New fossil assemblages from the Early Ordovician Fezouata Biota

The Fezouata Biota (Morocco) is an exceptionally well-preserved fossil community of Early Ordovician age and although its oldest units are comparable with Burgess Shale-type localities of the Cambrian Explosion, little attention has been paid to the younger units despite potential to reveal the conditions of the Ordovician Radiation. Herein, we describe a new middle to upper Floian Fezouata locality (Taichoute) encompassing an assemblage dominated by large bivalved euarthropods and giant filter-feeding radiodonts, which were transported and preserved in concretions associated with density-flow deposits. Taichoute captures the closing of the taphonomic window that characterizes exceptional fossil preservation during the Cambrian Explosion (i.e., carbonaceous compressions) as well as the faunal transition to assemblages dominated by typical Palaeozoic taxa.

The identity and significance of the high-latitude Early Ordovician Mediterranean brachiopod Province

During the Early Ordovician Epoch, the Mediterranean brachiopod Province was extensive in the higher-latitude sectors of the globe in the Southern Hemisphere. The latter was much occupied by the massive continent of Gondwana, which stretched from north of the Equator S-wards to cover the South Pole. The Mediterranean Province can be separated into two groups: Group 1, the higher-latitude fauna dominated by large linguliform brachiopods; and Group 2, which is more diverse, particularly in orthides. The large linguliform brachiopod faunas are particularly well known in southern Europe (France, Spain and Bohemia) and North Africa, and the second group in Avalonia, Chile and Argentina. The province is different from, but merges with, more diverse contemporary faunas in the lower latitudes of Gondwana to its north, although the latter contrast with other lower-latitude faunal provinces in South China, Laurentia, Siberia and elsewhere. Since the Rheic Ocean between Avalonia and Gondwana was relatively narrow during the Early Ordovician Epoch, the Avalonian brachiopods were integral parts of the Mediterranean Province, but only until end of the Dapingian Age. This paper focuses on the earlier phases of the Mediterranean Province, although the province continued until near the end of the Ordovician Period. Intermediate-latitude Baltica and some other faunas are included in new principal components and other analyses in order to compare them with the Mediterranean Province faunas. Radiation was very significant for many brachiopod taxa during the period, with first appearances of the Plectambonitoidea (Taffiidae), several orthide families (Euorthisinidae, Tarfayidae and Anamalorthidae) and the earliest endopunctate orthide, the dalmanelloid Lipanorthis.

Trepostome bryozoans buck the trend and ignore calcite-aragonite seas

Trepostome bryozoan skeletalisation did not passively respond to changes in seawater chemistry associated with calcite-aragonite seas. According to Stanley and others, trepostome bryozoans were passive hypercalcifiers. However, if this was the case, we would expect their degree of calcitic colony calcification to have decreased across the Calcite I Sea to the Aragonite II Sea at its transition in the Middle Mississippian. Data from the type species of all 184 trepostome genera from the Early Ordovician to the Late Triassic were utilised to calculate the Bryozoan Skeletal Index (BSI) as a proxy for the degree of calcification. BSI values and genus-level diversity did not decrease across the transition from the Calcite I Sea to the Aragonite II Sea. Nor were there any changes in the number of genus originations and extinctions. This suggests that trepostome bryozoans were not passive hypercalcifiers but active biomineralisers that controlled the mineralogy and robustness of their skeletons regardless of changes in seawater chemistry.

Ordovician and Mississippian stratigraphy in southwestern Missouri, USA

ABSTRACT A succession of Ordovician and Mississippian carbonates, separated unconformably, is exposed across the southern flank of the Ozark Dome in southwest Missouri. Deposits of both periods exemplify typical facies of the Midwestern United States: carbonate tidal-flat assemblages for the Early Ordovician and carbonate shelf environments for the Early–Middle Mississippian. The basic stratigraphic sequence of these deposits has been known for over a century, but interesting features remain to be addressed. Thin discontinuous sandstones are present within the Early Ordovician Cotter Dolomite, but the informal Swan Creek sandstone member seems anomalous. This sandstone can exceed 5 m in thickness and is fairly continuous across southwest Missouri. Most Ordovician sandstones in Missouri mark major transgressions above regional unconformities, but not the Swan Creek, and there is no obvious source of the sand. Therefore, we hypothesize that the Swan Creek represents reworked eolian dunes blown across the broad peritidal environment. Clastic sandstone dikes, apparently sourced from the Swan Creek, cut across beds of Cotter Dolomite near faults. We propose that these dikes are evidence of local faulting and seismicity during the Early Ordovician. Early and Middle Mississippian limestones comprise a sequence of shelf deposits, although mud mounds and other facies changes near the Missouri-Arkansas line mark the edge of the Mississippian shelf and the transition to a ramp setting. Early Mississippian carbonate deposition was interrupted by a short and localized influx of siliciclastic sediment comprising the Northview Formation. The Northview has additional characteristics consistent with a river-dominated deltaic deposit, which we suggest as its origin. If correct, this hypothesis implies that the history of tectonic features in the Midwest is more complicated than yet known. Finally, facies changes within and between the local Mississippian formations may record an early crustal response to the impending Ouachita orogeny farther to the south.

Furongian (Jiangshanian) occurrences of radiodonts in Poland and South China and the fossil record of the Hurdiidae

The Furongian period represents an important gap in the fossil record of most groups of non-biomineralizing organisms, owing to a scarcity of Konservat-Lagerstätten of that age. The most significant of these deposits, the Jiangshanian strata of the Sandu Formation near Guole Township (Guangxi, South China), have yielded a moderately abundant, but taxonomically diverse soft-bodied fossil assemblage, which provides rare insights into the evolution of marine life at that time. In this contribution, we report the first discovery of a radiodont fossil from the Guole Konservat-Lagerstätte. The specimen is an incomplete frontal appendage of a possibly new representative of the family Hurdiidae. It is tentatively interpreted as composed of seven podomeres, six of which bearing laminiform endites. The best preserved of these endites is especially long, and it bears short auxiliary spines that greatly vary in size. This is the second occurrence of hurdiids and more generally radiodonts in the Furongian, the first being the external mould of an oral cone from Jiangshanian strata of the Wiśniówka Sandstone Formation in Poland. Restudy of this Polish specimen confirms that it belongs to a hurdiid radiodont and best compares to Peytoia. The family Hurdiidae includes the oldest (basal Cambrian Epoch 2) and youngest (Early Ordovician, possibly Early Devonian) representatives of the Radiodonta and as such, has the longest stratigraphical range of the group. Yet, hurdiids only became prominent components of marine ecosystems during the middle Cambrian (Miaolingian), and their fossil record in younger strata remains limited.

Cambrian–Lower Ordovician of SW Quebec–NE New York

ABSTRACT The Ottawa aulacogen/graben on the NE US—Canadian (SW Quebec and eastern Ontario) border is a long ENE-trending structure formed with initial late Neo proterozoic rifting of the Rodinia supercontinent. This rifting formed the active spreading arms (New York Promontory and Quebec Reentrant) along the (presently) NE margin of the new Laurentia paleocontinent, with the Ottawa aulacogen commonly regarded as a failed arm of the rifting. However, no sediment accumulation in the aulacogen is recorded until the late early Cambrian subsidence of a SE- trending belt that includes the aulacogen and its extension, the Franklin Basin, in NW Vermont. Late early Cambrian marine onlap (Altona Formation) followed by more rapid late middle Cambrian subsidence and deposition of fluviatile arkoses (Covey Hill Formation of SW Quebec and Ausable Formation/Member of eastern New York) record rapid foundering of this “failed arm.” Subsequent deposition (latest middle Cambrian–Early Ordovician) in the Ottawa aulacogen produced a vertical succession of lithofacies that are fully comparable with those of the shelf of the New York Promontory. One of the greatest challenges in summarizing the geological history of the Ottawa aulacogen is the presence of a duplicate stratigraphic nomenclature with lithostratigraphic names changing as state and provincial borders are crossed.