scholarly journals Cooperation-based concept formation in male bottlenose dolphins

2021 ◽  
Vol 12 (1) ◽  
Author(s):  
Stephanie L. King ◽  
Richard C. Connor ◽  
Michael Krützen ◽  
Simon J. Allen

AbstractIn Shark Bay, Western Australia, male bottlenose dolphins form a complex nested alliance hierarchy. At the first level, pairs or trios of unrelated males cooperate to herd individual females. Multiple first-order alliances cooperate in teams (second-order alliances) in the pursuit and defence of females, and multiple teams also work together (third-order alliances). Yet it remains unknown how dolphins classify these nested alliance relationships. We use 30 years of behavioural data combined with 40 contemporary sound playback experiments to 14 allied males, recording responses with drone-mounted video and a hydrophone array. We show that males form a first-person social concept of cooperative team membership at the second-order alliance level, independently of first-order alliance history and current relationship strength across all three alliance levels. Such associative concepts develop through experience and likely played an important role in the cooperative behaviour of early humans. These results provide evidence that cooperation-based concepts are not unique to humans, occurring in other animal societies with extensive cooperation between non-kin.

2010 ◽  
Vol 7 (4) ◽  
pp. 623-626 ◽  
Author(s):  
Richard C. Connor ◽  
Jana J. Watson-Capps ◽  
William B. Sherwin ◽  
Michael Krützen

Male bottlenose dolphins in Shark Bay, Western Australia form two levels of alliances; two to three males cooperate to herd individual females and teams of greater than three males compete with other groups for females. Previous observation suggested two alliance tactics: small four to six member teams of relatives that formed stable pairs or trios and unrelated males in a large 14-member second-order alliance that had labile trio formation. Here, we present evidence for a third level of alliance formation, a continuum of second-order alliance sizes and no relationship between first-order alliance stability and second-order alliance size. These findings challenge the ‘two alliance tactics’ hypothesis and add to the evidence that Shark Bay male bottlenose dolphins engage in alliance formation that likely places considerable demands on their social cognition.


2009 ◽  
Vol 74 (1) ◽  
pp. 43-55 ◽  
Author(s):  
Dennis N. Kevill ◽  
Byoung-Chun Park ◽  
Jin Burm Kyong

The kinetics of nucleophilic substitution reactions of 1-(phenoxycarbonyl)pyridinium ions, prepared with the essentially non-nucleophilic/non-basic fluoroborate as the counterion, have been studied using up to 1.60 M methanol in acetonitrile as solvent and under solvolytic conditions in 2,2,2-trifluoroethan-1-ol (TFE) and its mixtures with water. Under the non- solvolytic conditions, the parent and three pyridine-ring-substituted derivatives were studied. Both second-order (first-order in methanol) and third-order (second-order in methanol) kinetic contributions were observed. In the solvolysis studies, since solvent ionizing power values were almost constant over the range of aqueous TFE studied, a Grunwald–Winstein equation treatment of the specific rates of solvolysis for the parent and the 4-methoxy derivative could be carried out in terms of variations in solvent nucleophilicity, and an appreciable sensitivity to changes in solvent nucleophilicity was found.


1993 ◽  
Vol 71 (4) ◽  
pp. 523-532 ◽  
Author(s):  
Joanna Fraser ◽  
Eric G. Kokko

The initial stages of panicle, spikelet, and floret development in field-grown 'Kay' orchardgrass were examined using scanning electron microscopy. Spikelets arose from a complex multilevelled sequence of initiation from branch apices. Spikelets developed indirectly in a two-tiered progression: (i) an acropetal and basipetal sequence of first order, second-order, and third-order inflorescence apices, and (ii) an acropetal development within subclusters of higher-order lateral branch inflorescence apices. The panicle had the unique feature of dorsiventrality as well as bilateral symmetry. The basal apex from first-order, second-order, or third-order apices developed on the same side of the main axis as the first-order apex. The two glumes subtending each spikelet primordium developed alternately and acropetally. Development and initiation of florets within spikelets was basipetal within the panicle, basipetal within clusters and subclusters of spikelets on lateral branches, and acropetal within spikelets. Within florets, paleas developed later than lemmas. Key words: Dactylis glomerata, cocksfoot, scanning electron microscopy, development, panicle.


1968 ◽  
Vol 11 (1) ◽  
pp. 85-93 ◽  
Author(s):  
Z. A. Melzak

In graph - theoretic terms a homogeneous p-dendrite, p ≥ 2, is defined as a finite singly-rooted tree in which the root has valency 1 while every other vertex has valency 1 or p. More descriptively, a homogeneous p-dendrite may be imagined to start from its root as the main, or 0th order, branch which proceeds to the first - order branch point where it gives rise top first - order branches. Each of these either terminates at its other end (which is a second-order branch point) or it splits there again into p branches (which are of third order), and so on. The order of the dendrite is the highest order of a branch present in it. For completeness, a 0-th order dendrite is also allowed, this consists of the 0-th order branch alone.


1993 ◽  
Vol 265 (1) ◽  
pp. H146-H151 ◽  
Author(s):  
R. L. Hester ◽  
A. Eraslan ◽  
Y. Saito

This study was designed to determine the physiological role of endothelium-dependent nitric oxide (EDNO) in the control of arteriolar diameter during rest and muscle stimulation. Diameters of first-, second-, and third-order arterioles in the superfused hamster cremaster muscle were measured before and throughout 1 min of field stimulation before and after inhibition of EDNO release. ENDO inhibition by intravenous N omega-nitro-L-arginine methyl ester (L-NAME) significantly attenuated the arteriolar vasodilation in response to 1 microM acetylcholine. First-order arterioles averaged 65 +/- 5 microns at rest and dilated to 86 +/- 6 microns during muscle stimulation (n = 9), second-order arterioles averaged 45 +/- 6 microns and dilated to 72 +/- 3 microns during muscle stimulation (n = 6), with third-order arterioles averaging 29 +/- 2 microns, and dilating to 53 +/- 3 microns during muscle stimulation (n = 7). EDNO inhibition significantly decreased both the resting diameter of first-order arterioles (57 +/- 4 microns) and functional dilation (68 +/- 3 microns; P <0.05). EDNO inhibition had no effect on the resting diameter of second-order arterioles (45 +/- 5 microns) yet significantly attenuated the functional dilation (64 +/- 4 microns; P < 0.05). EDNO inhibition had no effect on either the resting diameter of third-order arterioles (30 +/- 2 microns) or the functional dilation (49 +/- 2 microns).(ABSTRACT TRUNCATED AT 250 WORDS)


Terminology ◽  
2014 ◽  
Vol 20 (2) ◽  
pp. 279-303 ◽  
Author(s):  
Ann Bertels ◽  
Dirk Speelman

This paper presents an innovative approach, within the framework of distributional semantics, for the exploration of semantic similarity in a technical corpus. In complement to a previous quantitative semantic analysis conducted in the same domain of machining terminology, this paper sets out to discover fine-grained semantic distinctions in an attempt to explore the semantic heterogeneity of a number of technical items. Multidimensional scaling analysis (MDS) was carried out in order to cluster first-order co-occurrences of a technical node with respect to shared second-order and third-order co-occurrences. By taking into account the association values between relevant first and second-order co-occurrences, semantic similarities and dissimilarities between first-order co-occurrences could be determined, as well as proximities and distances on a graph. In our discussion of the methodology and results of statistical clustering techniques for semantic purposes, we pay special attention to the linguistic and terminological interpretation.


1998 ◽  
Vol 120 (1) ◽  
pp. 20-29 ◽  
Author(s):  
J. R. Krokstad ◽  
C. T. Stansberg ◽  
A. Nestega˚rd ◽  
T. Marthinsen

New results from the most recent work within the Norwegian Joint Industry Project (JIP) “Higher Order Wave Load Effects on Large Volume Structures” are presented. A nonslender theoretical model is validated from experiments for two fixed, vertical cylinders with different diameter/peak wavelength ratios. A combination of complete diffraction first-order simulations, sum and difference frequency second-order simulations, and third-order FNV (Faltinsen, Newman, and Vinje, nonlinear long wave model) is implemented in order to develop a simplified and robust ringing load model for a large range of cylinder diameter/peak wavelength ratios. Results from the full diffraction second-order analysis show a significant reduction of second-order loads compared to pure FNV in the wavelength range relevant for ringing loads. The results show improved correspondence with high-frequency experimental loads compared with the unmodified FNV. Results for different cylinder peak wavelength ratios are presented, including validation against experiments. In addition, a few simplified response simulations are carried out demonstrating significant improvements with the modified FNV model.


1969 ◽  
Vol 47 (7) ◽  
pp. 699-705 ◽  
Author(s):  
C. S. Sharma ◽  
R. G. Wilson

The first-order Hartree–Fock and unrestricted Hartree–Fock equations for the ground state of a five electron atomic system are solved exactly. The solutions are used to evaluate the corresponding second-order energies exactly and the third-order energies with great accuracy. The first-order terms in the expectation values of 1/r, r, r2, and δ(r) are also calculated.


2020 ◽  
Vol 2 (2) ◽  
pp. 90-93
Author(s):  
Luvera Deva Intan Indrawati ◽  
Rina Dwi Indriana ◽  
Irham Nurwidyanto

Geophysics programing of regional and residual anomaly separation on Magnetic data has been carried out with the results compared with the upward continuation method in the OasisMontaj software. Separation of anomalies with moving average and polynomial methods is processed using Matlab programming. The orders used in the polynomial method are first-order, second-order and third-order. Comparison is done by calculating the match value. The chosen matching method is autocorrelation. Correlation of residual magnetic anomalies resulting from upward continuation (Magpick) to moving averages, 1st-order polynomials, 2nd-order polynomials and 3rd-order polynomials. Correlation values obtained for the moving average method are 0.9604, first order polynomial 0.9072, 2nd order polynomial 0.9482 and third order polynomial 0.6057. The moving average and second order polynomial methods can be used as a substitute method if we do not use the upward continuation method.


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