Acoustics of Sound Production and of Hearing in the Bladder Cicada Cystosoma Saundersii (Westwood)

1978 ◽  
Vol 72 (1) ◽  
pp. 43-55 ◽  
Author(s):  
N.H. FLETCHER ◽  
K. G. HILL
Keyword(s):  
Sound Production ◽  
Abdominal Cavity ◽  
Low Frequency ◽  
Sound Intensity ◽  
Electrical Network ◽  
Directional Hearing ◽  
Hearing Sensitivity ◽  
Physical Mechanisms ◽  
Radiated Sound ◽  
Song Frequency

The male cicada of the species Cystosoma saundersii has a grossly enlarged, hollow abdomen and emits a loud calling song with a fundamental frequency of about 800 Hz. At the song frequency, its hearing is nondirectional. The female of C. saundersii lacks sound producing organs, has no enlargement of the abdomen, but possesses an abdominal air sac and has well developed directional hearing at the frequency of the species' song. Physical mechanisms are proposed that explain these observations in semi-quantitative detail using the standard method of electrical network analogues. The abdomen in the male, with its enclosed air, is found to act as a system resonant at the song frequency, thus contributing a large gain in radiated sound intensity. Coupling between this resonator and the auditory tympana accounts for the observed hearing sensitivity in the male, but destroys directionality. In the female, the abdominal cavity acts in association with the two auditory tympana as part of a phase shift network which results in appreciable directionality of hearing at the unusually low frequency of the male song.

scholarly journals A MODEL OF THE MECHANISM OF SOUND PRODUCTION IN CICADAS

1992 ◽  
Vol 173 (1) ◽  
pp. 123-153 ◽  
Author(s):  
H. C. Bennet-Clark ◽  
D. Young
Keyword(s):  
Resonant Frequency ◽  
Sound Production ◽  
Abdominal Cavity ◽  
Electrical Power ◽  
Low Frequency ◽  
Helmholtz Resonator ◽  
Sound Pulse ◽  
Song Frequency ◽  
Resonator System ◽  
Sound Pulses

1. Dried cicada bodies of the species Cyclochila australasiae and model cicadas made from a miniature earphone driving a plastic cavity were used to study the acoustics of sound production in male cicadas. 2. A model cicada with shape and dimensions similar to those of the abdomen of a male C. australasiae resonates at the natural song frequency of the species (4.3 kHz). The abdominal air sac of C. australasiae also resonates at frequencies close to the natural song frequency when excited by external sounds. In an atmosphere of chlorofluorocarbon (CFC) gas, the resonant frequency is lowered in keeping with the decrease in velocity of sound in the CFC gas. 3. At the model's resonant frequency, the driving earphone dissipates more electrical power with the cavity attached than without the cavity. The cavity of the model cicada acts as a narrow-band acoustic acceptance filter, tuned to the natural song frequency. 4. When the miniature earphone emits brief clicks, mimicking those produced by the natural tymbal mechanism, the model cicada produces sound pulses that vary in duration and shape according to the number and timing of the clicks. A coherent train of two or three resonant clicks results in a long slowly-decaying sound pulse similar to that in the natural song. 5. The natural song frequency can be predicted from the dimensions of the abdominal cavity and the tympana in C. australasiae using a simple equation for the resonant frequency of a Helmholtz resonator. This equation also predicts the song frequency of Macrotristria angularis and Magicicada cassini, but it fails with the low-frequency song of Magicicada septendecim. This discrepancy can be accounted for by the unusually thick tympana of M. septendecim, which tend to lower the resonant frequency of the system. 6. We conclude that the abdomen of male cicadas forms a Helmholtz resonator, the components of which are the large air sac as the cavity and the tympana as the neck of the resonator. We suggest that cicada sound production depends on the coupling of two resonators, that of the tymbal and that of the abdominal air sac, from which sound is radiated through the tympana. The coupled resonator system would produce the long sound pulses required for stimulating a sensitive sharply tuned auditory organ.

The Effects of Venting and Low-Frequency Gain Compensation on Performance in Noise with Directional Hearing Instruments

2006 ◽  
Vol 17 (03) ◽  
pp. 168-178 ◽  
Author(s):  
Melinda C. Freyaldenhoven ◽  
Patrick N. Plyler ◽  
James W. Thelin ◽  
Anna K. Nabelek ◽  
Samuel B. Burchfield
Keyword(s):  
Hearing Impairment ◽  
Hearing Aids ◽  
Low Frequency ◽  
Directional Hearing ◽  
Clinical Implications ◽  
Speech Understanding ◽  
Hearing Sensitivity ◽  
Speech Understanding In Noise ◽  
Hearing Instruments ◽  
Gain Compensation

The present study investigated the effects of gain compensation and venting on front-to-back ratios (FBRs), speech understanding in noise, and acceptance of noise in 19 listeners with hearing impairment utilizing directional hearing instruments. The participants were separated into two groups based on degree of low-frequency hearing sensitivity. Subjects were fitted binaurally with Starkey Axent II programmable behind-the-ear hearing aids and full-shell earmolds (select-a-vent). Results demonstrated that gain compensation and venting significantly affected FBRs for both groups; however, acceptance of noise was not significantly affected by gain compensation or venting for either group. Results further demonstrated that speech understanding in noise was unaffected by venting but may be improved with the use of gain compensation for some listeners. Clinical implications are discussed.

scholarly journals Wing, tail, and vocal contributions to the complex acoustic signals of courting Calliope hummingbirds

2011 ◽  
Vol 57 (2) ◽  
pp. 187-196 ◽  
Author(s):  
Christopher James Clark
Keyword(s):  
Sound Production ◽  
Low Frequency ◽  
Acoustic Signals ◽  
Wingbeat Frequency ◽  
Physical Mechanisms ◽  
Frequency Tone ◽  
Single Mechanism ◽  
Sound Pulses ◽  
Stellula Calliope ◽  
Courtship Displays

Abstract Multi-component signals contain multiple signal parts expressed in the same physical modality. One way to identify individual components is if they are produced by different physical mechanisms. Here, I studied the mechanisms generating acoustic signals in the courtship displays of the Calliope hummingbird Stellula calliope. Display dives consisted of three synchronized sound elements, a high-frequency tone (hft), a low frequency tone (lft), and atonal sound pulses (asp), which were then followed by a frequency-modulated fall. Manipulating any of the rectrices (tail-feathers) of wild males impaired production of the lft and asp but not the hft or fall, which are apparently vocal. I tested the sound production capabilities of the rectrices in a wind tunnel. Single rectrices could generate the lft but not the asp, whereas multiple rectrices tested together produced sounds similar to the asp when they fluttered and collided with their neighbors percussively, representing a previously unknown mechanism of sound production. During the shuttle display, a trill is generated by the wings during pulses in which the wingbeat frequency is elevated to 95 Hz, 40% higher than the typical hovering wingbeat frequency. The Calliope hummingbird courtship displays include sounds produced by three independent mechanisms, and thus include a minimum of three acoustic signal components. These acoustic mechanisms have different constraints and thus potentially contain different messages. Producing multiple acoustic signals via multiple mechanisms may be a way to escape the constraints present in any single mechanism.

THE SCALING OF SONG FREQUENCY IN CICADAS

1994 ◽  
Vol 191 (1) ◽  
pp. 291-294 ◽  
Author(s):  
H Bennet-Clark ◽  
D Young
Keyword(s):  
Resonant Frequency ◽  
Sound Production ◽  
Abdominal Cavity ◽  
Helmholtz Resonator ◽  
Ventral Surface ◽  
Recent Model ◽  
End Effect ◽  
Thin Walled ◽  
Song Frequency ◽  
Pressure Changes

In male cicadas, sound is generated by a pair of tymbals on the abdomen (Pringle, 1954). The tymbals buckle inwards causing pressure changes in the abdominal cavity, from which sound is radiated through the tympana (Young, 1990). A recent model of sound production in cicadas suggests that the abdominal cavity and tympana act as the components of a Helmholtz resonator that is excited by the drive from the tymbals (Bennet-Clark and Young, 1992). A Helmholtz resonator consists of a cavity open to the outside via a hole which has a real or notional neck, and the resonant frequency fo is given by the general equation: where c is the speed of sound in the fluid, taken as 340 m s-1 for air, A is the area of the neck, L is the length of the neck and V is the volume of the cavity. Where the resonator has two holes, these terms should be somewhat modified: A is the combined area of the two holes, L is 16/3pi r (~1.7r) for a simple hole in a thin-walled vessel and r is the radius of one hole (Seto, 1971). These modifications to equation 1, which include corrections for the acoustic end-effect at either side of a simple hole in the wall of a vessel, are applicable to a model of the male cicada, in which there are two tympana close to the ventral surface of the abdomen.

scholarly journals Earless toads sense low frequencies but miss the high notes

2017 ◽  
Vol 284 (1864) ◽  
pp. 20171670 ◽  
Author(s):  
Molly C. Womack ◽  
Jakob Christensen-Dalsgaard ◽  
Luis A. Coloma ◽  
Juan C. Chaparro ◽  
Kim L. Hoke
Keyword(s):  
High Frequency ◽  
Species Differences ◽  
Low Frequency ◽  
Auditory Brainstem ◽  
Low Frequencies ◽  
Hearing Sensitivity ◽  
Sensory Pathways ◽  
Airborne Sound ◽  
Vibrational Sensitivity ◽  
Species Specific

Sensory losses or reductions are frequently attributed to relaxed selection. However, anuran species have lost tympanic middle ears many times, despite anurans' use of acoustic communication and the benefit of middle ears for hearing airborne sound. Here we determine whether pre-existing alternative sensory pathways enable anurans lacking tympanic middle ears (termed earless anurans) to hear airborne sound as well as eared species or to better sense vibrations in the environment. We used auditory brainstem recordings to compare hearing and vibrational sensitivity among 10 species (six eared, four earless) within the Neotropical true toad family (Bufonidae). We found that species lacking middle ears are less sensitive to high-frequency sounds, however, low-frequency hearing and vibrational sensitivity are equivalent between eared and earless species. Furthermore, extratympanic hearing sensitivity varies among earless species, highlighting potential species differences in extratympanic hearing mechanisms. We argue that ancestral bufonids may have sufficient extratympanic hearing and vibrational sensitivity such that earless lineages tolerated the loss of high frequency hearing sensitivity by adopting species-specific behavioural strategies to detect conspecifics, predators and prey.

Effect of non-parallel mean flow on the Green’s function for predicting the low-frequency sound from turbulent air jets

2012 ◽  
Vol 695 ◽  
pp. 199-234 ◽  
Author(s):  
M. E. Goldstein ◽  
Adrian Sescu ◽  
M. Z. Afsar
Keyword(s):  
Green’S Function ◽  
Green's Function ◽  
Low Frequency ◽  
Source Location ◽  
Parallel Flow ◽  
Numerical Calculations ◽  
Mean Flow ◽  
Frequency Sound ◽  
The Mean ◽  
Radiated Sound

AbstractIt is now well-known that there is an exact formula relating the far-field jet noise spectrum to the convolution product of a propagator (that accounts for the mean flow interactions) and a generalized Reynolds stress autocovariance tensor (that accounts for the turbulence fluctuations). The propagator depends only on the mean flow and an adjoint vector Green’s function for a particular form of the linearized Euler equations. Recent numerical calculations of Karabasov, Bogey & Hynes (AIAA Paper 2011-2929) for a Mach 0.9 jet show use of the true non-parallel flow Green’s function rather than the more conventional locally parallel flow result leads to a significant increase in the predicted low-frequency sound radiation at observation angles close to the downstream jet axis. But the non-parallel flow appears to have little effect on the sound radiated at $9{0}^{\ensuremath{\circ} } $ to the downstream axis. The present paper is concerned with the effects of non-parallel mean flows on the adjoint vector Green’s function. We obtain a low-frequency asymptotic solution for that function by solving a very simple second-order hyperbolic equation for a composite dependent variable (which is directly proportional to a pressure-like component of this Green’s function and roughly corresponds to the strength of a monopole source within the jet). Our numerical calculations show that this quantity remains fairly close to the corresponding parallel flow result at low Mach numbers and that, as expected, it converges to that result when an appropriately scaled frequency parameter is increased. But the convergence occurs at progressively higher frequencies as the Mach number increases and the supersonic solution never actually converges to the parallel flow result in the vicinity of a critical- layer singularity that occurs in that solution. The dominant contribution to the propagator comes from the radial derivative of a certain component of the adjoint vector Green’s function. The non-parallel flow has a large effect on this quantity, causing it (and, therefore, the radiated sound) to increase at subsonic speeds and decrease at supersonic speeds. The effects of acoustic source location can be visualized by plotting the magnitude of this quantity, as function of position. These ‘altitude plots’ (which represent the intensity of the radiated sound as a function of source location) show that while the parallel flow solutions exhibit a single peak at subsonic speeds (when the source point is centred on the initial shear layer), the non-parallel solutions exhibit a double peak structure, with the second peak occurring about two potential core lengths downstream of the nozzle. These results are qualitatively consistent with the numerical calculations reported in Karabasov et al. (2011).

scholarly journals Strong earthquakes can be predicted: a multidisciplinary method for strong earthquake prediction

2003 ◽  
Vol 3 (6) ◽  
pp. 703-712 ◽  
Author(s):  
J. Z. Li ◽  
Z. Q. Bai ◽  
W. S. Chen ◽  
Y. Q. Xia ◽  
Y. R. Liu ◽  
...  
Keyword(s):  
Earthquake Prediction ◽  
Strong Earthquake ◽  
Low Frequency ◽  
Abnormal Behavior ◽  
Frequency Range ◽  
Strong Earthquakes ◽  
Physical Mechanisms ◽  
Innovative Methods ◽  
The World ◽  
Infrasonic Wave

Abstract. The imminent prediction on a group of strong earthquakes that occurred in Xinjiang, China in April 1997 is introduced in detail. The prediction was made on the basis of comprehensive analyses on the results obtained by multiple innovative methods including measurements of crustal stress, observation of infrasonic wave in an ultra low frequency range, and recording of abnormal behavior of certain animals. Other successful examples of prediction are also enumerated. The statistics shows that above 40% of 20 total predictions jointly presented by J. Z. Li, Z. Q. Ren and others since 1995 can be regarded as effective. With the above methods, precursors of almost every strong earthquake around the world that occurred in recent years were recorded in our laboratory. However, the physical mechanisms of the observed precursors are yet impossible to explain at this stage.

scholarly journals Songbirds use pulse tone register in two voices to generate low-frequency sound

2007 ◽  
Vol 274 (1626) ◽  
pp. 2703-2710 ◽  
Author(s):  
Kenneth K Jensen ◽  
Brenton G Cooper ◽  
Ole N Larsen ◽  
Franz Goller
Keyword(s):  
High Speed ◽  
Sound Production ◽  
Low Frequency ◽  
Pulse Generation ◽  
Sound Sources ◽  
Pulse Tone ◽  
Vocal Fold Dynamics ◽  
Vibration Pattern ◽  
Pulse Characteristics ◽  
Two Sides

The principal physical mechanism of sound generation is similar in songbirds and humans, despite large differences in their vocal organs. Whereas vocal fold dynamics in the human larynx are well characterized, the vibratory behaviour of the sound-generating labia in the songbird vocal organ, the syrinx, is unknown. We present the first high-speed video records of the intact syrinx during induced phonation. The syrinx of anaesthetized crows shows a vibration pattern of the labia similar to that of the human vocal fry register. Acoustic pulses result from short opening of the labia, and pulse generation alternates between the left and right sound sources. Spontaneously calling crows can also generate similar pulse characteristics with only one sound generator. Airflow recordings in zebra finches and starlings show that pulse tone sounds can be generated unilaterally, synchronously or by alternating between the two sides. Vocal fry-like dynamics therefore represent a common production mechanism for low-frequency sounds in songbirds. These results also illustrate that complex vibration patterns can emerge from the mechanical properties of the coupled sound generators in the syrinx. The use of vocal fry-like dynamics in the songbird syrinx extends the similarity to this unusual vocal register with mammalian sound production mechanisms.

Directional sound processing and interaural sound transmission in a small and a large grasshopper

1995 ◽  
Vol 198 (9) ◽  
pp. 1817-1827 ◽  
Author(s):  
A Michelsen ◽  
K Rohrseitz
Keyword(s):  
Outer Surface ◽  
Sound Pressure ◽  
Schistocerca Gregaria ◽  
Sound Transmission ◽  
Directional Hearing ◽  
Sound Processing ◽  
Low Frequencies ◽  
High Frequencies ◽  
Physical Mechanisms ◽  
Chorthippus Biguttulus

Physical mechanisms involved in directional hearing are investigated in two species of short-horned grasshoppers that differ in body length by a factor of 3­4. The directional cues (the effects of the direction of sound incidence on the amplitude and phase angle of the sounds at the ears) are more pronounced in the larger animal, but the scaling is not simple. At high frequencies (10­20 kHz), the sound pressures at the ears of the larger species (Schistocerca gregaria) differ sufficiently to provide a useful directionality. In contrast, at low frequencies (3­5 kHz), the ears must be acoustically coupled and work as pressure difference receivers. At 3­5 kHz, the interaural sound transmission is approximately 0.5 (that is, when a tympanum is driven by a sound pressure of unit amplitude at its outer surface, the tympanum of the opposite ear receives a sound pressure with an amplitude of 0.5 through the interaural pathway). The interaural transmission decreases with frequency, and above 10 kHz it is only 0.1­0.2. It still has a significant effect on the directionality, however, because the directional cues are large. In the smaller species (Chorthippus biguttulus), the interaural sound transmission is also around 0.5 at 5 kHz, but the directionality is poor. The reason for this is not the modest directional cues, but rather the fact that the transmitted sound is not sufficiently delayed for the ear to exploit the directional cues. Above 7 kHz, the transmission increases to approximately 0.8 and the transmission delay increases; this allows the ear to become more directional, despite the still modest directional cues.

Sound emission and the acoustic far field of a singing acridid grasshopper (Omocestus viridulus L.)

1999 ◽  
Vol 202 (12) ◽  
pp. 1571-1577
Author(s):  
A. Michelsen ◽  
N. Elsner
Keyword(s):  
Sound Production ◽  
Sound Intensity ◽  
Movement Pattern ◽  
Forward Direction ◽  
Octave Band ◽  
Sound Emission ◽  
Mean Values ◽  
The Mean ◽  
The One ◽  
Sound Intensities

An array of eight microphones, all at a distance of 15 cm, was used to make simultaneous recordings of the sounds emitted by courting male acridid grasshoppers of the species Omocestus viridulus. In this species, the movement pattern for sound production differs in the two hindlegs, and in most cases the leg facing the female moves with the larger amplitude. The sonic sound intensity (the total sound in the one-third octave bands with centre frequencies from 5 to 20 kHz) is maximal ipsilateral to the leg stridulating with the larger amplitude (the dominant leg). A spontaneous switch of dominance to the other leg may cause a significant change in the emitted sound power. The sound intensities contralateral to the dominant leg and frontal to the animal are, on average, approximately half (−3 dB) of the ipsilateral value, whereas the mean sound intensities behind and above the singer are approximately one-fifth (−7 dB) of the ipsilateral value. In most singers, the patterns of sound radiation are close to these mean values, but in some singers the radiation patterns are radically different. The sound radiated in various directions differs not only in terms of sound intensity but also with respect to the frequency spectrum, which was studied up to the one-third octave band with a centre frequency of 31.5 kHz. In particular, the ratio between the ultrasonic and sonic components is much smaller in the forward direction than in other directions. This may allow the courted female to hear whether the courting male is oriented directly towards her.

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