scholarly journals Stress assignment in Hittite and Proto-Indo-European

2016 ◽  
Vol 1 ◽  
pp. 25
Author(s):  
Anthony D Yates

This paper develops a new optimality theoretic analysis of lexical accent in Hittite (Anatolian, Indo-European). I demonstrate that Hittite synchronic stress assignment is consistent with Kiparsky and Halle's (1977) Basic Accentuation Principle, which assigns primary stress to the leftmost morpheme lexically specified for prosodic prominence or else to the left edge of a prosodic word. The Hittite evidence is thus shown to converge with Kiparsky and Halle's reconstruction of this principle for the common ancestor of the non-Anatolian Indo-European languages (i.e. Proto-Nuclear-Indo-European), and in view of this agreement, argued to be reconstructible for Proto-Indo-European itself.

Author(s):  
Anthony D Yates

This paper develops a new, optimality-theoretic analysis of word-level stress assignment in Cupeño (Takic, Uto-Aztecan). I argue that primary stress is assigned to the leftmost lexically accented (i.e. stress-preferring) morpheme, else to the word's left edge. I contend that this analysis is simpler and better explains the Cupeño data than previous accounts, which assume that special faithfulness constraints privilege the accentual properties of roots over those of other affixes. The typological implications of this renanalysis of Cupeño stress are then discussed; without empirical support from Cupeño, it is suggested that "root faithfulness" plays no role in determining word stress in lexical accent systems cross-linguistically.


2018 ◽  
Vol 60 (2) ◽  
pp. 7-24 ◽  
Author(s):  
Mohammed Nour Abu Guba

Abstract This paper proposes an Optimality-Theoretic analysis of stress assignment in Levantine Arabic. The proposed hierarchy incorporates two constraints, namely *EXTENDED-LAPSE-R, which restricts stress to one of the last three syllables, and ALIGN-LEFT, which demands that the left edge of the prosodic word be aligned with a foot. This hierarchy is superior to earlier research as it successfully accounts for stress assignment in a more comprehensive and economical way. Most interestingly, it can account for the unexpected stress on a light penult in prosodic words ending in four light syllables and the paradoxical status of foot extrametricality without ad hoc parameterization of constraints. Moreover, findings show that footing in Levantine Arabic is iterative, an indication that secondary stress is attested in Levantine dialects.


Entrepalavras ◽  
2020 ◽  
Vol 10 (1) ◽  
pp. 36
Author(s):  
Gabriel Antunes de Araujo ◽  
Manuele Bandeira ◽  
Ana Lívia Agostinho

The Proto-Creole of the Gulf of Guinea is the common ancestor of the Santome, Angolar, Lung’Ie and Fa d’Ambô languages. Based on a supposed occurrence of vowel harmony (vh) in these modern languages and the proven influence of the languages of the Niger Delta (HAGEMEIJER, 2009), in which vh is attested, this study aims to discuss the existence of vh processes with [atr] mid-vowels in the Proto-Creole of the Gulf of Guinea, considering as a starting point its phonological and lexical reconstruction (BANDEIRA, 2017). Therefore, we intend to verify what phonological parameters related to vh in the literature (trigger, target, domain, application direction and blocking element) indicate about vh processes in Proto-Creole, based on the lexical reflexes of a set of cognates in daughter languages and proto-forms. We will show that many of the items interpreted as harmonic derive from the maintenance of mid-vowels in Portuguese etyma and from the insertion of a vowel copy in order to avoid roots ending in consonants in the Proto-Creole of the Gulf of Guinea. In this analysis, we will argue for the hypothesis according to which parasitic vh processes (COLE; TRIGO, 1988) with [atr] mid-vowels in contiguous syllables occurred in Proto-Creole within the prosodic word domain. However, the vh was limited because, based on the observation of proto-forms and harmonisation patterns with [atr] mid-vowels in contiguous syllables, there were many cases in which there was neutralisation of final non-stressed mid-vowels and, therefore, no application of the harmony process in favouring contexts.


1983 ◽  
Vol 38 (5-6) ◽  
pp. 501-504 ◽  
Author(s):  
Mária Ujhelyi

Seryl tRNA (anticodon GCU) from mammalian mito­chondria shows in comparison to other mitochondrial tRNAs additional special features differing from the generalized tRNA model. When arranged in the tradi­tional cloverleaf form, eight bases fall within the TΨC loop, and the entire dihydrouridine loop is lacking. This seryl tRNA molecule is therefore shorter than other tRNAs. It was originally thought to represent a mito­chondrial analogon of 5 S rRNA and its precise classifica­tion is still disputed. The present studies suggest that this mitochondrial tRNA represents a fossil molecule which is related to the common ancestor of the present tRNA and 5 S rRNA molecules.


2013 ◽  
Vol 2013 ◽  
pp. 1-5 ◽  
Author(s):  
Jeffrey S. Prince ◽  
Paul Micah Johnson

The ultrastructure of the digestive gland of several sea hare species that produce different colored ink (Aplysia californicaproduces purple ink,A. julianawhite ink,A. parvulaboth white and purple ink, whileDolabrifera dolabriferaproduces no ink at all) was compared to determine the digestive gland’s role in the diet-derived ink production process. Rhodoplast digestive cells and their digestive vacuoles, the site of digestion of red algal chloroplast (i.e., rhodoplast) inA. californica, were present and had a similar ultrastructure in all four species. Rhodoplast digestive cell vacuoles either contained a whole rhodoplast or fragments of one or were empty. These results suggest that the inability to produce colored ink in some sea hare species is not due to either an absence of appropriate digestive machinery, that is, rhodoplast digestive cells, or an apparent failure of rhodoplast digestive cells to function. These results also propose that the digestive gland structure described herein occurred early in sea hare evolution, at least in the common ancestor to the generaAplysiaandDolabrifera. Our data, however, do not support the hypothesis that the loss of purple inking is a synapomorphy of the white-ink-producing subgenusAplysia.


Parasitology ◽  
2014 ◽  
Vol 142 (S1) ◽  
pp. S120-S127 ◽  
Author(s):  
GARETH D. WEEDALL ◽  
NEIL HALL

SUMMARYA key part of the life cycle of an organism is reproduction. For a number of important protist parasites that cause human and animal disease, their sexuality has been a topic of debate for many years. Traditionally, protists were considered to be primitive relatives of the ‘higher’ eukaryotes, which may have diverged prior to the evolution of sex and to reproduce by binary fission. More recent views of eukaryotic evolution suggest that sex, and meiosis, evolved early, possibly in the common ancestor of all eukaryotes. However, detecting sex in these parasites is not straightforward. Recent advances, particularly in genome sequencing technology, have allowed new insights into parasite reproduction. Here, we review the evidence on reproduction in parasitic protists. We discuss protist reproduction in the light of parasitic life cycles and routes of transmission among hosts.


Author(s):  
Satoshi Nakano ◽  
Takao Fujisawa ◽  
Bin Chang ◽  
Yutaka Ito ◽  
Hideki Akeda ◽  
...  

After the introduction of the seven-valent pneumococcal conjugate vaccine, the global spread of multidrug resistant serotype 19A-ST320 strains became a public health concern. In Japan, the main genotype of serotype 19A was ST3111, and the identification rate of ST320 was low. Although the isolates were sporadically detected in both adults and children, their origin remains unknown. Thus, by combining pneumococcal isolates collected in three nationwide pneumococcal surveillance studies conducted in Japan between 2008 and 2020, we analyzed 56 serotype 19A-ST320 isolates along with 931 global isolates, using whole-genome sequencing to uncover the transmission route of the globally distributed clone in Japan. The clone was frequently detected in Okinawa Prefecture, where the U.S. returned to Japan in 1972. Phylogenetic analysis demonstrated that the isolates from Japan were genetically related to those from the U.S.; therefore, the common ancestor may have originated in the U.S. In addition, Bayesian analysis suggested that the time to the most recent common ancestor of the isolates form Japan and the U.S. was approximately the 1990s to 2000, suggesting the possibility that the common ancestor could have already spread in the U.S. before the Taiwan 19F-14 isolate was first identified in a Taiwanese hospital in 1997. The phylogeographical analysis supported the transmission of the clone from the U.S. to Japan, but the analysis could be influenced by sampling bias. These results suggested the possibility that the serotype 19A-ST320 clone had already spread in the U.S. before being imported into Japan.


2022 ◽  
Author(s):  
Liqi Yao ◽  
Clay Clark

All caspases evolved from a common ancestor and subsequently developed into two general classes, inflammatory or apoptotic caspases. The caspase-hemoglobinase fold has been conserved throughout nearly one billion years of evolution and is utilized for both the monomeric and dimeric subfamilies of apoptotic caspases, called initiator and effector caspases, respectively. We compared the folding and assembly of procaspase-3b from zebrafish to that of human effector procaspases in order to examine the conservation of the folding landscape. Urea-induced equilibrium folding/unfolding of procaspase-3b showed a minimum three-state folding pathway, where the native dimer isomerizes to a partially folded dimeric intermediate, which then unfolds. A partially folded monomeric intermediate observed in the folding landscape of human procaspase-3 is not well-populated in zebrafish procaspase-3b. By comparing effector caspases from different species, we show that the effector procaspase dimer undergoes a pH-dependent conformational change, and that the conformational species in the folding landscape exhibit similar free energies. Together, the data show that the landscape for the caspase-hemoglobinase fold is conserved, yet it provides flexibility for species-specific stabilization or destabilization of folding intermediates resulting in changes in stability. The common pH-dependent conformational change in the native dimer, which yields an enzymatically inactive species, may provide an additional, albeit reversible, mechanism for controlling caspase activity in the cell.


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