Phylogenetic Relationships Within and Delimitation of the Cosmopolitan Flowering Plant Genus Stellaria L. (Caryophyllaceae): Core Stars and Fallen Stars

2019 ◽  
Vol 44 (4) ◽  
pp. 857-876 ◽  
Author(s):  
Mathew T. Sharples ◽  
Erin A. Tripp
Keyword(s):  
Phylogenetic Relationships ◽  
New Genus ◽  
Flowering Plant ◽  
Herbaceous Plants ◽  
New Combinations ◽  
Eastern Asia ◽  
Number Of Species ◽  
New Synonymy ◽  
Levels Of Support ◽  
Stable Estimate

Abstract—Stellaria (Caryophyllaceae, tribe Alsineae) is a cosmopolitan genus of herbaceous plants that heretofore has lacked a stable estimate of total number of species. Here, we attempted to sample all species currently recognized under the name Stellaria to determine which belong to a core Stellaria genus versus which are masquerading under this name and are instead more closely related to one of several outgroup lineages. Phylogenetic relationships inferred with RAD loci were recovered with generally high levels of support, regardless of age of specimens (here up to 98 yr) utilized for molecular work. Analyses resolved a monophyletic, core radiation of Stellaria that is sister to Cerastium and others. Within core Stellaria, of which we sampled ca. 87% of species, we recovered five primary lineages. However, current infrageneric hypotheses are incongruous with phylogenomic data, and most morphology-based infrageneric hypotheses break down in light of these results. Outside of core Stellaria we recovered numerous other species currently ascribed to Stellaria that are in need of revised generic placement. Based on current results, however, we propose several new taxonomic and nomenclatural renovations within core Stellaria and external to this group as contribution towards stabilization of generic boundaries in Alsineae. These modifications include description of a new genus of Caryophyllaceae from eastern Asia, Nubelaria, as well description of a new genus, Rabelera, to accommodate the lineage previously and more widely known as Stellaria holostea. Nine new combinations are proposed in the genera Adenonema, Cerastium, Mesostemma, Nubelaria, Rabelera, and core Stellaria. Additionally, one new status, one new synonymy, and lectotypes for four species are proposed. We estimate core Stellaria to consist of approximately 112 species, a decrease from many other published species counts. Our results lay a robust foundation for future evolutionary, phylogenetic, and morphological comparisons within Stellaria and among relatives.

scholarly journals Taxonomic Studies of Some often Over-Looked Diaporthomycetidae and Sordariomycetidae

2021 ◽  
Author(s):  
Huang Shi-Ke ◽  
Kevin D. Hyde ◽  
Ausana Mapook ◽  
Sajeewa S.N. Maharachchikumbura ◽  
D. Jayarama Bhat ◽  
...  
Keyword(s):  
Phylogenetic Analysis ◽  
Phylogenetic Relationships ◽  
Type Species ◽  
New Genus ◽  
Molecular Data ◽  
New Combinations ◽  
Number Of Species ◽  
New Family ◽  
Taxonomic Studies

Abstract Sordariomycetes is an earlier and one of the widely distributed class of Ascomycota. The class was initially classified based on morphology in having inoperculate and unitunicate asci. With the development of DNA based phylogenetic analysis, several undetermined or polyphyletic members of Sordariomycetes were reclassified. However, not all species belonging to this class have been sequenced and analyzed. There are a number of species, especially those old and poorly studied ones which have never been sequenced before and not even recollected again for further taxonomic verification. One of the main objective in this study is to revise and update the taxonomy of several well-known old and poorly studied species whose classification are still obscure. Herein, we re-examined the type materials and/or authentic specimens together to explore 74 relatively poorly-studied genera, which mainly belong to Boliniales, Calosphaeriales, Chaetosphaeriales, Jobellisiales, and Sordariales classified under Diaporthomycetidae and Sordariomycetidae. We provide descriptions, notes, figures and/or drawings and discussed their phylogenetic relationships. As a result, the monotypic Jobellisiales is transferred from Hypocreomycetidae to Diaporthomycetidae. Based on phylogenetic analysis, the polyphyletic Lasiosphaeriaceae is divided into five families, Bombardiaceae (Apodospora, Bombardia, Bombardioidea and Fimetariella), Lasiosphaeriaceae (Anopodium, Bellojisia, Corylomyces, Lasiosphaeria, Mammaria and Zopfiella), Lasiosphaeridaceae (Lasiosphaeris), Strattoniaceae (Strattonia) and Zygospermellaceae (Episternus and Zygospermella). In addition, a new family Neoschizotheciaceae is established based on Neoschizothecium. Analysis of the type species of Boothiella, Stellatospora, Sulcatistroma and Tengiomyces placed them in Sordariaceae, Chaetomiaceae, Hypocreales and Coronophorales, respectively. We classify the genera lacking molecular data based on their morphology and expect them to be recollected; that is, Kacosphaeria in Calosphaeriales; Arnium, Biconiosporella, Camptosphaeria, Diffractella, Emblemospora, Eosphaeria, Periamphispora, Ramophialophora, Synaptospora and Tripterosporella in Sordariales; Conidiotheca in Sordariomycetes; Copromyces, Effetia, Endophragmiella and Tulipispora are accommodated in Ascomycota. Besides, we establish a new genus Neoschizothecium based on phylogenetic analysis. New combinations proposed include: Camaropella amorpha, Cam. microspora, Cam. plana, Cladorrhinum grandiusculum, Cla. leucotrichum, Cla. terricola, Cla. olerum, Helminthosphaeria plumbea, Immersiella hirta, Jugulospora minor, Lasiosphaeris arenicola, Neoschizothecium aloides, Neo. carpinicola, Neo. conicum, Neo. curvisporum, Neo. fimbriatum, Neo. glutinans, Neo. inaequale, Neo. minicaudum, Neo. selenosporum, Neo. tetrasporum, Neurospora autosteira, Podospora brunnescens, P. flexuosa, P. jamaicensis, P. hamata, P. macrospora, P. spinosa, Strattonia petrogale and Triangularia microsclerotigena, T. nannopodalis, T. praecox, T. samala, T. tarvisina, T. unicaudata, T. yaeyamensis. New epithets are proposed for Apiorhynchostoma apiosporum and Podospora dacryoidea.

Generic Reclassification of Limonius Eschscholtz, 1829 (Elateridae: Dendrometrinae) sensu Candèze 1860 of the World

Zootaxa ◽  
2019 ◽  
Vol 4683 (3) ◽  
pp. 301-335 ◽  
Author(s):  
FRANK E. ETZLER
Keyword(s):  
North American ◽  
New Genus ◽  
North American Species ◽  
New Combination ◽  
Valid Species ◽  
New Combinations ◽  
New Name ◽  
The World ◽  
New Synonymy

The genus Limonius Eschscholtz, 1829 was last treated as a whole by Candèze (1860). Since then, members have been placed in eight other genera: Cidnopus Thomson, 1859; Gambrinus LeConte, 1853; Elathous Reitter, 1890; Kibunea Kishii, 1966; Limoniscus Reitter, 1905; Nothodes LeConte, 1861; Pheletes Kiesenwetter, 1858; and Solskyana Dolin, 1978. Based on the examination of adult and larval characters, five genera are recognized: Elathous Reitter, 1890; Gambrinus LeConte, 1853; Limonius Eschscholtz, 1829; Pheletes Kiesenwetter, 1858; and Tetralimonius new genus. Limoniscus Reitter, 1905 and Sichuanelater Platia and Gudenzi, 2006 are new synonymies of Gambrinus LeConte, 1853; Micrathous Lane, 1971, Neoathousius Schimmel and Platia, 1991 and Solskyana Dolin, 1978 are all new synonymies of Limonius. A total of 84 new combinations are proposed: Nearctic: Elathous huguenini (Van Dyke, 1932) new combination; Gambrinus angulatus (Motschulsky, 1859) new combination; Gambrinus bicolor (Van Dyke, 1932) new combination; Gambrinus clypeatus (Motschulsky, 1859) new combination; Gambrinus confusus (LeConte, 1853) new combination; Gambrinus cribriceps (Van Dyke, 1943) new combination; Gambrinus crotchii (Horn, 1872) new combination; Gambrinus flavomarginatus (Knull, 1938) new combination; Gambrinus fulvipilis (Candèze, 1860) new combination; Gambrinus griseus (Beauvois, 1805) new combination; Gambrinus humidus (Lane, 1941) new combination; Gambrinus interstitialis (Melsheimer, 1846) new combination; Gambrinus lanchesteri (Lane, 1941) new combination; Gambrinus meridianus (Knull, 1947) new combination; Gambrinus mirus (LeConte, 1853) new combination; Gambrinus norahae (Al Dhafer, 2009) new combination; Gambrinus olentangyi (Knull, 1947) new combination; Gambrinus plebejus (Say, 1825) new combination; Gambrinus propexus (Candèze, 1860) new combination; Gambrinus rudis (Brown, 1933) new combination; Gambrinus rufihumeralis (Lane, 1941) new combination; Gambrinus seminudus (Van Dyke, 1932) new combination; Gambrinus shircki (Lane, 1965) new combination; Gambrinus sinuifrons (Fall, 1907) new combination; Gambrinus snakensis (Lane, 1965) new combination; Gambrinus stigma (Herbst, 1806) new combination; Gambrinus pictus (Van Dyke, 1932) new combination; Gambrinus ulkei (Horn, 1871) new combination; Gambrinus ursinus (Van Dyke, 1932) new combination; Gambrinus venablesi (Wickham, 1913) new combination; Limonius brevis (Van Dyke, 1932) new combination; Limonius sordidus (Van Dyke, 1932) new combination; Pheletes lecontei (Lane, 1971) new combination; Tetralimonius definitus (Ziegler, 1845) new combination; Tetralimonius humeralis (Candèze, 1860) new combination; Tetralimonius maculicollis (Motschulsky, 1860) new combination; Tetralimonius nimbatus (Say, 1825) new combination; Tetralimonius ornatulus (LeConte, 1857) new combination. Palearctic: Gambrinus elegans (Buysson, 1891) new combination; Gambrinus gibbosus (Platia and Gudenzi, 2006) new combination. Gambrinus henanensis (Schimmel, 2006) new combination; Gambrinus hinakurai (Kishii, 1998) new combination; Gambrinus katoi (Kishii, 2002) new combination; Gambrinus kawaharai (Kishii, 2002) new combination; Gambrinus kucerai (Schimmel, 2006) new combination; Gambrinus nanshanensis (Arimoto and Hiramatsu, 2013) new combination; Gambrinus naomii (Kishii, 1997) new combination; Gambrinus shaanxiensis (Schimmel, 2006) new combination; Gambrinus suturalis (Gebler, 1844) new combination; Gambrinus takabai (Kishii, 1997) new combination; Gambrinus violaceus (Müller, 1821) new combination; Gambrinus wittmeri (Chassain, 1998) new combination; Gambrinus yamato (Kishii, 1998) new combination; Gambrinus yujii (Arimoto, 2013) new combination; Gambrinus zhejiangensis (Schimmel, 2015) new combination; Limonius brancuccii (Schimmel and Platia, 1991) new combination; Limonius decorus (Gurjeva, 1975) new combination; Limonius exiguus (Schimmel and Platia, 1991) new combination; Limonius hartmanni (Schimmel, 1998) new combination; Limonius hiermeieri (Schimmel and Platia, 1991) new combination; Limonius hirtus (Dolin, 1978) new combination; Limonius hubeiensis (Kishii and Jiang, 1996) new combination; Limonius kubani (Schimmel, 1996) new combination; Limonius loebli (Schimmel and Platia, 1991) new combination; Limonius longicornis (Schimmel and Platia, 1991) new combination; Limonius macedonicus (Cate and Platia, 1989) new combination; Limonius marginellus brusteli (Leseigneur, 2004) new combination; Limonius manaliensis (Schimmel and Platia, 1991) new combination; Limonius miandamensis (Schimmel and Platia, 1991) new combination; Limonius minusculus (Schimmel and Platia, 1991) new combination; Limonius nigronitidus (Han and Lee, 2012) new combination; Limonius platiai (Mertlik, 1996) new combination; Limonius pseudopilosus (Platia and Gudenzi 1985) new combination; Limonius recticornis (Schimmel and Platia, 1991) new combination; Limonius riesei (Platia, 1988) new combination; Limonius rusticus (Schimmel and Platia, 1991) new combination; Limonius schurmanni (Platia and Gudenzi, 1998) new combination; Limonius sinensis (Schimmel and Platia, 1994) new combination; Limonius singularis (Schimmeland Platia, 1991) new combination; Limonius stapfi (Schimmel, 2007) new combination; Limonius turcicus (Platia, 2004) new combination; Limonius wittmeri (Schimmel and Platia, 1991) new combination; Tetralimonius quercus (Olivier, 1790) new combination; Tetralimonius reitteri (Gurjeva, 1976) new combination. The following 12 North American species are removed from synonymy and recognized as valid species: Gambrinus interstitialis (Melsheimer, 1846) status resurrected; Gambrinus propexus (Candèze, 1860) status resurrected; Gambrinus shircki (Lane, 1965) status resurrected; Gambrinus snakensis (Lane, 1965) status resurrected; Gambrinus ulkei (Horn, 1871) status resurrected; Limonius anceps LeConte, 1853 status resurrected; Limonius dubitans LeConte, 1853 status resurrected; Limonius infuscatus Motschulsky, 1859 status resurrected; Limonius pilosulus Candèze, 1891 status resurrected; Limonis semianeus LeConte, 1853 status resurrected. Tetralimonius humeralis (Candèze, 1860) status resurrected; Tetralimonius maculicollis (Motschulsky, 1860) status resurrected. New replacement names are proposed for three homynyms: Limonius schimmeli Etzler new name for Neoathousius ferrugineus Schimmel and Platia, 1991; Elathous malatyanus Etzler new name for Elathous bicolor Platia, 2010, not Elathous bicolor (LeConte, 1853); and Microdesmes carteri Etzler new name for Limonius angulatus Carter, 1939 (= Microdesmes angulatus). Limonius kondratieffi Al Dhafer, 2009 is a new synonymy of Elathous bicolor (LeConte, 1853). A key to genera, generic descriptions, notes on species, and definitions of important characters are provided. 

PHARPA, A NEW GENUS OF NEOTROPICAL AGATHIDINAE (HYMENOPTERA: BRACONIDAE) WITH A DISCUSSION OF PHYLOGENETIC RELATIONSHIPS

1986 ◽  
Vol 118 (12) ◽  
pp. 1231-1239 ◽  
Author(s):  
M. Sharkey
Keyword(s):  
Phylogenetic Relationships ◽  
New Genus ◽  
New Combinations ◽  
Southern Mexico

AbstractThe genus Pharpa is erected to include three species of Braconidae in the subfamily Agathidinae, viz. P. dubiosum (Szépligeti), P. basimacula (Cameron), and P. simulatrix (Cameron) all new combinations. Microdus albitarsis Cameron is newly synonymized with P. basimacula (Cameron). The genus Pharpa is distributed widely throughout the neotropics from southern Mexico to northern Argentina. Phylogenetic relationships with other agathidine genera are discussed, as is the possibility that the various species represent a polytypic, conspecific, mimetic complex.

A new genus and species of anophthalmous Otiorhynchini from Greece, with a new synonymy and new combinations (Coleoptera: Curculionidae, Entiminae)

Zootaxa ◽  
2021 ◽  
Vol 4938 (1) ◽  
pp. 69-84
Author(s):  
CESARE BELLO’ ◽  
ENZO COLONNELLI ◽  
LEONARDO FORBICIONI ◽  
GIUSEPPE OSELLA ◽  
ENRICO RUZZIER
Keyword(s):  
Type Species ◽  
New Genus ◽  
New Taxon ◽  
New Combinations ◽  
Unique Combination ◽  
Distinctive Pattern ◽  
New Genus And Species ◽  
Lateral Margins ◽  
New Synonymy

A new genus of Entiminae, an endogean weevil of the tribe Otiorhynchini, Giavarhynchus Bellò, Osella & Ruzzier, gen. n., and its type species Giavarhynchus amicorum Bellò, Osella & Ruzzier, sp. n. are described. The new taxon is readily distinguished from all other members of the tribe due to the unique combination of lack of eyes, elongate rostrum with a ventral transverse furrow and excised lateral margins located at apical third, punctation of pronotum of two distinct sizes arranged in a distinctive pattern, interval 7 of elytra protruding from base of pronotum and crenulate basally, metafemora bearing a spine-like tooth much larger than that of pro- and mesofemora, female tibiae granulate on inner margin, bisinuous mesotibiae. The following synonymy is proposed: Nematocerus Reiche, 1849 (= Cyrtozemia Pascoe, 1872, syn. n.; = Holcorhinosoma Voss, 1939, syn. n.). New combinations are: Nematocerus cognatus (Marshall, 1916), comb. n.; Nematocerus dispar (Pascoe, 1872), comb. n.; Nematocerus pilipes (Morimoto, 2015), comb. n., all from Cyrtozemia; Nematocerus subtuberculatus (Voss, 1939), comb. n. from Holcorhinosoma. New tribal placement is: Pseudocratopus Hustache, 1921 from Otiorhynchini to Peritelini. New subgeneric placement is that of Otiorhynchus deceptorius Białooki, Germann & Pelletier, 2017 and of Otiorhynchus incisirostris Białooki, Germann & Pelletier, 2017 from Otiorhynchus (Lixorrhynchus) Reitter, 1914 to Otiorhynchus (Aranihus) Reitter, 1912. 

Afrotheora, a new genus of primitive Hepialidae from Africa (Lepidoptera: Hepialoidea)

1986 ◽  
Vol 17 (1) ◽  
pp. 29-54 ◽  
Author(s):  
Malcolm J. Scoble ◽  
Ebbe S. Nielsen
Keyword(s):  
South Africa ◽  
New Species ◽  
Southern Africa ◽  
New Genus ◽  
Compound Eye ◽  
Sensu Stricto ◽  
New Taxon ◽  
New Combinations ◽  
Senior Synonym ◽  
New Synonymy

AbstractAfrotheora, a new hepialoid genus from central and southern Africa, is described. There are eight species of which seven are named. Three were described previously, but were assigned to other genera (Dalaca rhodaula Meyrick, Eudalaca jordani Viette, and Hepialus thermodes Meyrick - a new senior synonym of Hepialus pardalias Janse). Four new species are named and described (minirhodaula, argentimaculata, flavimaculata and brevivalva). A further new species is described, but is not formally named. All species and their genitalia are described and illustrated. Three new combinations and one new synonymy are established, two lectotypes are designated and Hepialus ptiloscelis Meyrick from South Africa is transferred to Gorgopis Hübner. The monophyly of Afrotheora is recognized by the possession of two unique characters: (1) long bristle-like setae from the antennal scape reaching almost across the compound eye, and (2) the trulleum in the male genitalia comprising two lateral sclerotized rods separated by a membrane. Afrotheora represents one of the 12 hepialoid basal lineages currently thought to be monophyletic, and it is demonstrated that the new taxon is not subordinate to any other of these hepialoid clades. Its relationships are briefly discussed, but its exact affinities await further studies of hepialoid phylogeny. The term 'primitive Hepialidae' is here applied to four genera: Fraus Walker, Gazoryctra Hübner, Antihepialus Janse and Afrotheora. We use 'Hepialidae sensu stricto' to refer to the remaining genera of the Hepialidae sensulato (i.e. the Hepialidae of authors) until the phylogeny of the Hepialoidea is better understood. This does not indicate that the primitive Hepialidae are monophyletic while the Hepialidae sensu stricto undoubtedly are.

A NEW GENUS BROOKSELLA, NEAR ILNACORA REUTER, WITH NEW SYNONYMY AND NEW COMBINATIONS FOR 15 SPECIES CURRENTLY PLACED IN MELANOTRICHUS REUTER (HETEROPTERA: MIRIDAE)

1979 ◽  
Vol 111 (8) ◽  
pp. 949-954 ◽  
Author(s):  
Leonard A. Kelton
Keyword(s):  
North American ◽  
Male Genitalia ◽  
Type Species ◽  
New Genus ◽  
North American Species ◽  
New Combinations ◽  
New Synonymy

AbstractBrooksella n. gen. is described (type-species, Asciodema inconspicua Uhler). Lectotypes are designated for Asciodema inconspicua Uhler and Orthotylus viridicatus Uhler. The following North American species currently in Melanotrichus Reuter are transferred to Brooksella: Asciodema inconspicua Uhler (= M. atricornis Knight, n. syn.), M. chelifer Knight (= M. brevirostris Knight, M. custeri Knight, n. syn.), M. azteci Knight, M. incurvus Knight, M. malvastri Knight, M. nevadensis Knight, M. nicholi Knight, M. shoshonea Knight, Orthotylus althaeae Hussey, O. ferox Van Duzee, O. tibialis Van Duzee, and O. viridicatus Uhler. Male genitalia and tergal process are illustrated for each species and a key to separate them is provided.

A REVIEW OF THE SPECIES OF NITIDOTACHINUS NEW GENUS (COLEOPTERA: STAPHYLINIDAE: TACHYPORINAE)

1993 ◽  
Vol 125 (3) ◽  
pp. 521-548 ◽  
Author(s):  
J.M. Campbell
Keyword(s):  
North America ◽  
Phylogenetic Relationships ◽  
New Genus ◽  
New Combinations ◽  
Diagnostic Characters ◽  
Line Drawings ◽  
Scanning Electron

AbstractA new genus, Nitidotachinus, is described in the subfamily Tachyporinae (Staphylinidae) to include the species formerly placed in the Tachyporoides and Impunctatus groups of Tachinus Gravenhorst and Tachinus excellens Bernhauer. The new genus includes 10 species, all previously described, from North America, Japan, China, and Taiwan. All major diagnostic characters for the genus are compared with those of Tachinus and illustrated with scanning electron photomicrographs or line drawings. The following new combinations are created by transferring the species from Tachinus to the new genus Nitidotachinus: N. adachii (Watanabe and Shibata), N. agilis (Horn), N. excellens (Bernhauer), N. horni (Campbell), N. impunctatus (Sharp), N. lanei (Hatch), N. sawadai (Watanabe and Shibata), N. scrutator (Gemminger and Harold), N. tachyporoides (Horn), and N. taiwanensis (Shibata).The 10 species of the genus are redescribed; the aedeagus is illustrated for all species, except N. excellens Bernhauer, known only from a unique female. A key is provided to aid in distinguishing the species and the phylogenetic relationships of the species of the genus are discussed.

Checklist of British Lichen-Forming, Lichenicolous And Allied Fungi

1980 ◽  
Vol 12 (1) ◽  
pp. 1-115 ◽  
Author(s):  
D. L. Hawksworth ◽  
P. W. James ◽  
B. J. Coppins
Keyword(s):  
Original Work ◽  
New Genus ◽  
New Combinations ◽  
British Isles ◽  
Number Of Species ◽  
First Time

AbstractA new checklist of the lichen-forming, lichenicolous and allied fungi occuring in the British Isles (including Ireland) is presented. The total number of species accepted is 1701, distributed through 294 genera. Of these species 1471 are lichen-forming, 183 lichenicolous, and 47 allied fungi. In addition to incorporating the results of investigations published since the last checklishts, the list embodies a great deal of original work by the authors and their collaborators; as a result a considerable number of changes in nomenclature are made here for the first time, including one new genus (Herteliana) and 56 new combinations. Names utilized in previous checklists are cross-referenced and relevant papers cited under generic heads.

TRANSFER OF THE TARSOCHEYLIDAE TO THE HETEROSTIGMATA, AND REASSIGNMENT OF THE TARSONEMINA AND HETEROSTIGMATA TO LOWER HIERARCHIC STATUS IN THE PROSTIGMATA (ACARI)

1976 ◽  
Vol 108 (1) ◽  
pp. 23-48 ◽  
Author(s):  
Evert E. Lindquist
Keyword(s):  
North America ◽  
Phylogenetic Relationships ◽  
Type Species ◽  
New Genus ◽  
Western North America ◽  
Undescribed Species ◽  
Morphological Criteria ◽  
The Family ◽  
Collection Data ◽  
New Synonymy

AbstractThe systematic status of the genus Tarsocheylus Berlese, 1904 and the position of the family Tarsocheylidae Atyeo and Baker, 1964 in the Prostigmata is reviewed. The family does not belong in either the Raphignathoidea or the Anystoidea but in its own superfamily, the Tarsocheyloidea, which is described for this purpose. In turn, this superfamily is placed alongside the Heterocheyloidea in the Heterostigmata.Because the type-species of Tarsocheylus, T. paradoxus Berlese, 1904, is conspecific with Hoplocheylus discalis Atyeo and Baker, 1964 (new synonymy), the concept of Hoplocheylus Atyeo and Baker, 1964, which has remained until now as originally proposed, is the same as that of Tarsocheylus. However, a new case is made for recognizing these genera as mutually distinct, based on other morphological criteria. Tarsocheylus atomarius Berlese, 1913, the type-species of Hoplocheylus, is apparently conspecific with Hoplocheylus canadensis Marshall, 1966 (new synonymy) and perhaps with one or two other described species from North America. A third generic entity, represented by Hoplocheylus johnstoni Atyeo and Baker, 1964, is noted but not named or formally described. Hoplocheylus pickardi Smiley and Moser, 1968 is conspecific with H. similis Delfinado and Baker, 1974 (new synonymy).Hemitarsocheylus Soliman and Zaher, 1975, which was described as a new genus in the Tarsocheylidae, is congeneric with Stigmocheylus Berlese, 1910 (new synonymy). This genus does not belong in the Tarsocheyloidea, but rather in the Anystoidea.A list of character states found useful in recognizing groupings of species of Tarsocheylidae is presented. Based on the collection data with specimens representing described and undescribed species at hand, the known distribution of the Tarsocheylidae is extended to include Australia and western North America, and the second known instance of an association with passalid beetles is recorded.From a phylogenetic standpoint, the Tarsocheyloidea and Heterocheyloidea are proposed as sister groups that together constitute a new suprafamilial taxon, the Tarsocheylina. In turn, the Tarsocheylina and Tarsonemina are proposed as sister groups, and both are described; together, they form the Heterostigmata.The systematic and phylogenetic relationships of the Heterostigmata (including the Tarsonemina) to the Prostigmata and the Astigmata are reviewed. Not only should the Heterostigmata be retained in the Prostigmata but it should be lowered in hierarchic rank, such that it is a part of the Eleutherengona. The stock that gave rise to the Anystoidea and related superfamilies is shown to be ancestral to the Heterostigmata as well.

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