Revision of the Early Ordovician (late Tremadocian; Stairsian) cheirurid trilobite Tesselacauda Ross, with species from the Great Basin, western USA

Zootaxa ◽  
2019 ◽  
Vol 4661 (2) ◽  
pp. 201-255
Author(s):  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM

The Stairsian genus Tesselacauda Ross, 1951, has historically included two species, the poorly known type, T. depressa Ross, 1951 (Bearriverops loganensis Zone), and the even less well known T. flabella Kobayashi, 1955 (Bearriverops alsacharovi Zone), which may not belong to the genus. The family assignment of the genus has long been in question, with some workers assigning it to Cheiruridae and some to Pliomeridae. New field collections from western Utah and southeastern Idaho yield abundant material of T. depressa, which facilitates revision on the basis of multiple specimens of most exoskeletal parts. Two additional well known species are proposed, T. morrisoni (Rossaspis leboni Zone), and T. kriegerae (Bearriverops alsacharovi Zone). A third new species, very similar to T. depressa, is described in open nomenclature from the Rossaspis leboni Zone. Knowledge of hypostomes from silicified material helps to clarify the basal morphologies of cheirurid versus pliomerid trilobites. Pliomerids have anteroposteriorly elongate hypostomes with a basic pattern of three pairs of lateral hypostomal spines and a single posteromedian spine. Some or all of the spines are reduced or lost in various taxa. Cheirurids either lack paired spines or have only one or two pairs, and never have a posteromedian spine. Cheirurid hypostomes tend to be much shorter and more subquadrate than pliomerids. Other differences between the families are: a small, triangular or trapezoidal rostral plate in pliomerids versus a wide, short plate in cheirurids; a thoracic segment count commonly of 11–13 in Cheiruridae (fewer in one derived subfamily) versus commonly 15 or more in pliomerids (fewer in two derived subfamilies); thoracic pleurae with subequal bands and a prominent furrow in cheirurids versus a much more inflated and rib-like posterior band, reduced anterior band, and short, anteriorly placed furrow; and pygidia with four or fewer segments in cheirurids versus commonly five in pliomerids (again, fewer in two derived subfamilies). On these and other criteria, Tesselacauda is clearly a cheirurid, assigned for the present to the presumptively basal and possibly paraphyletic Subfamily Pilekiinae.

2014 ◽  
Vol 88 (6) ◽  
pp. 1163-1173 ◽  
Author(s):  
Daniel B. Blake

Phragmactis grayaeSpencer andSwataria derstlerinew genus new species are early (Ordovician) asterozoans (Echinodermata) that comprise the Phragmactinidae. Asterozoans are complexly varied, but as is true for other echinoderms, ambulacral construction is critical to interpretation. Phragmactinids share plesiomorphic aspects of ambulacral form and articulation with basal somasteroids and stenuroids whereas the apomorphic ambulacral expressions of asteroids and ophiuroids are lacking. Phragmactinids, like asteroids and ophiuroids, have only one virgal-series ossicle associated with each ambulacral, unlike the multiple ossicles of somasteroids and stenuroids. Virgal morphology of phragmactinids is reminiscent of expressions in somasteroids and stenuroids. Aspects of phragmactinid mouth frame construction are apomorphic. Morphologies of other ossicular series are similarly varied, and as a result, the family cannot be easily fitted into a recently proposed class-level taxonomy of early asterozoans; it is left in open nomenclature. Phragmactinid morphology does not indicate behavior significantly different from that of other early asterozaons. Asterozoan diversity suggests an early period of rapid evolutionary radiation.


1999 ◽  
Vol 73 (5) ◽  
pp. 945-951 ◽  
Author(s):  
William W. Korth

A new genus of mylagaluid rodent from the Miocene of the Great Basin, Hesperogaulus, is described, and to it two new species, H. gazini and H. wilsoni (type species) are referred. This genus differs from all other genera of later Tertiary mylagaulines in having: large bosses of rugose bone on the anterior end of the nasal bones; lower slope of the occipital bone; and unique separation of the anterior branches of the forked parafossette on the upper premolar.Hepserogaulus n. gen. represents a distinct lineage of mylagaulids that has been separate from other members of the family since the early Barstovian. This genus is geographically limited to the Great Basin. Other later Tertiary genera of mylagaulids show geographic limitations to other parts of the continent as well.


Zootaxa ◽  
2011 ◽  
Vol 2969 (1) ◽  
pp. 1 ◽  
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN

Panisaspis n. gen. is a clade of pliomerid trilobites from the Tulean and Blackhillsian stages (Floian) of the Great Basin.  It includes Protopliomerops? quattuor Hintze, 1953, and ten new species, six of which are formally named: Panisaspis millardensis (type species), P. sevierensis, P. deltaensis, P. rancherensis, P. topscityensis, and P. loganensis.  Four new species are not well enough known for formal naming and are described in open nomenclature.  All species are Tulean in age except for P. millardensis, which is earliest Blackhillsian.  Synapomorphies of Panisaspis include a short, nearly semicircular anterior border; small L1; large genal spines; a rounded, ovoid hypostomal border; elongated third pygidial spines; and a large, triangular terminal piece with distinct pitted impressions.  Phylogenetic analysis indicates that P. millardensis and P. sevierensis are sister taxa, and that P. deltaensis, P. rancherensis, P. topscityensis, P. quattuor, and P. loganensis are successive sister species.  The group may be sister to Ibexaspis Přibyl and Vaněk in Přibyl et al., 1985. 


Zootaxa ◽  
2018 ◽  
Vol 4525 (1) ◽  
pp. 1
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM

Field-based revision and phylogenetic analysis demonstrate that the pliomerid trilobite taxon Ibexaspis Přibyl and Vaněk (in Přibyl et al., 1985), previously known from a single formally named species (I. brevis [Young, 1973]), belongs to a complex of 14 mostly newly discovered, related species from the Early Ordovician (Floian; Tulean and Blackhillsian) of the northern Laurentian margin. The species are known from silicified samples recovered from sections in eastern Nevada, western Utah, and southeastern Idaho. The stratigraphically early Tuleaspis n. gen. (type species: T. jeneki n. sp.; Tulean; low Protopliomerella contracta Zone) includes its type and two species described in open nomenclature. Tuleaspis is sister to the remainder of the clade. Ibexaspis now includes three additional species: I. coadyi n. sp. (Blackhillsian; Carolinites nevadensis Zone), I. leuppi n. sp. (Blackhillsian; Presbynileus ibexensis Zone), and I. rupauli n. sp. (Blackhillsian; "Pseudocybele nasuta Zone"). Ibexapsis is sister to a clade of Millardaspis n. gen. + Deltapliomera n. gen. Millardaspis (type species M. milsteadi n. sp.; Tulean; Heckethornia hyndeae Zone) also includes M. knoxi n. sp. (Tulean; Panisaspis sevierensis Zone). Deltapliomera (type species D. humphriesi n. sp.; Blackhillsian, Carolinites nevadensis Zone) also includes D. inglei n. sp. (Tulean; Heckethornia bowiei Zone), D. heimbergi (Tulean; Panisaspis sevierensis Zone), D. eppersoni n. sp. (Blackhillsian; Bathyurina plicolabeona Zone), and a species described in open nomenclature. 


2021 ◽  
pp. 1-26
Author(s):  
Jonathan M. Adrain ◽  
Talia S. Karim

Abstract The Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi has long been known from a small number of pygidia assigned to a single formally named species, and its affinities have not been assessed. Silicified material from western Utah and southeastern Idaho includes six distinct species assigned to the genus, one of which is the type species. Two others (G. moffitti and G. pankowskii) are new and formally named. An additional three species that are clearly new but known from sparse material are described in open nomenclature. Gonioteloides has a stratigraphic distribution through five consecutive trilobite zones in the mid-Stairsian Stage (upper Tremadocian). Although exoskeletal morphology of three species is almost completely known, the phylogenetic affinity of the taxon remains difficult to determine. It is tentatively assigned to Dimeropygidae Hupé. UUID: http://zoobank.org/23257d6c-262b-4ef5-ae4e-cc431777e67e


2009 ◽  
Vol 46 (12) ◽  
pp. 875-914 ◽  
Author(s):  
Neo E.B. McAdams ◽  
Jonathan M. Adrain

Heckethornia n. gen. is a morphologically striking clade of dimeropygid trilobites known from the Pogonip Group (Ordovician; Ibexian Series, Tulean and Blackhillsian stages) of western Utah and eastern Nevada. It includes seven species based on silicified material. All occur in the Fillmore Formation of western Utah, and four are also known from the Yellow Hill Limestone of eastern Nevada. Diagnostic features of Heckethornia include a highly vaulted exoskeleton with a tall pygidial “wall” made of fused outer pleurae, three pairs of large glabellar spines, two to three pairs of prominent pygidial spines, a single (or paired) large occipital spine(s), and an arc of tubercles on the librigenal field. Cladistic parsimony analysis suggests that the genus comprises two subclades, one including species with a single median occipital spine, and species with paired occipital spines or tubercles in the other. New species are H. smithi , H. hyndeae , H. numani , H. bowiei , H. morrisseyi , and H. ballionae .


Zootaxa ◽  
2011 ◽  
Vol 3144 (1) ◽  
pp. 1 ◽  
Author(s):  
NEO E. B. MCADAMS ◽  
JONATHAN M. ADRAIN

New field sampling of classic Ibexian sections in the Great Basin has resulted in the discovery of several new species assignable to the pliomerid trilobite genus Protopliomerella Harrington, 1957. The genus was originally monotypic and based on Protopliomerops contracta Ross, 1951. Protopliomerella pauca Demeter, 1973, was later added, but it is shown herein to be a junior subjective synonym of P. contracta. The material originally assigned to Protopliomerella contracta by Ross belongs to as many as three distinct, stratigraphically separate species. New species include P. stegneri, P. bowlesi, P. kerouaci, P. seegeri, and P. okeeffeae. Two additional new species are not well enough known to name and are reported in open nomenclature. The phylogenetic status of Protopliomerella, as thus conceived, is ambiguous and the taxon may be rendered paraphyletic by the genera Pseudocybele Ross, 1951, and Lemureops McAdams and Adrain, 2009.


2017 ◽  
Vol 186 (1) ◽  
pp. 113-116
Author(s):  
František Šifner

Abstract A new Nearctic species of the genus Coniosternum Becker, 1894, C. masneri sp. nov., is described from Canada, and its important diagnostic characters are illustrated.


2013 ◽  
Vol 22 (2) ◽  
pp. 204-223
Author(s):  
S.Yu. Storozhenko

Seven new species of the genus Zhengitettix Liang, 1994 are described: Z. hosticus sp. nov., Z. mucronatus sp. nov. and Z. spinulentus sp. nov. from Vietnam; Z. albitarsus sp. nov. and Z. extraneus sp. nov. from Thailand; Z. palawanensis sp. nov. and Z. taytayensis sp. nov. from the Philippines. Two species, Z. curvispinus Liang, Jiang et Liu, 2007 and Z. obliquespicula Zheng et Jiang, 2005 are firstly recorded from Vietnam. An annotated check-list and key to species of the genus Zhengitettix are given. Position of Zhengitettix within the family Tetrigidae is briefly discussed.


2012 ◽  
Vol 21 (1) ◽  
pp. 131-144 ◽  
Author(s):  
A.G. Kirejtshuk ◽  
A. Nel

In the paper two new species of the genus Rhyzobius Stephens, 1829 (R. antiquus sp. nov. and R. gratiosus sp. nov.) and one new species of the genus Nephus Mulsant 1846 (N. subcircularis sp. nov. without a certain subgeneric placement) from the Lowermost Eocene amber of Oise are described. A short review of known fossil records of the family Coccinellidae is given.


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