The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA

Zootaxa ◽  
2018 ◽  
Vol 4525 (1) ◽  
pp. 1
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM
Keyword(s):  
Phylogenetic Analysis ◽  
Great Basin ◽  
Related Species ◽  
Type Species ◽  
New Genera ◽  
Additional Species ◽  
Western Usa ◽  
Early Ordovician ◽  
Laurentian Margin

Field-based revision and phylogenetic analysis demonstrate that the pliomerid trilobite taxon Ibexaspis Přibyl and Vaněk (in Přibyl et al., 1985), previously known from a single formally named species (I. brevis [Young, 1973]), belongs to a complex of 14 mostly newly discovered, related species from the Early Ordovician (Floian; Tulean and Blackhillsian) of the northern Laurentian margin. The species are known from silicified samples recovered from sections in eastern Nevada, western Utah, and southeastern Idaho. The stratigraphically early Tuleaspis n. gen. (type species: T. jeneki n. sp.; Tulean; low Protopliomerella contracta Zone) includes its type and two species described in open nomenclature. Tuleaspis is sister to the remainder of the clade. Ibexaspis now includes three additional species: I. coadyi n. sp. (Blackhillsian; Carolinites nevadensis Zone), I. leuppi n. sp. (Blackhillsian; Presbynileus ibexensis Zone), and I. rupauli n. sp. (Blackhillsian; "Pseudocybele nasuta Zone"). Ibexapsis is sister to a clade of Millardaspis n. gen. + Deltapliomera n. gen. Millardaspis (type species M. milsteadi n. sp.; Tulean; Heckethornia hyndeae Zone) also includes M. knoxi n. sp. (Tulean; Panisaspis sevierensis Zone). Deltapliomera (type species D. humphriesi n. sp.; Blackhillsian, Carolinites nevadensis Zone) also includes D. inglei n. sp. (Tulean; Heckethornia bowiei Zone), D. heimbergi (Tulean; Panisaspis sevierensis Zone), D. eppersoni n. sp. (Blackhillsian; Bathyurina plicolabeona Zone), and a species described in open nomenclature. 

Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA

Zootaxa ◽  
2011 ◽  
Vol 2969 (1) ◽  
pp. 1 ◽  
Author(s):  
NEO E.B. McADAMS ◽  
JONATHAN M. ADRAIN
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Great Basin ◽  
Type Species ◽  
New Genus ◽  
Sister Species ◽  
Anterior Border ◽  
Western Usa ◽  
Lower Ordovician

Panisaspis n. gen. is a clade of pliomerid trilobites from the Tulean and Blackhillsian stages (Floian) of the Great Basin.  It includes Protopliomerops? quattuor Hintze, 1953, and ten new species, six of which are formally named: Panisaspis millardensis (type species), P. sevierensis, P. deltaensis, P. rancherensis, P. topscityensis, and P. loganensis.  Four new species are not well enough known for formal naming and are described in open nomenclature.  All species are Tulean in age except for P. millardensis, which is earliest Blackhillsian.  Synapomorphies of Panisaspis include a short, nearly semicircular anterior border; small L1; large genal spines; a rounded, ovoid hypostomal border; elongated third pygidial spines; and a large, triangular terminal piece with distinct pitted impressions.  Phylogenetic analysis indicates that P. millardensis and P. sevierensis are sister taxa, and that P. deltaensis, P. rancherensis, P. topscityensis, P. quattuor, and P. loganensis are successive sister species.  The group may be sister to Ibexaspis Přibyl and Vaněk in Přibyl et al., 1985. 

scholarly journals Systematics of the Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi, with species from the Great Basin, western USA

2021 ◽  
pp. 1-26
Author(s):  
Jonathan M. Adrain ◽  
Talia S. Karim
Keyword(s):  
Great Basin ◽  
Type Species ◽  
Distinct Species ◽  
Western Usa ◽  
Early Ordovician ◽  
Phylogenetic Affinity ◽  
Genus One

Abstract The Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi has long been known from a small number of pygidia assigned to a single formally named species, and its affinities have not been assessed. Silicified material from western Utah and southeastern Idaho includes six distinct species assigned to the genus, one of which is the type species. Two others (G. moffitti and G. pankowskii) are new and formally named. An additional three species that are clearly new but known from sparse material are described in open nomenclature. Gonioteloides has a stratigraphic distribution through five consecutive trilobite zones in the mid-Stairsian Stage (upper Tremadocian). Although exoskeletal morphology of three species is almost completely known, the phylogenetic affinity of the taxon remains difficult to determine. It is tentatively assigned to Dimeropygidae Hupé. UUID: http://zoobank.org/23257d6c-262b-4ef5-ae4e-cc431777e67e

Revision of Hestiasulini Giglio-Tos, 1915 stat. rev. (Insecta: Mantodea: Hymenopodidae) of Borneo, with description of new taxa and comments on the taxonomy of the tribe

Zootaxa ◽  
2017 ◽  
Vol 4291 (2) ◽  
pp. 243 ◽  
Author(s):  
CHRISTIAN J. SCHWARZ ◽  
EVGENY SHCHERBAKOV
Keyword(s):  
New Taxa ◽  
Related Species ◽  
Type Species ◽  
The Other ◽  
New Genera ◽  
Additional Species ◽  
Apical Process ◽  
Se Asia ◽  
Partial Fusion ◽  
Subgenital Plate

The Bornean Oxypilinae species previously included in the genus Hestiasula Saussure, 1871 are revised. External morphology and genital characters of Sundaian taxa differ considerably from the Indian type species H. brunneriana Saussure, 1871, necessitating taxonomic and nomenclatural changes. The genus Catestiasula Giglio-Tos, 1915 is reinstated. It is characterized by the apomorphic partial fusion of the dorsal and ventral laminae of the left phallomere, and by the complete reduction of the apical process. It is represented on Borneo with two species, C. nitida (Brunner de Wattenwyl, 1893) and C. moultoni Giglio-Tos, 1915. An additional species, C. seminigra (Zhang, 1992) n. comb. occurs in continental SE Asia. Two new genera are described, united by the lack of styli, the presence of dorsal carinae on the subgenital plate, and the unique morphology of the dorsal lamina of the left phallomere, but differing in the morphology of head and forelegs. Astyliasula gen. nov. accommodates A. phyllopus (De Haan, 1842) n. comb. and related species from the Sunda Islands and continental SE Asia: A. basinigra (Zhang, 1992) n. comb., A. hoffmanni (Tinkham, 1937) n. comb., A. javana (Beier, 1929) n. comb., A. major (Beier, 1929) n. comb., A. inermis (Wood-Mason, 1879) n. comb., and A. wuyshana (Yang & Wang, 1999) n. comb.. Hestias sarawaca Westwood, 1889 is removed from synonymy with A. phyllopus and reinstated as A. sarawaca (Westwood, 1889) n. comb.. The monotypic taxon Pseudohestiasula borneana gen. nov. sp. nov. is erected for a Bornean endemic more closely related to Astyliasula than to the other genera. Hestiasula is now restricted to H. brunneriana and related species from India and adjacent countries. The tribe Hestiasulini Giglio-Tos, 1915 stat. rev. is proposed for all genera more closely related to Hestiasula than to other Oxypilinae, that is Hestiasula, Ephestiasula, Catestiasula, Astyliasula, and Pseudohestiasula. New data on the ecology and distribution of all Bornean Hestiasulini as well as a key to the Oxypilinae of Borneo are provided. 

Phylogeny and taxonomy of the Rhinorhynchinae (Coleoptera:Nemonychidae)

2010 ◽  
Vol 24 (6) ◽  
pp. 573 ◽  
Author(s):  
Guillermo Kuschel ◽  
Richard A. B. Leschen
Keyword(s):  
Phylogenetic Analysis ◽  
Type Species ◽  
New Caledonia ◽  
Molecular Data ◽  
Host Use ◽  
New Genera ◽  
Plant Associations ◽  
The Family ◽  
Fossil Genus ◽  
Subjective Synonym

An overview of the taxa of Rhinorhynchinae (Nemonychidae) is presented. A phylogenetic analysis of the 19 extant genera of the Rhinorhynchinae and one fossil genus from the Lower Cretaceous (Cratomacer Zherikhin & Gratshev), based on 29 characters of adults, larvae and host plant associations, shows three monophyletic groups, treated as tribes (Rhinorhynchini, Mecomacerini and Rhynchitomacerini). Primitive associations are unknown for Rhinorhynchinae and early host use diversification included associations with Ranunculaceae (Nemonyx Redtenbacher), Pinaceae (Cimberis Gozis), Nothofagaceae (Rhynchitomacerini) and Podocarpaceae (Rhinorhynchini). While Mecomacerini diversified on Araucariaceae, within Rhinorhynchini there was a single reversal to Araucariaceae and a shift in Atopomacer Kuschel to Pinaceae. Placement of Cratomacer into Mecomacerini is consistent with geological and molecular data that suggest gymnosperms may have been the primitive hosts for the family. Three new genera are described in Rhinorhynchinae: Araucomacer, gen. nov. (type species A. hirticeps Kuschel) from Chile, Idiomacer, gen. nov. (type species I. basicornis, sp. nov.) from New Caledonia, and Zimmiellus, gen. nov. (type species Z. fronto, sp. nov.) from Queensland, Australia, while Stenomacer Kuschel, stat. nov. from Chile is reinstated from synonymy. Seven species are described as new: Atopomacer grandifurca, sp. nov. from Costa Rica and Panama, A. obrieni, sp. nov. from Mexico, A. panamensis, sp. nov. from Panama, A. pini, sp. nov. from Mexico, A. podocarpi, sp. nov. from Venezuela, Basiliogeus dracrycarpi, sp. nov. from western New Guinea and B. inops, sp. nov. from Queensland. Rhynchitomacer rufus Kuschel is a new junior subjective synonym of R. nigritus Kuschel, syn. nov., R. viridulus Kuschel is a new junior subjective synonym of R. flavus Voss, syn. nov. and Stenomacer fuscus Kuschel is a new junior subjective synonym of S. vernus Kuschel, syn. nov. Keys to genera of Rhinorhynchinae and to species of Atopomacer, Rhynchitomacer and Stenomacer are included. Diagnoses of the three extant subfamilies of Nemonychidae are included.

Classification, reconstructed phylogeny and geographical history of genera of Pilipalpinae (Coleoptera : Tenebrionoidea : Pyrochroidae)

1995 ◽  
Vol 9 (4) ◽  
pp. 563 ◽  
Author(s):  
DA Pollock
Keyword(s):  
Phylogenetic Analysis ◽  
New Zealand ◽  
Type Species ◽  
New Genera ◽  
Parsimony Analysis ◽  
Data Set ◽  
Geological Evidence ◽  
Southern South America ◽  
Single Clade ◽  
History Of

The 12 genera of Pilipalpinae are classified on the basis of characters of larvae and adults. Three new genera and six new species are here described: Malagaethes, gen. nov. (type species M. lawrencei, sp. nov.); Ranomafana, gen. nov. (type species R. steineri, sp. nov.); Binburrum, gen. nov. (type species Techmessa ruficollis Champion); Binburrum angusticollis, sp. nov.; Binburrum concavifrons, sp. nov.; Cycloderus immaculicollis, sp. nov. and Cycloderus hirsutus, sp. nov. The following new synonymies of specific names are proposed (with valid names given first): Paromarteon constans Lea, 1917 = Eucistela cyanea Carter, 1922; Paromarteon mutabile Blackburn, 1897 = Paromarteon mutabile var. nigripenne Lea, 1920; Temnopalpus bicolor Blackburn, 1888 = Temnopalpus tricolor Lea, 1920; Pilipalpus dasytoides Fairmaire, 1876 = Copobaenus maculicollis Pic, 1942 and Pilipalpus danvini Abdullah, 19646; Exocalopus pectinatus Broun, 1893 = Exocalopus antennalis Broun, 1903. The following subspecies have been elevated to species rank: Paromarteon apicale Lea, Paromarteon fasciatum Lea and Paromarteon parvum Lea. Phylogenetic analysis of 30 structural characters of larvae and adults yielded the following set of incompletely resolved relationships among genera of Pilipalpinae: (((Paromarteon + ((Temnopalpus + Malagaethes) + Pilipalpus + (Ranomafana + (Incollogenius + ((Exocalopus + (Binburrum + (Cycloderus + Morpholycus)) + Techmessodes) + Techmessa))))). The data set contained much homoplasy and several reversals. The historical geographical relationships inferred from the reconstructed phylogeny were compared with geological evidence for the break-up of Pangaea and Gondwanaland. The ancestral stock of Pilipalpinae was widespread on Gondwanaland, and differentiated through its fragmentation. Remnant relict genera persisted on Madagascar, New Zealand, southern South America (Magellanica), and Australia. Brooks Parsimony Analysis was conducted on the data resulting in the following area relationships: (Holarctic + (Madagascar + (New Zealand + (Australia + Chile)))). This agrees generally with accepted geological evidence and is considered support for they hypothesised phylogeny. A single clade (Temnopalpus + Malagaethes) was in disagreement (homoplasous) with the area cladogram, indicating possible incongruence in the data. The area relationships of other Southern Hemisphere groups were compared with Pilipalpinae.

Phylogenetic analysis of Lumbrineridae Schmarda, 1861 (Annelida: Polychaeta)

Zootaxa ◽  
2006 ◽  
Vol 1332 (1) ◽  
pp. 1 ◽  
Author(s):  
LUIS F. CARRERA-PARRA
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Type Species ◽  
Phylogenetic Reconstruction ◽  
Morphological Characters ◽  
New Genera ◽  
The Family ◽  
Three New Species

This study represents the first phylogenetic reconstruction of lumbrinerid genera using parsimony analyses of 38 morphological characters. Following higher-level phylogenetic analysis, Oenone (Oenonidae) was selected as outgroup. The analysis was restricted to type species for each genus, yielded 24 equally parsimonious trees, which after successive weighting were reduced to one tree (CI= 0.7396). The topology of this tree revealed the separation of the family into four main clades: 1. Lysarete, 2. Arabellonereis, 3. Scoletoma, Lumbrineris, Hilbigneris gen. nov., Kuwaita, Lumbricalus, Sergioneris gen. nov. and Eranno, and 4. Abyssoninoe, Cenogenus, Lumbrinerides, Lumbrineriopsis, Augeneria, Loboneris gen. nov., Gallardoneris gen. nov., Helmutneris gen. nov., and Gesaneris gen. nov.; the position of Ninoe is unclear. A diagnosis of each genus is provided, including the description of six new genera and three new species. A key to lumbrinerid genera is included.

North American Components of the Genus Diasemia of Authors (Lepidoptera: Pyralidae)

1957 ◽  
Vol 89 (4) ◽  
pp. 164-170 ◽  
Author(s):  
Eugene Munroe
Keyword(s):  
Structural Study ◽  
North American ◽  
Related Species ◽  
Type Species ◽  
New Combination ◽  
New Genera ◽  
Undescribed Species ◽  
The North ◽  
Lepidoptera Pyralidae ◽  
The Tropics

Work undertaken in connection with Mr. C. P. Kimball's projected list of Florida Lepidopera has revealed a number of problems in the North American species usually referred to Diasemia Hübner ([1824-25] p. 348), type Pyralis literalis Denis and Schiffermüller (=Phalaena litterata Scopoli) (Figs. 1, 2). A structural study shows that these species belong to five groups, generically distinct from one another and from the type species of Diasemia. Diasemia alaskalis Gibson (Fig. 3) is congeneric with Udea ferrugalis (Hübner) and should be known as Udea alaskalis (Gibson), new combination. I have characterized the genus Udea Guenée in an earlier paper (Munroe, 1950). Diasemia plumbosignalis Fernald (Fig. 10) and related species belong to the genus Choristostigma Warren, 1892: 440. The species of Choristostigma will be discussed in a separate publication. Diasemia magdalena Fernald and an undescribed species belong to the genus Daulia Walker (1859: 975) hitherto known from the tropics of the Old World and from Argentina. Hydrocampa ramburialis Duponchel and Desmia? janassialis Walker require new genera.

scholarly journals Review of the New World genera of the Leafhopper Tribe Erythroneurini (Hemiptera: Cicadellidae: Typhlocycbinae)

2006 ◽  
Vol 37 (1-6) ◽  
pp. 118-190 ◽  
Author(s):  
Christopher H. Dietrich ◽  
Dmitry A. Dmitriev
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Type Species ◽  
New World ◽  
New Genus ◽  
Morphological Characters ◽  
New Genera ◽  
Previously Treated ◽  
One New Species

The genus-level classification of New World Erythroneurini is revised based on results of a phylogenetic analysis of 100 morphological characters. The 704 known species are placed into 18 genera. Erasmoneura Young and Eratoneura Young, previously treated as subgenera of Erythroneura Fitch, and Erythridula Young, most recently treated as a subgenus of Arboridia Zachvatkin, are elevated to generic status. Three species previously included in Erasmoneura are placed in a new genus, Rossmoneura (type species, Erythroneura tecta McAtee). The concept of Erythroneura is thereby narrowed to include only those species previously included in the nominotypical subgenus. New World species previously included in Zygina Fieber are not closely related to the European type species of that genus and are therefore placed in new genera. Neozygina, n. gen., based on type species Erythroneura ceonothana Beamer, includes all species previously included in the “ceonothana group”, and Zyginama, n. gen., based on type species Erythroneura ritana Beamer, includes most species previously included in the “ritana group” of New World Zygina. Five additional new genera are described to include other previously described North American Erythroneurini: Hepzygina, n. gen., based on type species Erythroneura milleri Beamer and also including E. aprica McAtee; Mexigina, n. gen., based on type species Erythroneura oculata McAtee; Nelionidia, n. gen., based on type species N. pueblensis, n. sp., three additional new species, and Erythroneura amicis Ross; Neoimbecilla, n. gen., based on type species Erythroneura kiperi Beamer and one new species; and Illinigina, n. gen., based on type species Erythroneura illinoiensis Gillette. Five new genera, based on previously undescribed species, are also recognized: Aztegina, n. gen, based on A. punctinota, n. sp., from Mexico; Amazygina, n. gen., based on type species A. decaspina, n. sp., and three additional new species from Ecuador; Hamagina, n. gen., based on type species H. spinigera, n. sp., and two additional new species from Peru and Ecuador; Napogina, n. gen., based on type species N. recta, n. sp., and one additional new species from Ecuador; Perugina, n. gen., based on type species P. denticula, n. sp., from Peru; and Spinigina, n. gen., based on type species S. hirsuta, n. sp., and an additional new species from Peru. Phylogenetic analysis suggests that the New World Erythroneurini consist of three lineages resulting from separate invasions from the Old World.

Revision of the Early Ordovician (late Tremadocian; Stairsian) cheirurid trilobite Tesselacauda Ross, with species from the Great Basin, western USA

Zootaxa ◽  
2019 ◽  
Vol 4661 (2) ◽  
pp. 201-255
Author(s):  
JONATHAN M. ADRAIN ◽  
TALIA S. KARIM
Keyword(s):  
New Species ◽  
Great Basin ◽  
Thoracic Segment ◽  
Basic Pattern ◽  
Western Usa ◽  
Early Ordovician ◽  
The Family ◽  
Short Plate

The Stairsian genus Tesselacauda Ross, 1951, has historically included two species, the poorly known type, T. depressa Ross, 1951 (Bearriverops loganensis Zone), and the even less well known T. flabella Kobayashi, 1955 (Bearriverops alsacharovi Zone), which may not belong to the genus. The family assignment of the genus has long been in question, with some workers assigning it to Cheiruridae and some to Pliomeridae. New field collections from western Utah and southeastern Idaho yield abundant material of T. depressa, which facilitates revision on the basis of multiple specimens of most exoskeletal parts. Two additional well known species are proposed, T. morrisoni (Rossaspis leboni Zone), and T. kriegerae (Bearriverops alsacharovi Zone). A third new species, very similar to T. depressa, is described in open nomenclature from the Rossaspis leboni Zone. Knowledge of hypostomes from silicified material helps to clarify the basal morphologies of cheirurid versus pliomerid trilobites. Pliomerids have anteroposteriorly elongate hypostomes with a basic pattern of three pairs of lateral hypostomal spines and a single posteromedian spine. Some or all of the spines are reduced or lost in various taxa. Cheirurids either lack paired spines or have only one or two pairs, and never have a posteromedian spine. Cheirurid hypostomes tend to be much shorter and more subquadrate than pliomerids. Other differences between the families are: a small, triangular or trapezoidal rostral plate in pliomerids versus a wide, short plate in cheirurids; a thoracic segment count commonly of 11–13 in Cheiruridae (fewer in one derived subfamily) versus commonly 15 or more in pliomerids (fewer in two derived subfamilies); thoracic pleurae with subequal bands and a prominent furrow in cheirurids versus a much more inflated and rib-like posterior band, reduced anterior band, and short, anteriorly placed furrow; and pygidia with four or fewer segments in cheirurids versus commonly five in pliomerids (again, fewer in two derived subfamilies). On these and other criteria, Tesselacauda is clearly a cheirurid, assigned for the present to the presumptively basal and possibly paraphyletic Subfamily Pilekiinae.

scholarly journals The problems with Polypedilum Kieffer (Diptera: Chironomidae), with the description of Probolum subgen. n.

Zootaxa ◽  
2010 ◽  
Vol 2497 (1) ◽  
pp. 1 ◽  
Author(s):  
OLE A. SÆTHER ◽  
TROND ANDERSEN ◽  
LUIZ C. PINHO ◽  
HUMBERTO F. MENDES
Keyword(s):  
Phylogenetic Analysis ◽  
Type Species ◽  
Additional Species ◽  
Male And Female ◽  
Full Species ◽  
New Subgenus ◽  
Female Pupa

A phylogenetic analysis of the genus Polypedilum Kieffer utilizing only species with known larvae, and including one additional species presumed to belong to a new subgenus was performed. The subgenera Tripodura Townes, 1945, Pentapedilum Kieffer, 1913 and Cerobregma Sæther et Sundal, 1999, were all monophyletic, while Uresipedilum Oyewo et Sæther, 1998 was monophyletic when the new subgenus Probolum was excluded. Polypedilum s. str. was never monophyletic even when excluding the atypical P. nubifer (Skuse, 1889), but if the relationships between subgenera were unresolved a cladogram as short as the shortest trees could be erected with Polypedilum s. str. as monophyletic. It is proposed to reestablish the genus Tripedilum Kieffer, 1921a as a subgenus with the type species P. (T.) fuscipenne (Kieffer, 1921a) and including P. (T.) nubifer (Skuse); treat Polypedilum Kieffer, 1913 as valid (pending an application to the ICZN to supress Polypedilum Kieffer, 1912); reestablish the subgenus Kribionympha Kieffer, 1921a with the type species Polypedilum (Kribionympha) declivis Kieffer, 1922; give priority to Tripodura Townes, 1945 over senior synonyms; and erect the new subgenus Probolum Andersen et Sæther with Polypedilum (Probolum) marcondesi Pinho et Mendes sp. n. as type species and including at least P. (Probolum) pedatum excelsius Townes sensu Grodhaus et Rotramel, 1980, P. (Probolum) simantokeleum Sasa, Suzuki et Sakai, 1998 and P. (Probolum) bullum Zhang et Wang, 2004. Short diagnoses for the subgenera of Polypedilum are presented. Polypedilum (Probolum) marcondesi subgen. n., sp. n. is described and figured as male, female, pupa and larva. The two subspecies of P. (Uresipedilum) pedatum Townes, 1945 are regarded as full species. The male imago of P. (Uresipedilum) excelsius Townes, 1945 and the male and female of P. (Uresipedilum) pedatum Townes are redescribed.

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