Furongian (upper Cambrian) trilobites and agnostoids from the Alum Shale Formation of Bornholm, Denmark: revised taxonomy and biostratigraphy

The Furongian (upper Cambrian) trilobite-agnostoid fauna from the Alum Shale Formation of Bornholm, Denmark, is reviewed and revised. The study is based on the museum material stored at the Natural History Museum of Denmark, including the material originally monographed by C. Poulsen (1923) [Bornholms Olenuslag og deres fauna. Danmarks Geologiske Undersøgelse II. Række, Vol. 40, 83 pp]. A total of 8502 specimens, mostly disarticulated sclerites, have been registered. The taxonomy of all species is updated and the best preserved specimens are illustrated. A total of 39 olenid and 5 agnostoid taxa (incl. the Miaolingian Agnostus pisiformis) are recorded including one new species, Ctenopyge magna n. sp. Two specimens of Ctenopyge, treated in open nomenclature as Ctenopyge sp. 1 and sp. 2, may also rep-resent new species. 14 taxa have not been previously reported from Bornholm, viz. Ctenopyge ahlbergi, Ctenopyge tumidoides, Eurycare brevicauda, Leptoplastus abnormis, Leptoplastus crassicornis, Olenus transversus, Parabolina lobata praecurrens, Peltura acu-tidens, Peltura minor, Peltura westergaardi, Protopeltura planicauda, Protopeltura praecursor, Pseudagnostus leptoplastorum? and Sphaerophthalmus drytonensis. Ctenopyge pecten and Ctenopyge affinis are also new to Bornholm as the material formerly described under these names represent Ctenopyge tenuis and C. magna n. sp., respectively. Lotagnostus americanus, Ctenopyge fletcheri, Sphaerophthalmus alatus and Triangulopyge humilis were described under different names by C. Poulsen (1923). Peltura westergaardi and Ctenopyge tenuis are elevated from subspecies to species rank. A redescription of Leptoplastus bornholmensis is presented; the species is transferred to Eurycare. The identification of isolated skeletal parts of L. abnormis and Leptoplastus ovatus and Sphaerophthalmus flagellifer and S. drytonensis are remarked on. The presence of three agnostoid and 14 trilobite zones is confirmed by fossils and all six Furongian superzones are developed on Bornholm. At least the Leptoplastus paucisegmentatus and Leptoplastus raphidophorus zones seem to be absent. Other undocumented zones may be unfossiliferous, not exposed or truly absent. Three different trilobite assemblages (potential subzones) are discerned in the Peltura acu-tidens–Ctenopyge tumida Zone; Ctenopyge tumidoides and Sphaerophthalmus angustus range into the basal part of this zone. All exposures of the Furongian Alum Shale Formation along the Læså and Øleå streams on southern Bornholm are briefly described including GPS coordinates.

Composition and Niche-Specific Characteristics of Microbial Consortia Colonizing Marsberg Copper Mine in the Rhenish Massif

The Kilianstollen Marsberg (Rhenish Massif, Germany) has been extensively mined for copper ores, dating from Early Medieval Period till 1945. The exposed organic-rich alum shale rocks influenced by the diverse mine drainages at an ambient temperature of 10 °C could naturally enrich biogeochemically distinct heavy metal resistant microbiota. This metagenomic study evaluates the microbially colonized subterranean rocks of the abandoned copper mine Kilianstollen to characterize the colonization patterns and biogeochemical pathways of individual microbial groups. Under the selective pressure of the heavy metal contaminated environment at illuminated sites, Chloroflexi (Ktedonobacteria) and Cyanobacteria (Oxyphotobacteria) build up whitish-greenish biofilms. In contrast, Proteobacteria, Firmicutes and Actinobacteria dominate rocks around the uncontaminated spring water streams. The metagenomic analysis revealed that the heavy metal resistant microbiome was evidently involved in redox cycling of transition metals (Cu, Zn, Co, Ni, Mn, Fe, Cd, Hg). No deposition of metals or minerals, though, was observed by transmission electron microscopy in Ktedonobacteria biofilms which may be indicative for the presence of different detoxification pathways. The underlying heavy metal resistance mechanisms, as revealed by analysis of metagenome-assembled genomes, were mainly attributed to transition metal efflux pumps, redox enzymes, volatilization of Hg0, methylated intermediates of As(III) and reactive oxygen species detoxification pathways.

Review of Devonian-Carboniferous Boundary sections in the Rhenish Slate Mountains (Germany)

Thirty Devonian-Carboniferous Boundary sections of the Rhenish Slate Mountains and adjacent subsurface areas are reviewed with respect to litho-, event, conodont, ammonoid, sequence, and chemostratigraphy. In the interval from the base of the uppermost Famennian (Wocklum Beds, Wocklumian) to the base of the middle Tournaisian (base Lower Alum Shale), 11 conodont and 16 ammonoid (sub)zones are distinguished. The terminology of the Hangenberg Crisis Interval is refined, with an overall regressive Crisis Prelude below the main Hangenberg Extinction, which defines the base of the transgressive Lower Crisis Interval (Hangenberg Black Shale). The glacigenic and regressive Middle Crisis Interval (Hangenberg Shale/Sandstone) is followed by the overall transgressive Upper Crisis Interval that can be subdivided into three parts (I to III) with the help of conodont stratigraphy (upper costatus-kockeli Interregnum = upper ckI, Protognathodus kockeli Zone, and lower part of Siphonodella (Eosiphonodella) sulcata s.l./Pr. kuehni Zone). Protognathodus kockeli includes currently a wide range of forms, which variabilities and precise ranges need to be established before a precise GSSP level should be selected. Returning to its original definition, the former Upper duplicata Zone is re-named as Siphonodella (S.) mehli Zone. It replaces the S. (S.) jii Zone, which is hampered by taxonomic complications. The S. (S.) quadruplicata Zone of Ji (1985) is hardly supported by Rhenish data. The entry of typical S. (S.) lobata (M1) characterises an upper subdivision (subzone) of the S. (S.) sandbergi Zone; the new S. (S.) lobata M2 enters much earlier within the S. (S.) mehli Zone. The ammonoid-defined base of the Wocklum-Stufe (Upper Devonian = UD VI) begins with the Linguaclymenia similis Zone (UD VI-A1). The oldest S. (Eosiphonodella) enter within the Muessenbiaergia bisulcata Zone (UD VI-A2). The traditional Parawocklumeria paradoxa Zone of Schindewolf (1937) is divided into successive P. paprothae (VI-C1), P. paradoxa (VI-C2), and Mayneoceras nucleus (VI-C3) Subzones. In the lower Tournaisian (Lower Carboniferous = LC I), the Gattendorfia subinvoluta Zone is subdivided into G. subinvoluta (LC I-A2) and “Eocanites” nodosus (LC I-A3) Subzones. The Paprothites dorsoplanus Zone (LC I-B) can be divided into Pap. dorsoplanus (LC I-B1) and Paragattendorfia sphaeroides (LC I-B2) Subzones. Potential subdivisions of the Pseudarietites westfalicus (LC I-C) and Parag. patens Zones (LC I-D) are less distinctive. The unfossiliferous or argillaceous upper part of the Hangenberg Limestone and the overlying Lower Alum Shale Event Interval remain regionally unzoned for ammonoids.

Reconstructing the Ediacaran to Ordovician history of the Baltoscandian Margin of continent Baltica

<p>In the Scandes, the lower thrust sheets of the Caledonian allochthons provide unambiguous stratigraphic evidence of correlation with the successions of the Baltoscandian platform. Cambrian successions, including the Alum Shale Formation, providing the footwall for the main Caledonian decollement in Scandinavia, can be followed at least 200 km westwards from the thrust front into the hinterland of the orogen. The overlying early Palaeozoic strata provide evidence of facies changes into foreland basin deposits in the mid Ordovician and early Silurian; also of Ediacaran and Cryogenian successions, including Marinoan tillites. The amount of internal shortening in the Lower Allochthon is not uncontroversial, but certainly amounts to more than 100 km, implying that all the overlying alllochthons in the Scandes were derived from west of the Norwegian coast.</p><p>The metamorphic grade of the units in the Lower Allochthon increases from low to high greenschist facies, from the thrust front westwards into the deep hinterland. Overlying thrust sheets of the Middle Allochthon are of higher metamorphic grade and more ductilely deformed. The basal parts are usually dominated by basement-derived units and Neoproterozoic sedimentary rocks. They are overthrust by dolerite dyke-intruded thrust sheets, the Särv Nappes, with host-rocks dominated by Cryogenian and Ediacaran sandstones, the former including subordinate limestones and Marinoan tillites. The Baltoscandian margin dolerite dyke swarms amount to up to c. 35% of these thrust sheets.</p><p>The overlying, highest tectonic units in the Middle Allochthon (the Seve Nappe Complex, SNC) are of amphibolite and higher metamorphic grade. They include a greater variety of lithologies, including some that are very similar to those in the underlying Särv Nappes (e,g. quartzites and eclogitized dolerites). The metasedimentay host rocks include a wide range of paragneisses and marbles. Abundant mafic rocks include metamorphosed gabbros, basalts and peridotites and, together with the dyke swarms, can totally dominate the composition of some thrust sheets. The similar geochemistry and early Ediacaran age (c. 600 Ma) of the mafic rocks in the Särv and Seve nappes define the Baltoscandian outermost margin and continent-ocean transition zone (COT). Iapetus Ocean terranes comprise the overlying thrust sheets of the Upper Allochthon (e.g. the Köli Nappe Complex).</p><p>The metamorphism of the different thrust sheets in the SNC provide clear evidence that some parts were subducted; others not. A wide range of isotope age data constrain the timing of subduction, with the earliest ages in the mid Cambrian (c. 505 Ma) to early Ordovician (c. 483 Ma). It has been suggested that the deposition of the Alum Shale Formation on the Baltscandian platform, was related to this early Caledonian subduction. A more probable interpretation is that subduction along the outermost edge of this highly extended COT did not influence the edge of the platform till the early Tremadoc.</p><p>Some authors have introduced cryptic sutures into the Baltoscandian outer margin, described above. They should reassess their data and better define the evidence for their conviction.</p>

Collisional Orogeny in the Scandinavian Caledonides (COSC): Some preliminary results from drilling of the 2.276 km deep COSC-2 borehole, central Sweden

<p>COSC investigations and drilling activities are focused in the Åre-Mörsil area (Sweden) of central Scandinavia. COSC-2 was drilled with nearly 100% core recovery in 2020 to 2.276 km depth with drilling ongoing from mid-April to early August. Drilling targets for COSC-2 included (1) the highly conductive Alum shale, (2) the Caledonian décollement, the major detachment that separates the Caledonian allochthons from the autochthonous basement of the Fennoscandian Shield, and (3) the strong seismic reflectors in the Precambrian basement.</p><p>Combined seismic, magnetotelluric (MT) and magnetic data were used to site the COSC-2 borehole about 20 km east-southeast of COSC-1. Based on these data it was predicted that the uppermost, tectonic occurrence of Cambrian Alum shale would be penetrated at about 800 m, the main décollement in Alum shale at its stratigraphic level at about 1200 m and the uppermost high amplitude basement reflector at about 1600 m. Paleozoic turbidites and greywackes were expected to be drilled down to 800 m depth. Below this depth, Ordovician limestone and shale with imbricates of Alum shale were interpreted to be present. Directly below the main décollement, magnetite rich Precambrian basement was expected to be encountered with a composition similar to that of magnetic granitic rocks found east of the Caledonian Front. The actual depths of the main contacts turned out to agree very well with the predictions based on the geophysical data. However, the geology below the uppermost occurrence of Alum shale is quite different from the expected model. Alum shale was only clearly encountered as a highly deformed, about 30 m thick unit, starting at about 790 m. Between about 820 and 1200 m, preliminary interpretations are that the rocks mainly consist of Neo-Proterozoic to Early Cambrian tuffs. Further below, Precambrian porphyries are present. The high amplitude reflections within the Precambrian sequence appear to be generated by dolerite sheets with the uppermost top penetrated at about 1600 m. Several deformed sheets of dolerite may be present down to about 1930 m. Below this depth the rocks are again porphyries.</p><p>A preliminary conclusion concerning the tectonic model is that the main décollement is at about 800 m and not at 1200 m. Also the thickness of the lowermost Cambrian/uppermost Neoproterozoic sediments on top of the basement is much greater than expected (hundreds of meters instead of tens of meters) and likely to have been thickened tectonically. Detailed studies are required to assess the actual importance of the “main décollement” and the degree, type and age of deformation in its footwall. We can also conclude that the Precambrian basement is very similar to the Dala porphyries succession that are typically present farther south.</p><p>An extensive set of downhole logging data was acquired directly after drilling. Borehole seismic measurements in 2021 will help to define and correlate seismic boundaries with lithology and structures in the core. Unfortunately, work for describing the geology of the drill core in detail is still on hold due to Covid-19.</p>