylogenetic analysis based on cladistic principles. The adult morphological character set was augmented with 17 characters from eggs and 27 from larvae. The resulting phylogeny is poorly resolved but provides support for many of the relationships recovered by previous molecular analyses of the group, including basal derivations for characters of Larentiinae and Sterrhinae relative to those of the rest of Geometridae, and the monophyly of the Geometrinae + Oenochrominae s. str. Combining 28S D2 datawith morphological data produced a matrix of 60 taxa and 590 characters. The majority rule consensus tree produced by the combined morphological and 28S D2 data is almost identical to the majority rule consensus tree produced by the 28S D2 data alone, except that bootstrap support is lower for most nodes. Common clades obtained from the molecular and morphological trees are described in terms of morphological data. On this basis a concept of the Australian Nacophorini includes Lithinini and Australian Archiearinae. Two robust groups within the tribe also are delimited using characters from all data sources. Comparsions are made between the Nearctic and Neotropical Nacophorini on the basis of shared morphological characters. Australian Boarmiini are defined by synapomorphies.

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