Taxonomy of coccids (Hemiptera: Coccidae: Coccus L.) associated with Crematogaster ants (Hymenoptera: Formicidae) in the stems of Macaranga plants (Euphorbiaceae) in Southeast Asia

Zootaxa ◽  
2018 ◽  
Vol 4521 (1) ◽  
pp. 1 ◽  
Author(s):  
PENNY J. GULLAN ◽  
TAKUMASA KONDO ◽  
BRIGITTE FIALA ◽  
SWEE-PECK QUEK
Keyword(s):  
New Species ◽  
Southeast Asia ◽  
Type Species ◽  
Instar Nymph ◽  
Scale Insects ◽  
Soft Scale ◽  
Adult Females ◽  
First Instar ◽  
Morphological Differences ◽  
Three New Species

The Southeast Asian soft scale insects (Hemiptera: Coccomorpha: Coccidae) associated with ants of the Crematogaster borneensis-group (Hymenoptera: Formicidae) and living in the hollow stems of Macaranga plants (Euphorbiaceae) are revised taxonomically. Ten species of the genus Coccus L. are recognised: seven were described previously and three new species are described herein. The species are: Coccus caviramicolus Morrison, C. circularis Morrison, C. heckrothi Gullan & Kondo sp. n., C. lambirensis Gullan & Kondo sp. n., C. macarangicolus Takahashi, C. macarangae Morrison, C. penangensis Morrison, C. pseudotumuliferus Gullan & Kondo sp. n., C. secretus Morrison and C. tumuliferus Morrison. All of these species are described or redescribed and newly illustrated based on the adult females, and a key to distinguish the species is provided. We designate a lectotype for C. macarangicolus. The first-instar nymphs of all species are morphologically extremely similar and therefore only the first-instar nymph of C. macarangae is described and illustrated. Seven of these species currently are known only from Macaranga, but C. macarangae, C. secretus and perhaps C. pseudotumuliferus have been recorded from the hollow stems of several other ant-plants and a few non-myrmecophytes. The Coccus species from Macaranga are closely related to C. hesperidum L., the type species of the genus, and therefore are retained in the genus Coccus even though the adult females exhibit a few morphological differences from C. hesperidum. The species of Coccus from Macaranga appear to be parthenogenetic because no male nymphs or adults have been found, despite extensive collecting. 

Taxonomic revision of the myrmecophilous, meliponiphilous and rhizophilous soft scale genus Cryptostigma Ferris (Hemiptera: Coccoidea: Coccidae)

Zootaxa ◽  
2019 ◽  
Vol 2709 (1) ◽  
pp. 1 ◽  
Author(s):  
TAKUMASA KONDO
Keyword(s):  
New Species ◽  
Host Plants ◽  
New World ◽  
Taxonomic Revision ◽  
Scale Insects ◽  
Biological Information ◽  
World Species ◽  
Soft Scale ◽  
Adult Females ◽  
First Instar

The present study revises the soft scale insects of the genus Cryptostigma Ferris (Hemiptera: Coccoidea: Coccidae), which comprises a group of New World species associated with ants or bees or of hypogeal habit. It includes 17 species, of which 10 species are new to science. The study is based on the external, mostly cuticular morphology of the adult females and first-instar nymphs. Taxonomic keys based on adult females and known first-instar nymphs are provided. For each species a description or redescription is given, followed by information on its distribution, host plants, and known biological information including associated ants or bees. The new species described are Cryptostigma chacoensis sp. nov., C. gullanae sp. nov., C. jonmartini sp. nov., C. longinoi sp. nov., C. melissophilum sp. nov., C. mexicanum sp. nov., C. philwardi , sp. nov. C. rhizophilum sp. nov., C. serratum sp. nov., and C. tuberculosum sp. nov. Neolecanium urichi (Cockerell) is transferred to Cryptostigma as C. urichi (Cockerell) comb. nov., and C. quinquepori (Newstead) is synonymized with Cryptostigma urichi, comb. nov. Lectotypes are designated for Cryptostigma saundersi Laing, Lecanium silveirai Hempel, Lecanium urichi Cockerell and Akermes quinquepori Newstead. Cryptostigma inquilina (Newstead) is amended to C. inquilinum in order to match the neutral ending “stigma”. The following names are treated as nomina nuda: Cryptostigma jamaicensis and Lecanopsis jamaicensis (Ben-Dov, 1993: 97).

Taxonomy of Toumeyella lomagundiae Hall and T. obunca De Lotto, and their transfer to Hallicoccus gen. nov. (Hemiptera: Coccoidea: Coccidae)

Zootaxa ◽  
2007 ◽  
Vol 1415 (1) ◽  
pp. 23-33
Author(s):  
TAKUMASA KONDO
Keyword(s):  
Instar Nymph ◽  
Scale Insects ◽  
Soft Scale ◽  
Taxonomic Key ◽  
Adult Females ◽  
First Instar

Two African soft scale insects, Toumeyella lomagundiae Hall and T. obunca De Lotto, are transferred to Hallicoccus gen. nov. The adult females of both species are redescribed, and the first-instar nymph of T. lomagundiae is described. A revised taxonomic key to separate the adult females is provided. The affinity of Hallicoccus gen. nov. with the genus Toumeyella Cockerell is briefly discussed.

Description of a new pest species of Hemilecanium Newstead (Hemiptera: Coccidae) on mango from Thailand, and a key to species of the genus

Zootaxa ◽  
2005 ◽  
Vol 1045 (1) ◽  
pp. 25 ◽  
Author(s):  
TAKUMASA KONDO ◽  
MICHAEL L. WILLIAMS
Keyword(s):  
Mangifera Indica ◽  
Scale Insect ◽  
Instar Nymph ◽  
Pest Species ◽  
Key To Species ◽  
Soft Scale ◽  
Adult Females ◽  
First Instar ◽  
A New Species ◽  
New Pest

The adult female and first-instar nymph of a new species of soft scale insect, Hemilecanium mangiferae Kondo & Williams sp. nov., are described and illustrated. This species was collected on the trunk and branches of mango, Mangifera indica L., in Nakhon Pathom and Sukhothai Provinces, Thailand. The insect can cause serious damage by depleting the host sap and by producing large amounts of honeydew on which sooty molds grow. Damage was severe on the local mango cultivar, Nam DokMai. Also included are a revised diagnosis of the genus, a revised key to the adult females of the five species currently placed in the genus, and a key to the known firstinstar nymphs.La hembra adulta y la ninfa del primer estadio de una nueva especie de escama blanda, Hemilecanium mangiferae Kondo & Williams sp. nov., se describe e ilustra. Esta especie fue colectada en el tronco y ramas del mango: Mangifera indica L. en las provincias de Nakhon Pathom y Sukhothai, Tailandia. Este insecto puede causar daños severos al mango deprivando el árbol de savia, y produciendo gran cantidad de melado cual induce el crecimiento de fumaginas. El daño fue más severo en un cultivar local conocido como Nam Dokmai. Tambien se incluyen una diagnosis revisada para el género, una clave revisada para las hembras adultas de las cinco especies actualmente incluidas en el género, más una clave para las ninfas del primer estadio conocidas.

Cissococcus Cockerell (Hemiptera: Coccoidea: Coccidae), a unique gall-inducing soft scale genus on Vitaceae from South Africa, with description of a new species

Zootaxa ◽  
2011 ◽  
Vol 2996 (1) ◽  
pp. 1 ◽  
Author(s):  
C. J. HODGSON ◽  
I. M. MILLAR ◽  
P. J. GULLAN
Keyword(s):  
South Africa ◽  
Type Species ◽  
Scale Insect ◽  
Instar Nymph ◽  
Closely Related Species ◽  
Soft Scale ◽  
First Instar ◽  
The Family ◽  
Gall Induction ◽  
A New Species

The gall-inducing genus Cissococcus Cockerell is revised to include two species. The family placement of Cissococcus has been uncertain, but characters of the first-instar nymph and adult male clearly show that Cissococcus is a soft scale insect (Coccidae) and is therefore the only member of the Coccidae known to induce a complex covering gall. All stages of the type species, C. fulleri Cockerell, are described and illustrated, plus the adult and third-instar female and first-instar nymph of a new, closely-related species, C. braini Hodgson & Millar sp. n. Both species are known only from wild vines in the southeast and C. fulleri also in the northeast of South Africa, but each induces a unique gall and each appears to be restricted to a different species of Rhoicissus (Vitaceae). The galls of each species are described and figured, and gall induction in the Coccidae is briefly discussed.

Neogreenia lonicera sp. nov., a new species of Margarodidae sensu lato (Hemiptera: Coccoidea) from China, with a key to species of Neogreenia MacGillivray and placement of the genus in the family Kuwaniidae

Zootaxa ◽  
2012 ◽  
Vol 3274 (1) ◽  
pp. 43 ◽  
Author(s):  
SAN’AN WU ◽  
NAN NAN
Keyword(s):  
New Species ◽  
Cladistic Analysis ◽  
Instar Nymph ◽  
Morphological Data ◽  
Key To Species ◽  
Adult Females ◽  
First Instar ◽  
The Family ◽  
Helan Mountain ◽  
A New Species

A new species, Neogreenia lonicera Wu & Nan, is described and illustrated based on the adult female, second-instarfemale and first-instar nymph. This new species was collected at Helan Mountain, Inner Mongolia, China, in bark crevicesof Lonicera microphylla (Caprifoliaceae). A key is provided to separate the five species now known in NeogreeniaMacGillivray. A cladistic analysis of morphological data from adult females and first-instar nymphs of 28 archaeococcoidgenera has Neogreenia in a clade with Jansenus Foldi and Neosteingelia Morrison and usually also with KuwaniaCockerell, and thus Neogreenia should be placed in the family Kuwaniidae. A key to distinguish the adult females of the four genera of the Kuwaniidae is provided.

On some new and old soft scale insects (Hemiptera: Coccomorpha: Coccidae) from Africa, with description of a new Coccus species and introduction of a new genus 

Zootaxa ◽  
2019 ◽  
Vol 4612 (3) ◽  
pp. 373
Author(s):  
BOŻENA ŁAGOWSKA ◽  
CHRIS J. HODGSON
Keyword(s):  
South Africa ◽  
South African ◽  
Type Species ◽  
New Genus ◽  
Scale Insects ◽  
Similar Species ◽  
Soft Scale ◽  
Adult Females ◽  
First Time ◽  
A New Species

A new species of soft scale (Hemiptera: Coccomorpha; Cocccidae) from South Africa, Coccus giliomeei Łagowska & Hodgson sp. n., collected on Gymnosporia buxifolia (L.) Szyszyl, is described and illustrated. Also, Coccus rhodesiensis (Hall) is recorded for the first time from South Africa and is redescribed and illustrated based on the adult females of the type series and fresh South African specimens. An updated key to the species of Coccus and similar species known from Africa is included. Based on this latter study, (i) Marsipococcus proteae (Brain) and M. durbanensis (Brain) are considered not to be congeneric with Marsipococcus marsupialis (Green), the type species of Marsipococcus Cockerell & Bueker, and are placed in a new genus Proteacoccus Łagowska & Hodgson, gen. n. with Lecanium proteae Brain as the type species; (ii) it is considered that Coccus asiaticus Lindinger is clearly not a junior synonym of Parasaissetia nigra (Nietner) and is accepted as a full species, rev. stat., and (iii) Neoplatylecanium adersi (Newstead) is considered to be non-conspecific with N. cinnamomi Takahashi, the type species of Neoplatylecanium Takahashi, and is transferred to Maacoccus Tao & Wong, as Maacoccus adersi (Newstead), comb. n. 

A new species of Marsipococcus Cockerell & Bueker (Hemiptera: Coccomorpha: Coccidae) from Malaysia

Zootaxa ◽  
2017 ◽  
Vol 4358 (2) ◽  
pp. 295
Author(s):  
BOZENA ŁAGOWSKA ◽  
JON H. MARTIN ◽  
CHRIS J. HODGSON
Keyword(s):  
New Species ◽  
Adult Female ◽  
Adult Male ◽  
Instar Nymph ◽  
Adult Females ◽  
First Instar ◽  
Female Pupa ◽  
A New Species

The adult female, first-instar nymph, second-instar male, third-instar female, pupa and adult male of a new species of Marsipococcus Cockerell & Bueker (Hemiptera: Coccomorpha: Coccidae) from Malaysia, M. ulubendulensis Łagowska & Martin sp. n., are described and illustrated, while the second-instar female is only described. Marsipococcus is rediagnosed and a key to the adult females of the four species now placed in this genus is included. 

Scale insect fauna (Hemiptera: Sternorrhyncha: Coccoidea) of New Zealand’s pygmy mistletoes (Korthalsella: Viscaceae) with description of three new species: Leucaspis albotecta, L. trilobata (Diaspididae) and Eriococcus korthalsellae (Eriococcidae)

Zootaxa ◽  
2010 ◽  
Vol 2644 (1) ◽  
pp. 1 ◽  
Author(s):  
ROSA C. HENDERSON ◽  
AMIR SULTAN ◽  
ALASTAIR W. ROBERTSON
Keyword(s):  
New Species ◽  
New Zealand ◽  
Adult Female ◽  
Scale Insect ◽  
Instar Nymph ◽  
Scale Insects ◽  
Insect Fauna ◽  
Male And Female ◽  
Black Scale ◽  
Three New Species

The scale insect fauna recorded on the pigmy mistletoe genus Korthalsella (Viscaceae) and New Zealand’s loranthaceous mistletoes is reviewed. Three new species from New Zealand discovered on Korthalsella species during this study are described: the adult female, 2nd-instar male and female nymphs of two pupillarial species Leucaspis albotecta sp. nov. and Leucaspis trilobata sp. nov. (Diaspididae); and the adult female, 2nd-instar male and female nymphs and 1st-instar nymph of Eriococcus korthalsellae sp. nov. (Eriococcidae). A distribution map is provided for the three new species. We report the first records of adventive (exotic) scale insects on Korthalsella species in New Zealand – Ceroplastes sinensis Del Guercio, Coccus hesperidum Linnaeus, soft brown scale, Saissetia coffeae (Walker), hemispherical scale, S. oleae (Olivier), black scale (Coccidae), and Aspidiotus nerii Bouché, oleander scale, Hemiberlesia lataniae (Signoret), latania scale, H. rapax (Comstock), greedy scale (Diaspididae), while three native scale insects are also first records: Kalasiris perforata (Maskell) (Coccidae), Eriococcus pallidus Maskell (Eriococcidae) and Paracoccus glaucus (Maskell) (Pseudococcidae). Ten scale insect species are listed for New Zealand’s loranthaceous mistletoes.

Redescription of the genus Chlamydolecanium Goux, 1933 (Hemiptera: Coccoidea: Coccidae) with a discussion of its affinities

Zootaxa ◽  
2011 ◽  
Vol 2965 (1) ◽  
pp. 1
Author(s):  
ISABELLE M. VEA
Keyword(s):  
Type Species ◽  
Original Description ◽  
Sister Group ◽  
Morphological Characters ◽  
Instar Nymph ◽  
Soft Scale ◽  
Incertae Sedis ◽  
Adult Females ◽  
Coccus Hesperidum ◽  
Group Relationships

The monotypic genus Chlamydolecanium Goux (Hemiptera: Coccoidea: Coccidae) is known only from the original collection in Corsica. The original description does not allow definite placement in any of the subfamilies in the Coccidae. Although it had been suggested that it belonged to the Cardiococcinae, no other members of this subfamily were geographically close. The adult female and 1 st -instar nymph of the type species, Chlamydolecanium conchioides Goux are here redescribed. A parsimony analysis was undertaken based on 56 morphological characters of the adult females in 29 taxa (24 genera considered to belong to the Cardiococcinae, plus Parafairmaria bipartita (Signoret), an incertae sedis monospecific genus possibly close to Cardiococcinae, plus the recently described Kenima galilit Ben-Dov plus two other soft scale genera as outgroups (Coccus hesperidum L. and Paralecanium frenchii (Maskell)). This analysis confirmed the placement of the genus Chlamydolecanium within the Cardiococcinae clade, although its sister-group relationships remain unresolved. Its geographic distribution and taxonomic relationships are discussed.

Description of a new species of Kermicus Newstead (Hemiptera: Coccomorpha: Pseudococcidae) from bamboo in southeast China

Zootaxa ◽  
2020 ◽  
Vol 4859 (3) ◽  
pp. 440-450
Author(s):  
SAN-AN WU ◽  
SHAOBIN HUANG ◽  
CHUANGUAN LIANG
Keyword(s):  
New Species ◽  
Adult Female ◽  
Guangdong Province ◽  
Instar Nymph ◽  
Southeast China ◽  
Adult Females ◽  
First Instar ◽  
Anal Ring ◽  
A New Species ◽  
Marginal Region

A new species, Kermicus huizhouensis Wu & Huang sp. n. (Hemiptera: Coccomorpha: Pseudococcidae), is described and illustrated based on the adult female, second instar female and first-instar nymph. It was collected at Qianfeng village, Huiyang District, Huizhou City, Guangdong Province, P. R. China, inside the stem of bamboo Bambusa rigida (Poaceae) and attended by the ant Tetraponera binghami (Forel) (Hymenoptera: Formicidae). It differs from K. wroughtoni Newstead, 1897 by the adult female lacking multilocular disc pores in the marginal region of the venter, and both the adult female and first-instar nymph having the anal ring bearing 16–20 setae. Keys to genera of legless mealybugs on bamboo based on adult females, and the immature stages of Kermicus, are provided. 

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