The seed capacity of red pine cones varies from about 30 to over 110, depending on the size of the cone and its position in the tree crown, and is determined by the number of ovules that are structurally complete at the time of pollination. These ovules occur in a central "productive" region and constitute less than one-half of the total. The remaining ovules, most: of which are in the proximal part of the cone, never become structurally perfect, and do not contribute to seed production. Abortion of ovules in the productive region usually reduces seed production efficiency to 50–60%, and is accompanied mainly by withering of the nucellus in the first year and failure to produce archegonia early in the second year. The extent of ovule abortion during the first year varies indirectly with cone size, seed capacity, and height in tree.