A latest Eocene (Chadronian) brontothere (Mammalia, Perissodactyla) from the Antero Formation, South Park, Colorado

2021 ◽  
Vol 56 (1) ◽  
pp. 37-50
Author(s):  
Rory E. Sweedler ◽  
Jaelyn J. Eberle ◽  
Matthew C. Mihlbachler
Keyword(s):  
North America ◽  
High Altitude ◽  
Great Plains ◽  
Late Eocene ◽  
Lower Incisor ◽  
South Park ◽  
Exact Identification ◽  
Latest Eocene ◽  
Land Mammal ◽  
Insight Into

ABSTRACT Late Eocene brontotheres are documented most prevalently from formations in the Great Plains of North America. Here we describe UCM 109045, a mandible and lower dentition of a brontothere recovered from a latest Eocene (Chadronian) locality in the Antero Formation in South Park, Colorado. This is a high-altitude locality in which vertebrate fossils are rare. Lower incisor number and presence of a long postcanine diastema indicate that UCM 109045 does not belong to Megacerops coloradensisLeidy, 1870, by far the most abundant brontothere from the Chadronian North American Land Mammal Age. Instead, UCM 109045 is morphologically most similar to Protitanops curryiStock, 1936, from the early Chadronian of the southwestern United States, and nomen dubium Megacerops primitivusLambe, 1908, from the Chadronian of Saskatchewan, Canada. It is possible that Megacerops kuwagatarhinusMader and Alexander, 1995, is a junior synonym of M. primitivus. If UCM 109045 belongs to Megacerops primitivus (= M. kuwgatarhinus), it would support the hypothesis that only two species of brontothere—M. primitivus (= M. kuwgatarhinus) and M. coloradensis—survived into the latest Eocene. Regardless of its exact identification, the discovery of UCM 109045 in the Antero Formation provides insight into a poorly understood, high-altitude locality in North America from just before brontothere extinction at the Eocene–Oligocene boundary.

Marsupials (Mammalia, Metatheria) from the Brule Formation (Whitneyan, Oligocene) North Dakota

2020 ◽  
Vol 95 (1) ◽  
pp. 193-204
Author(s):  
William W. Korth ◽  
Clint A. Boyd ◽  
Robert J. Emry ◽  
Jeff J. Person
Keyword(s):  
New Species ◽  
North America ◽  
North Dakota ◽  
Great Plains ◽  
Single Specimen ◽  
Marsupial Species ◽  
Fourth Species ◽  
Latest Eocene ◽  
Land Mammal ◽  
Brule Formation

AbstractFive marsupial species are recognized from the Brule Formation at two localities in southwestern North Dakota: Fitterer Ranch and Obritsch Ranch (middle Oligocene; Whitneyan North American Land Mammal Age [NALMA]). The herpetotheriids Herpetotherium fugax Cope, 1873a, Copedelphys superstes new species, and the peradectid Nanodelphys hunti (Cope, 1873b) are represented at both localities. A fourth species is H. sp., cf. H. merriami (Stock and Furlong, 1922), represented by a single specimen from Fitterer Ranch, being limited elsewhere to the later Arikareean NALMA. A fifth species is represented by two isolated lower cheek teeth, interpreted as m1s, from Fitterer Ranch that are unique in lacking a trigonid (only two cusps present) while having a well-developed talonid. These specimens are referred to an indeterminate herpetotheriine species. The new species of Copedelphys is distinct from other species of the genus in that the anterior two lower molars are enlarged relative to the posterior molars. Overall, this new species is more similar in proportions to the latest Eocene (Chadronian) C. titanelix (Matthew, 1903) than the Oligocene (Orellan and Whitneyan) C. stevensoni (Cope, 1873b). This study adds a third and fourth Whitneyan marsupial fauna from the Great Plains region of North America, increases the known diversity of Whitneyan marsupials, and provides further evidence that marsupial diversity during the late Paleogene in North America was relatively stable until the late early Arikareean NALMA.UUID: http://zoobank.org/b8534802-b9ed-4120-baca-fc72917f7d6a

scholarly journals Additional records and stratigraphic distribution of the middle Eocene carettochelyid turtle Anosteira pulchra from the Uinta Formation of Utah, North America

PeerJ ◽  
2020 ◽  
Vol 8 ◽  
pp. e9775
Author(s):  
Brent Adrian ◽  
Patricia A. Holroyd ◽  
J. Howard Hutchison ◽  
KE Beth Townsend
Keyword(s):  
North America ◽  
New Records ◽  
Middle Eocene ◽  
Single Species ◽  
Uinta Basin ◽  
Stratigraphic Framework ◽  
Multiple Levels ◽  
Land Mammal ◽  
Anatomical Information ◽  
Insight Into

Background Anosteira pulchra is one of two species of the obligately-aquatic freshwater clade Carettochelyidae (pig-nosed turtles) from the Eocene of North America. Anosteira pulchra is typically rare in collections, and their distribution is poorly documented. The Uinta Formation [Fm.] contains a diverse assemblage of turtles from the Uintan North American Land Mammal Age. Whereas turtles are abundantly preserved in the Uinta Fm., A. pulchra has been reported only from a few specimens in the Uinta C Member. Methods We describe new records of Anosteira pulchra from the Uinta Basin and analyze the distribution of 95 specimens from multiple repositories in the previously published stratigraphic framework of the middle and upper Uinta Fm. Results Here we report the first records of the species from the Uinta B interval, document it from multiple levels within the stratigraphic section and examine its uncommon appearance in only approximately 5% of localities where turtles have been systematically collected. This study details and extends the range of A. pulchra in the Uinta Fm. and demonstrates the presence of the taxon in significantly lower stratigraphic layers. These newly described fossils include previously unknown elements and associated trace fossils, with new anatomical information presented. This study provides insight into the taxonomy of Anosteira spp. in the middle Eocene, and suggests the presence of a single species, though no synonymy is defined here due to limits in Bridger material.

scholarly journals Whence the beardogs? Reappraisal of the Middle to Late Eocene ‘ Miacis ’ from Texas, USA, and the origin of Amphicyonidae (Mammalia, Carnivora)

2016 ◽  
Vol 3 (10) ◽  
pp. 160518 ◽  
Author(s):  
Susumu Tomiya ◽  
Zhijie Jack Tseng
Keyword(s):  
North America ◽  
Great Plains ◽  
Geographical Origin ◽  
Middle Eocene ◽  
Cladistic Analysis ◽  
Rocky Mountain ◽  
Late Eocene ◽  
Northern Great Plains ◽  
Mammalian Fauna ◽  
Additional Support

The Middle to Late Eocene sediments of Texas have yielded a wealth of fossil material that offers a rare window on a diverse and highly endemic mammalian fauna from that time in the southern part of North America. These faunal data are particularly significant because the narrative of mammalian evolution in the Paleogene of North America has traditionally been dominated by taxa that are known from higher latitudes, primarily in the Rocky Mountain and northern Great Plains regions. Here we report on the affinities of two peculiar carnivoraforms from the Chambers Tuff of Trans-Pecos, Texas, that were first described 30 years ago as Miacis cognitus and M. australis . Re-examination of previously described specimens and their inclusion in a cladistic analysis revealed the two taxa to be diminutive basal amphicyonids; as such, they are assigned to new genera Gustafsonia and Angelarctocyon , respectively. These two taxa fill in some of the morphological gaps between the earliest-known amphicyonid genus, Daphoenus , and other Middle-Eocene carnivoraforms, and lend additional support for a basal caniform position of the beardogs outside the Canoidea. The amphicyonid lineage had evidently given rise to at least five rather distinct forms by the end of the Middle Eocene. Their precise geographical origin remains uncertain, but it is plausible that southern North America served as an important stage for a very early phase of amphicyonid radiation.

scholarly journals Hyainailourine and teratodontine cranial material from the late Eocene of Egypt and the application of parsimony and Bayesian methods to the phylogeny and biogeography of Hyaenodonta (Placentalia, Mammalia)

PeerJ ◽  
2016 ◽  
Vol 4 ◽  
pp. e2639 ◽  
Author(s):  
Matthew R. Borths ◽  
Patricia A. Holroyd ◽  
Erik R. Seiffert
Keyword(s):  
North America ◽  
Bayesian Methods ◽  
Middle Eocene ◽  
Phylogenetic Analyses ◽  
Late Eocene ◽  
Biogeographic History ◽  
Close Relationship ◽  
History Of ◽  
New Material ◽  
Latest Eocene

Hyaenodonta is a diverse, extinct group of carnivorous mammals that included weasel- to rhinoceros-sized species. The oldest-known hyaenodont fossils are from the middle Paleocene of North Africa and the antiquity of the group in Afro-Arabia led to the hypothesis that it originated there and dispersed to Asia, Europe, and North America. Here we describe two new hyaenodont species based on the oldest hyaenodont cranial specimens known from Afro-Arabia. The material was collected from the latest Eocene Locality 41 (L-41, ∼34 Ma) in the Fayum Depression, Egypt.Akhnatenavus nefertiticyonsp. nov. has specialized, hypercarnivorous molars and an elongate cranial vault. InA. nefertiticyonthe tallest, piercing cusp on M1–M2is the paracone.Brychotherium ephalmosgen. et sp. nov. has more generalized molars that retain the metacone and complex talonids. InB. ephalmosthe tallest, piercing cusp on M1–M2is the metacone. We incorporate this new material into a series of phylogenetic analyses using a character-taxon matrix that includes novel dental, cranial, and postcranial characters, and samples extensively from the global record of the group. The phylogenetic analysis includes the first application of Bayesian methods to hyaenodont relationships.B. ephalmosis consistently placed within Teratodontinae, an Afro-Arabian clade with several generalist and hypercarnivorous forms, andAkhnatenavusis consistently recovered in Hyainailourinae as part of an Afro-Arabian radiation. The phylogenetic results suggest that hypercarnivory evolved independently three times within Hyaenodonta: in Teratodontinae, in Hyainailourinae, and in Hyaenodontinae. Teratodontines are consistently placed in a close relationship with Hyainailouridae (Hyainailourinae + Apterodontinae) to the exclusion of “proviverrines,” hyaenodontines, and several North American clades, and we propose that the superfamily Hyainailouroidea be used to describe this relationship. Using the topologies recovered from each phylogenetic method, we reconstructed the biogeographic history of Hyaenodonta using parsimony optimization (PO), likelihood optimization (LO), and Bayesian Binary Markov chain Monte Carlo (MCMC) to examine support for the Afro-Arabian origin of Hyaenodonta. Across all analyses, we found that Hyaenodonta most likely originated in Europe, rather than Afro-Arabia. The clade is estimated by tip-dating analysis to have undergone a rapid radiation in the Late Cretaceous and Paleocene; a radiation currently not documented by fossil evidence. During the Paleocene, lineages are reconstructed as dispersing to Asia, Afro-Arabia, and North America. The place of origin of Hyainailouroidea is likely Afro-Arabia according to the Bayesian topologies but it is ambiguous using parsimony. All topologies support the constituent clades–Hyainailourinae, Apterodontinae, and Teratodontinae–as Afro-Arabian and tip-dating estimates that each clade is established in Afro-Arabia by the middle Eocene.

A latest Chadronian (Late Eocene) mammalian fauna from the Cypress Hills, Saskatchewan

1994 ◽  
Vol 31 (8) ◽  
pp. 1335-1341 ◽  
Author(s):  
John E. Storer
Keyword(s):  
North America ◽  
Strong Evidence ◽  
Late Eocene ◽  
Mammalian Fauna ◽  
Local Fauna ◽  
Early Oligocene ◽  
Land Mammal

The Kealey Springs West (KSW) local fauna from the Cypress Hills Formation (Eocene to Miocene), Saskatchewan, is probably latest Chadronian in age, and is the northernmost assemblage representing a time near the Eocene–Oligocene boundary in North America. Presence of Pseudocylindrodon neglectus is strong evidence that the assemblage is Chadronian. Herpetotherium valens, Protosciurus, Namatomys cf. N. lloydi, and Centimanomys also suggest a Chadronian age. The assemblage contains Adjidaumo maximus and Palaeolagus cf. P. hemirhizis, species that have been cited as characteristic of the early part of the Orellan Land Mammal Age (early Oligocene), and the typically Orellan Heliscomys hatcheri and Eumys elegans. Several other tara in the KSW assemblage are known from both the Chadronian (late Eocene) and Orellan.

The Neogene transition from C3 to C4 grasslands in North America: assemblage analysis of fossil phytoliths

Paleobiology ◽  
2011 ◽  
Vol 37 (1) ◽  
pp. 50-71 ◽  
Author(s):  
Caroline A. E. Strömberg ◽  
Francesca A. McInerney
Keyword(s):  
North America ◽  
Great Plains ◽  
Late Miocene ◽  
Vegetation Change ◽  
Stable Carbon Isotopes ◽  
Fire Regime ◽  
Tooth Enamel ◽  
Regional Scale ◽  
Grassland Ecosystems ◽  
Isotopic Records

The rapid ecological expansion of grasses with C4 photosynthesis at the end of the Neogene (8–2 Ma) is well documented in the fossil record of stable carbon isotopes. As one of the most profound vegetation changes to occur in recent geologic time, it paved the way for modern tropical grassland ecosystems. Changes in CO2 levels, seasonality, aridity, herbivory, and fire regime have all been suggested as potential triggers for this broadly synchronous change, long after the evolutionary origin of the C4 pathway in grasses. To date, these hypotheses have suffered from a lack of direct evidence for floral composition and structure during this important transition. This study aimed to remedy the problem by providing the first direct, relatively continuous record of vegetation change for the Great Plains of North America for the critical interval (ca. 12–2 Ma) using plant silica (phytolith) assemblages.Phytoliths were extracted from late Miocene-Pliocene paleosols in Nebraska and Kansas. Quantitative phytolith analysis of the 14 best-preserved assemblages indicates that habitats varied substantially in openness during the middle to late Miocene but became more uniformly open, corresponding to relatively open grassland or savanna, during the late Miocene and early Pliocene. Phytolith data also point to a marked increase of grass short cells typical of chloridoid and other potentially C4 grasses of the PACMAD clade between 8 and 5 Ma; these data suggest that the proportion of these grasses reached up to ∼50–60% of grasses, resulting in mixed C3-C4 and highly heterogeneous grassland communities by 5.5 Ma. This scenario is consistent with interpretations of isotopic records from paleosol carbonates and ungulate tooth enamel. The rise in abundance of chloridoids, which were present in the central Great Plains since the early Miocene, demonstrates that the “globally” observed lag between C4 grass evolution/taxonomic diversification and ecological expansion occurred at the regional scale. These patterns of vegetation alteration imply that environmental change during the late Miocene-Pliocene played a major role in the C3-C4 shift in the Great Plains. Specifically, the importance of chloridoids as well as a decline in the relative abundance of forest indicator taxa, including palms, point to climatic drying as a key trigger for C4 dominance.

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