scholarly journals Systematics of the Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi, with species from the Great Basin, western USA

2021 ◽  
pp. 1-26
Author(s):  
Jonathan M. Adrain ◽  
Talia S. Karim

Abstract The Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi has long been known from a small number of pygidia assigned to a single formally named species, and its affinities have not been assessed. Silicified material from western Utah and southeastern Idaho includes six distinct species assigned to the genus, one of which is the type species. Two others (G. moffitti and G. pankowskii) are new and formally named. An additional three species that are clearly new but known from sparse material are described in open nomenclature. Gonioteloides has a stratigraphic distribution through five consecutive trilobite zones in the mid-Stairsian Stage (upper Tremadocian). Although exoskeletal morphology of three species is almost completely known, the phylogenetic affinity of the taxon remains difficult to determine. It is tentatively assigned to Dimeropygidae Hupé. UUID: http://zoobank.org/23257d6c-262b-4ef5-ae4e-cc431777e67e

Zootaxa ◽  
2018 ◽  
Vol 4525 (1) ◽  
pp. 1
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM

Field-based revision and phylogenetic analysis demonstrate that the pliomerid trilobite taxon Ibexaspis Přibyl and Vaněk (in Přibyl et al., 1985), previously known from a single formally named species (I. brevis [Young, 1973]), belongs to a complex of 14 mostly newly discovered, related species from the Early Ordovician (Floian; Tulean and Blackhillsian) of the northern Laurentian margin. The species are known from silicified samples recovered from sections in eastern Nevada, western Utah, and southeastern Idaho. The stratigraphically early Tuleaspis n. gen. (type species: T. jeneki n. sp.; Tulean; low Protopliomerella contracta Zone) includes its type and two species described in open nomenclature. Tuleaspis is sister to the remainder of the clade. Ibexaspis now includes three additional species: I. coadyi n. sp. (Blackhillsian; Carolinites nevadensis Zone), I. leuppi n. sp. (Blackhillsian; Presbynileus ibexensis Zone), and I. rupauli n. sp. (Blackhillsian; "Pseudocybele nasuta Zone"). Ibexapsis is sister to a clade of Millardaspis n. gen. + Deltapliomera n. gen. Millardaspis (type species M. milsteadi n. sp.; Tulean; Heckethornia hyndeae Zone) also includes M. knoxi n. sp. (Tulean; Panisaspis sevierensis Zone). Deltapliomera (type species D. humphriesi n. sp.; Blackhillsian, Carolinites nevadensis Zone) also includes D. inglei n. sp. (Tulean; Heckethornia bowiei Zone), D. heimbergi (Tulean; Panisaspis sevierensis Zone), D. eppersoni n. sp. (Blackhillsian; Bathyurina plicolabeona Zone), and a species described in open nomenclature. 


Zootaxa ◽  
2011 ◽  
Vol 2969 (1) ◽  
pp. 1 ◽  
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN

Panisaspis n. gen. is a clade of pliomerid trilobites from the Tulean and Blackhillsian stages (Floian) of the Great Basin.  It includes Protopliomerops? quattuor Hintze, 1953, and ten new species, six of which are formally named: Panisaspis millardensis (type species), P. sevierensis, P. deltaensis, P. rancherensis, P. topscityensis, and P. loganensis.  Four new species are not well enough known for formal naming and are described in open nomenclature.  All species are Tulean in age except for P. millardensis, which is earliest Blackhillsian.  Synapomorphies of Panisaspis include a short, nearly semicircular anterior border; small L1; large genal spines; a rounded, ovoid hypostomal border; elongated third pygidial spines; and a large, triangular terminal piece with distinct pitted impressions.  Phylogenetic analysis indicates that P. millardensis and P. sevierensis are sister taxa, and that P. deltaensis, P. rancherensis, P. topscityensis, P. quattuor, and P. loganensis are successive sister species.  The group may be sister to Ibexaspis Přibyl and Vaněk in Přibyl et al., 1985. 


Zootaxa ◽  
2019 ◽  
Vol 4661 (2) ◽  
pp. 201-255
Author(s):  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM

The Stairsian genus Tesselacauda Ross, 1951, has historically included two species, the poorly known type, T. depressa Ross, 1951 (Bearriverops loganensis Zone), and the even less well known T. flabella Kobayashi, 1955 (Bearriverops alsacharovi Zone), which may not belong to the genus. The family assignment of the genus has long been in question, with some workers assigning it to Cheiruridae and some to Pliomeridae. New field collections from western Utah and southeastern Idaho yield abundant material of T. depressa, which facilitates revision on the basis of multiple specimens of most exoskeletal parts. Two additional well known species are proposed, T. morrisoni (Rossaspis leboni Zone), and T. kriegerae (Bearriverops alsacharovi Zone). A third new species, very similar to T. depressa, is described in open nomenclature from the Rossaspis leboni Zone. Knowledge of hypostomes from silicified material helps to clarify the basal morphologies of cheirurid versus pliomerid trilobites. Pliomerids have anteroposteriorly elongate hypostomes with a basic pattern of three pairs of lateral hypostomal spines and a single posteromedian spine. Some or all of the spines are reduced or lost in various taxa. Cheirurids either lack paired spines or have only one or two pairs, and never have a posteromedian spine. Cheirurid hypostomes tend to be much shorter and more subquadrate than pliomerids. Other differences between the families are: a small, triangular or trapezoidal rostral plate in pliomerids versus a wide, short plate in cheirurids; a thoracic segment count commonly of 11–13 in Cheiruridae (fewer in one derived subfamily) versus commonly 15 or more in pliomerids (fewer in two derived subfamilies); thoracic pleurae with subequal bands and a prominent furrow in cheirurids versus a much more inflated and rib-like posterior band, reduced anterior band, and short, anteriorly placed furrow; and pygidia with four or fewer segments in cheirurids versus commonly five in pliomerids (again, fewer in two derived subfamilies). On these and other criteria, Tesselacauda is clearly a cheirurid, assigned for the present to the presumptively basal and possibly paraphyletic Subfamily Pilekiinae.


2021 ◽  
pp. 1-5 ◽  
Author(s):  
James C. Lamsdell

One of the oldest fossil horseshoe crabs figured in the literature is Entomolithus lunatus Martin, 1809, a Carboniferous species included in his Petrificata Derbiensia. While the species has generally been included within the genus Belinurus Bronn, 1839, it was recently used as the type species of the new genus Parabelinurus Lamsdell, 2020. However, recent investigation as to the appropriate authority for Belinurus (see Lamsdell and Clapham, 2021) revealed that all the names in Petrificata Derbiensia were suppressed in Opinion 231 of the International Commission on Zoological Nomenclature (1954) for being consistently nonbinomial under Article 11.4 of the International Code of Zoological Nomenclature (ICZN) (International Commission on Zoological Nomenclature, 1999). Despite the validation of several species names for anthozoans, brachiopods, and cephalopods described in Petrificata Derbiensia in subsequent rulings (International Commission on Zoological Nomenclature, 1956a, b), Belinurus lunatus has not been the subject of any subsequent Commission ruling or opinion, and so its use in Petrificata Derbiensia remains suppressed. The Belinurus lunatus species name was used in several subsequent publications during the 1800s, none of which made the name available under ICZN article 11.5; Parkinson (1811) is also suppressed for being nonbinomial, while Woodward (1830), Buckland (1837), Bronn (1839), and Baily (1859) refer to the species only as a synonym of Belinurus trilobitoides (Buckland, 1837) through citation to the suppressed Pretificata Derbiensia. The first author to make Belinurus lunatus an available name was Baldwin (1905), who used the name in reference to a new figured specimen from Sparth Bottoms, Rochdale, UK, but again as an explicit junior synonym of Belinurus trilobitoides (Buckland, 1837). Therefore, it was not until Eller (1938) treated B. lunatus as a distinct species from B. trilobitoides that B. lunatus became an available name as per ICZN Article 11.6.1 under the authorship of Baldwin (1905) following ICZN Article 50.7.


Zootaxa ◽  
2011 ◽  
Vol 2992 (1) ◽  
pp. 1-51
Author(s):  
PATRICK DAVID ◽  
GERNOT VOGEL ◽  
ALAIN DUBOIS

This paper analyzes the consequences of the non-respect of the Rules of the Code to ascertain the valid subsequent designation of the nucleospecies (type species) of the nominal genus Trimeresurus Lacépède, 1804. The long accepted designation was invalid because it was based on a nominal species which was not among the prenucleospecies (originally included species) of the nominal genus. In contrast with the commonly accepted viewpoint which makes the Indian taxon Coluber gramineus Shaw, 1802 the nucleospecies of the genus, we show that this role is played by Trimeresurus viridis Lacépède, 1804, a species inhabiting the Lesser Sunda Islands and Timor and, as a nomen oblitum, a senior synonym of Trimeresurus albolabris insularis Kramer, 1977, a taxon now considered a distinct species. The important nomenclatural implications of this finding are discussed here, especially with regard to the recent splitting of the genus Trimeresurus. The generic nomen Trimeresurus should be associated with the Trimeresurus albolabris group of species currently placed in the genus or subgenus Cryptelytrops Cope, 1860. A lectophoront (lectotype) is selected and described for Trimeresurus viridis Lacépède, 1804. Coluber viridis Bechstein, 1802 is an invalid objective junior synonym of Coluber gramineus Shaw, 1802. The current content of the genus Trimeresurus and of its eight subgenera is provided. Some clarifications or improvements to the Code are suggested.


Zootaxa ◽  
2006 ◽  
Vol 1139 (1) ◽  
pp. 27 ◽  
Author(s):  
J. POORANI ◽  
R. G. BOOTH

The following nomenclatural changes in the Oriental Sticholotidini (Coccinellidae: Sticholotidinae) are proposed: Sticholotis quadrisignata rugicollis Korschefsky (1934) is elevated to a distinct species (status revised) and Sticholotis gomyi Chazeau (1974) is a new junior synonym of S. rugicollis (new synonym). The genus Neojauravia Gordon & Almeida (1991), described from Brazil, is a new junior synonym of Pharoscymnus Bedel (1906) and the type species, Neojauravia naeida Gordon & Almeida (1991), is a new junior synonym of Pharoscymnus flexibilis (Mulsant, 1853) (new synonyms).


2012 ◽  
Vol 20 (6) ◽  
pp. 1133-1147 ◽  
Author(s):  
J. Gillespie ◽  
S. T. Nelson ◽  
A. L. Mayo ◽  
D. G. Tingey

2007 ◽  
Vol 21 (2) ◽  
pp. 173 ◽  
Author(s):  
Klaus Rützler ◽  
Manuel Maldonado ◽  
Carla Piantoni ◽  
Ana Riesgo

The systematics of tropical and subtropical western Atlantic species of Iotrochota is re-examined in light of the discovery of an undescribed species. Iotrochota birotulata (Higgin), the type species, is found to have more characters than previously recognised and is redefined with emphasis on a skeleton of spongin fibres containing stout, curved styles and strongyles (category I) and an interstitial spiculation consisting mainly of longer, slender and straight styles (II). Iotrochota bistylata Boury-Esnault is confirmed as a synonym of the above. The new species, named I. arenosa, sp. nov., differs in external morphology, strong mucus development, incorporation of sand and interstitial spicules that are mainly long, straight strongyles. Iotrochota atra (Whitfield), thought to be a synonym of I. birotulata, is recognised as a separate species occurring exclusively in the Bahamas and is found to be a senior synonym of I. imminuta Pulitzer-Finali; it is morphologically very similar to I. birotulata, but lacks birotulae and has a strongly reduced skeleton of megascleres (mostly one category of delicate strongyles). Iotrochota agglomerata Lehnert & van Soest is recognised as the fourth distinct species for its unusual colour (orange), thinly encrusting habit and special spiculation (styles with tylostylote modifications).


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