Systematics of the spider genus Neoleptoneta Brignoli, 1972 (Araneae:Leptonetidae) with a discussion of the morphology and relationships for the North American Leptonetidae

2011 ◽  
Vol 25 (4) ◽  
pp. 334 ◽  
Author(s):  
Joel Ledford ◽  
Pierre Paquin ◽  
James Cokendolpher ◽  
Josh Campbell ◽  
Charles Griswold

A phylogenetic analysis of the spider genus Neoleptoneta Brignoli, 1972 is presented based on molecular sequence variation from three genes (mitochondrial cytochrome c oxidase subunit I, nuclear histone H3 and nuclear 28S rDNA) and including exemplars for all North American leptonetid genera except the ecribellate archoleptonetine Darkoneta. Analysis of concatenated data and independent genes using Bayesian, maximum likelihood and parsimony methods failed to recover Neoleptoneta as monophyletic. The genera Archoleptoneta, Appaleptoneta and Calileptoneta are monophyletic and a sister group relationship is supported between Appaleptoneta and Calileptoneta. Morphological data based on a survey of leptonetid genera using scanning electron and compound light microscopy are discussed and traced on the molecular phylogeny. Images for each North American leptonetine genus are provided, including comparison with Asian and European outgroups. Images of the incertae sedis species Leptoneta brunnea Gertsch, 1974 and Leptoneta sandra Gertsch, 1974 are provided and their generic placement is re-evaluated. Ancestral state reconstruction is used to assess patterns of cave evolution and shows that most species are descended from troglophilic ancestors and that troglobites have evolved at least nine times independently within the North American Leptonetidae. Neoleptoneta is relimited to include seven species restricted to central Mexico including N. bonita (Gertsch, 1974), N. capilla (Gertsch, 1971), N. delicata (Gertsch, 1971), N. limpida (Gertsch, 1974), N. rainesi (Gertsch, 1971) and N. reclusa (Gertsch, 1971) and to include Leptoneta brunnea, giving the new combination N. brunnea (Gertsch, 1974). The remaining species described in Neoleptoneta are placed in three new genera: (1) Chisoneta, gen. nov. from south-western Texas and Nuevo Leon, Mexico, including the four species C. chisosea (Gertsch, 1974), C. isolata (Gertsch, 1971), C. modica (Gertsch, 1974) and C. pecki (Gertsch, 1971), new combinations; (2) Ozarkia, gen. nov. from Arizona and New Mexico north-east to Arkansas, Alabama and Georgia, including the nine species O. alabama (Gertsch, 1974), O. apachea (Gertsch, 1974), O. archeri (Gertsch, 1974), O. arkansa (Gertsch, 1974), O. blanda (Gertsch, 1974), O. georgia (Gertsch, 1974), O. ivei (Gertsch, 1974), O. novaegalleciae (Brignoli, 1979) and O. serena (Gertsch, 1974), new combinations; and (3) Tayshaneta, gen. nov. from Texas south to Coahuila, Mexico, with the eleven species T. anopica (Gertsch, 1974), T. bullis (Cokendolpher, 2004), T. coeca (Chamberlin & Ivie, 1942), T. concinna (Gertsch, 1974), T. devia (Gertsch, 1974), T. furtiva (Gertsch, 1974), T. microps (Gertsch, 1974), T. myopica (Gertsch, 1974), T. paraconcinna (Cokendolpher & Reddell, 2001), T. uvaldea (Gertsch, 1974) and T. valverdae (Gertsch, 1974), new combinations. Leptoneta sandra Gertsch, 1974 cannot be placed in any North American, European or Asian genus and is thus transferred to the new genus Montanineta, gen. nov., giving the new combination Montanineta sandra (Gertsch, 1974).

2019 ◽  
Vol 44 (4) ◽  
pp. 930-942
Author(s):  
Geraldine A. Allen ◽  
Luc Brouillet ◽  
John C. Semple ◽  
Heidi J. Guest ◽  
Robert Underhill

Abstract—Doellingeria and Eucephalus form the earliest-diverging clade of the North American Astereae lineage. Phylogenetic analyses of both nuclear and plastid sequence data show that the Doellingeria-Eucephalus clade consists of two main subclades that differ from current circumscriptions of the two genera. Doellingeria is the sister group to E. elegans, and the Doellingeria + E. elegans subclade in turn is sister to the subclade containing all remaining species of Eucephalus. In the plastid phylogeny, the two subclades are deeply divergent, a pattern that is consistent with an ancient hybridization event involving ancestral species of the Doellingeria-Eucephalus clade and an ancestral taxon of a related North American or South American group. Divergence of the two Doellingeria-Eucephalus subclades may have occurred in association with northward migration from South American ancestors. We combine these two genera under the older of the two names, Doellingeria, and propose 12 new combinations (10 species and two varieties) for all species of Eucephalus.


Zootaxa ◽  
2007 ◽  
Vol 1501 (1) ◽  
pp. 31-44 ◽  
Author(s):  
FALITIANA C.E. RABEMANANJARA ◽  
ANGELICA CROTTINI ◽  
YLENIA CHIARI ◽  
FRANCO ANDREONE ◽  
FRANK GLAW ◽  
...  

Malagasy poison frogs of the genus Mantella with its 16 species are currently sub-divided into 5 major groups. Of these, the Mantella betsileo group is traditionally understood as containing four species, Mantella betsileo, M. expectata, M. viridis and M. manery, while the M. laevigata group is considered to be monospecific. A phylogenetic analysis of samples from multiple localities of all species in these two groups, based on sequences of the mitochondrial cytochrome b gene, shows the existence of several well-distinct clades in what is currently considered to be Mantella betsileo: (1) central-western populations from Kirindy, Isalo, and near Antsirabe close to the Betsileo region, to which the name M. betsileo is to be applied, (2) populations of the north-east and north-west, which are closely related to M. viridis and to which the name M. ebenaui is to be applied, and (3) a clade from southernmost Madagascar and from the Tsingy de Bemaraha, which is sister to M. expectata and furthermore includes important intra-clade variation, therefore probably representing one or two undescribed species. Our data also support a large genetic distance of M. manery to all other species and its probable sister-group relationship to the sympatric M. laevigata; M. manery is consequently transferred from the M. betsileo group to the M. laevigata group.


ZooKeys ◽  
2018 ◽  
Vol 789 ◽  
pp. 51-101 ◽  
Author(s):  
Bernhard A. Huber ◽  
Jonas Eberle ◽  
Dimitar Dimitrov

With almost 600 species, the latest molecular phylogeny of pholcid spiders (Eberle et al. 2018, BMC Evolutionary Biology) more than triples the largest previously available molecular phylogeny of the family. At the level of genera, the coverage is high (86%, i.e., 75 of the 87 named genera), and at the level of subfamilies it is complete. The present paper is an effort to critically evaluate the implications of this phylogeny for pholcid systematics. The analyses largely support the division of Pholcidae into five subfamilies: Ninetinae, Arteminae, Modisiminae, Smeringopinae, and Pholcinae. Their compositions are largely unchanged except thatChisosaHuber, 2000 is moved from Ninetinae to Arteminae. The positions ofArtemaWalckenaer, 1837 andPrisculaSimon, 1893 in this system remain dubious. Relationships among subfamilies remain weakly supported, except for the sister group relationship between Smeringopinae and Pholcinae. Several major clades within subfamilies are separated from each other along geographical boundaries; for example within Modisiminae a South American clade and a Central + North American + Caribbean clade, and within Smeringopinae a Sub-Saharan clade and a clade ranging from the Mediterranean to Central Asia. Central + North American + Caribbean clades in both Ninetinae and Modisiminae may originate from South American ancestors.Many taxonomic changes are suggested by the data, some of which are formally implemented herein. Two new genera result from the splitting ofCalapnitaSimon, 1892 andPanjangeDeeleman-Reinhold & Deeleman, 1983, respectively:NipisaHuber,gen. n.; andApokayanaHuber,gen. n.Nine new genera result from splitting ofPholcus:CantikusHuber,gen. n.;KelabitaHuber,gen. n.;KintaqaHuber,gen. n.;MurutaHuber,gen. n.;MerahaHuber,gen. n.;PaiwanaHuber,gen. n.;PribumiaHuber,gen. n.;TerangaHuber,gen. n.; andTissahamiaHuber,gen. n.Two genera are newly synonymized:PlatnickniaÖzdikmen & Demir, 2009 is synonymized withModisimusSimon, 1893;SihalaHuber, 2011 is synonymized withPholcusWalckenaer, 1805.PholcusagadirHuber, 2011 is moved toMicropholcusDeeleman-Reinhold & Prinsen, 1987, resulting in the new combinationMicropholcusagadir(Huber, 2011).


1964 ◽  
Vol 42 (7) ◽  
pp. 859-884 ◽  
Author(s):  
Tetsuo Koyama ◽  
Shoichi Kawano

Taxonomy and distribution of grasses with the North American and eastern Asiatic distribution have been discussed in detail with brief comments on their history. Morphological comparison of the corresponding taxa was correlated with cytological data, and the distribution was discussed with particular emphasis on the related ecological evidence. The new combinations proposed are: Schizachne purpurascens ssp. callosa, Brachyelytrum erectum ssp. erectum var. glabratum, B. erectum ssp. japonicum, Muhlenbergia frondosa ssp. ramosa, M. tenuiflora ssp. curviaristata, Glyceria acutiflora ssp. japonica, Festuca subulata ssp. japonica, Torreyochloa pallida ssp. pallida var. Fernaldii, T. pallida ssp. natans, T. pallida ssp. natans var. viridis, and Beckmannia Syzigachne ssp. baicalensis.


1993 ◽  
Vol 125 (5) ◽  
pp. 847-867 ◽  
Author(s):  
Yves Alarie

AbstractNorth American members of the Oreodyies alaskanus clade are revised. The species O. productotruncatus (Hatch) and O. recticollis (Fall) are recognized as valid and those names are removed from junior synonymy with O. alaskanus (Fall.). Oreodytes leechi Zimmerman is considered a new junior subjective synonym of O. recticollis. Lectotype designations are provided for O. alaskanus and O. recticollis. Palaearctic O. dauricus (Motschulsky) is included within the O. alaskanus clade whose members are characterized by the protibia having the inner margin sinuate and strongly narrowed proximally, Oreodytes kanoi Kamiya, from Japan, is suggested as the sister-group of members of the O. alaskanus clade based on the shared presence in the female of a last abdominal sternite with an emargination at the apex.


1963 ◽  
Vol 41 (9) ◽  
pp. 1331-1346 ◽  
Author(s):  
Aino Henssen

The systematic position of the genus Massalongia and the closely related genera Koerberia, Vestergrenopsis, and Placynthium in the family Peltigeraceae including lichens with hemiangiocarpic apothecia is discussed. The ontogeny of a hemiangiocarpic apothecium is described briefly. A key for the determination of the genera is provided.A general survey is given for the morphology and anatomy of the genus Massalongia. The two species, M. carnosa and M. microphylliza, are described in detail. The new combination M. microphylliza is made.


2004 ◽  
Vol 73 (1-2) ◽  
pp. 3-163 ◽  
Author(s):  
Ronald A. Jenner

This paper critically assesses all morphological cladistic analyses of the Metazoa that were published during the last one and a half decades. Molecular and total evidence analyses are also critically reviewed. This study focuses on evaluating alternative phylogenetic positions of the ‘acoelomate’ worms: Platyhelminthes, Nemertea, and Gnathostomulida. This paper consists of two parts. In Part I, all recently proposed sister group hypotheses and the supporting synapomorphies for these phyla are evaluated. Discrepancies in the treatment of corresponding characters in different cladistic analyses are identified, and where possible, resolved. In Part II, the overall phylogenetic significance across the Metazoa of all characters relevant for placing the ‘acoelomate’ worms is examined. The coding and scoring of these characters for other phyla are evaluated, and uncertainties in our understanding are pointed out in order to guide future research. The characters discussed in this paper are broadly categorized as follows: epidermis and cuticle, reproduction and sexual condition, development, larval forms, coeloms and mesoderm source, nervous system and sensory organs, nephridia, musculature, digestive system, and miscellaneous characters. Competing phylogenetic hypotheses are compared in terms of several criteria: 1) taxon sampling and the fulfillment of domain of definition for each character; 2) character sampling; 3) character coding; 4) character scoring and quality of primary homology; 5) quality of the proposed diagnostic synapomorphies as secondary homologies. On the basis of this study I conclude that a sister group for the Platyhelminthes has not yet been unambiguously established. A clade minimally composed of Neotrochozoa (Mollusca, Sipuncula, Echiura, Annelida) emerges as the most likely sister group of the Nemertea on the basis of morphological and total evidence analyses. Finally, morphological data currrently favor a sister group relationship of Gnathostomulida and Syndermata (probably plus Micrognathozoa). In contrast, molecular or total evidence analyses have not identified a reliable sister group of Gnathostomulida.Further progress in our understanding of metazoan phylogeny crucially depends on the improvement of the quality of currently adopted cladistic data matrices. A thorough reassessment of many of the more than 70 morphological characters discussed here is necessary. Despite the recent compilation of comprehensive data matrices, the power to test competing hypotheses of higher-level metazoan relationships is critically compromised due to uncritical data selection and poor character study in even the most recently published cladistic analyses.


Zootaxa ◽  
2019 ◽  
Vol 4683 (3) ◽  
pp. 301-335 ◽  
Author(s):  
FRANK E. ETZLER

The genus Limonius Eschscholtz, 1829 was last treated as a whole by Candèze (1860). Since then, members have been placed in eight other genera: Cidnopus Thomson, 1859; Gambrinus LeConte, 1853; Elathous Reitter, 1890; Kibunea Kishii, 1966; Limoniscus Reitter, 1905; Nothodes LeConte, 1861; Pheletes Kiesenwetter, 1858; and Solskyana Dolin, 1978. Based on the examination of adult and larval characters, five genera are recognized: Elathous Reitter, 1890; Gambrinus LeConte, 1853; Limonius Eschscholtz, 1829; Pheletes Kiesenwetter, 1858; and Tetralimonius new genus. Limoniscus Reitter, 1905 and Sichuanelater Platia and Gudenzi, 2006 are new synonymies of Gambrinus LeConte, 1853; Micrathous Lane, 1971, Neoathousius Schimmel and Platia, 1991 and Solskyana Dolin, 1978 are all new synonymies of Limonius. A total of 84 new combinations are proposed: Nearctic: Elathous huguenini (Van Dyke, 1932) new combination; Gambrinus angulatus (Motschulsky, 1859) new combination; Gambrinus bicolor (Van Dyke, 1932) new combination; Gambrinus clypeatus (Motschulsky, 1859) new combination; Gambrinus confusus (LeConte, 1853) new combination; Gambrinus cribriceps (Van Dyke, 1943) new combination; Gambrinus crotchii (Horn, 1872) new combination; Gambrinus flavomarginatus (Knull, 1938) new combination; Gambrinus fulvipilis (Candèze, 1860) new combination; Gambrinus griseus (Beauvois, 1805) new combination; Gambrinus humidus (Lane, 1941) new combination; Gambrinus interstitialis (Melsheimer, 1846) new combination; Gambrinus lanchesteri (Lane, 1941) new combination; Gambrinus meridianus (Knull, 1947) new combination; Gambrinus mirus (LeConte, 1853) new combination; Gambrinus norahae (Al Dhafer, 2009) new combination; Gambrinus olentangyi (Knull, 1947) new combination; Gambrinus plebejus (Say, 1825) new combination; Gambrinus propexus (Candèze, 1860) new combination; Gambrinus rudis (Brown, 1933) new combination; Gambrinus rufihumeralis (Lane, 1941) new combination; Gambrinus seminudus (Van Dyke, 1932) new combination; Gambrinus shircki (Lane, 1965) new combination; Gambrinus sinuifrons (Fall, 1907) new combination; Gambrinus snakensis (Lane, 1965) new combination; Gambrinus stigma (Herbst, 1806) new combination; Gambrinus pictus (Van Dyke, 1932) new combination; Gambrinus ulkei (Horn, 1871) new combination; Gambrinus ursinus (Van Dyke, 1932) new combination; Gambrinus venablesi (Wickham, 1913) new combination; Limonius brevis (Van Dyke, 1932) new combination; Limonius sordidus (Van Dyke, 1932) new combination; Pheletes lecontei (Lane, 1971) new combination; Tetralimonius definitus (Ziegler, 1845) new combination; Tetralimonius humeralis (Candèze, 1860) new combination; Tetralimonius maculicollis (Motschulsky, 1860) new combination; Tetralimonius nimbatus (Say, 1825) new combination; Tetralimonius ornatulus (LeConte, 1857) new combination. Palearctic: Gambrinus elegans (Buysson, 1891) new combination; Gambrinus gibbosus (Platia and Gudenzi, 2006) new combination. Gambrinus henanensis (Schimmel, 2006) new combination; Gambrinus hinakurai (Kishii, 1998) new combination; Gambrinus katoi (Kishii, 2002) new combination; Gambrinus kawaharai (Kishii, 2002) new combination; Gambrinus kucerai (Schimmel, 2006) new combination; Gambrinus nanshanensis (Arimoto and Hiramatsu, 2013) new combination; Gambrinus naomii (Kishii, 1997) new combination; Gambrinus shaanxiensis (Schimmel, 2006) new combination; Gambrinus suturalis (Gebler, 1844) new combination; Gambrinus takabai (Kishii, 1997) new combination; Gambrinus violaceus (Müller, 1821) new combination; Gambrinus wittmeri (Chassain, 1998) new combination; Gambrinus yamato (Kishii, 1998) new combination; Gambrinus yujii (Arimoto, 2013) new combination; Gambrinus zhejiangensis (Schimmel, 2015) new combination; Limonius brancuccii (Schimmel and Platia, 1991) new combination; Limonius decorus (Gurjeva, 1975) new combination; Limonius exiguus (Schimmel and Platia, 1991) new combination; Limonius hartmanni (Schimmel, 1998) new combination; Limonius hiermeieri (Schimmel and Platia, 1991) new combination; Limonius hirtus (Dolin, 1978) new combination; Limonius hubeiensis (Kishii and Jiang, 1996) new combination; Limonius kubani (Schimmel, 1996) new combination; Limonius loebli (Schimmel and Platia, 1991) new combination; Limonius longicornis (Schimmel and Platia, 1991) new combination; Limonius macedonicus (Cate and Platia, 1989) new combination; Limonius marginellus brusteli (Leseigneur, 2004) new combination; Limonius manaliensis (Schimmel and Platia, 1991) new combination; Limonius miandamensis (Schimmel and Platia, 1991) new combination; Limonius minusculus (Schimmel and Platia, 1991) new combination; Limonius nigronitidus (Han and Lee, 2012) new combination; Limonius platiai (Mertlik, 1996) new combination; Limonius pseudopilosus (Platia and Gudenzi 1985) new combination; Limonius recticornis (Schimmel and Platia, 1991) new combination; Limonius riesei (Platia, 1988) new combination; Limonius rusticus (Schimmel and Platia, 1991) new combination; Limonius schurmanni (Platia and Gudenzi, 1998) new combination; Limonius sinensis (Schimmel and Platia, 1994) new combination; Limonius singularis (Schimmeland Platia, 1991) new combination; Limonius stapfi (Schimmel, 2007) new combination; Limonius turcicus (Platia, 2004) new combination; Limonius wittmeri (Schimmel and Platia, 1991) new combination; Tetralimonius quercus (Olivier, 1790) new combination; Tetralimonius reitteri (Gurjeva, 1976) new combination. The following 12 North American species are removed from synonymy and recognized as valid species: Gambrinus interstitialis (Melsheimer, 1846) status resurrected; Gambrinus propexus (Candèze, 1860) status resurrected; Gambrinus shircki (Lane, 1965) status resurrected; Gambrinus snakensis (Lane, 1965) status resurrected; Gambrinus ulkei (Horn, 1871) status resurrected; Limonius anceps LeConte, 1853 status resurrected; Limonius dubitans LeConte, 1853 status resurrected; Limonius infuscatus Motschulsky, 1859 status resurrected; Limonius pilosulus Candèze, 1891 status resurrected; Limonis semianeus LeConte, 1853 status resurrected. Tetralimonius humeralis (Candèze, 1860) status resurrected; Tetralimonius maculicollis (Motschulsky, 1860) status resurrected. New replacement names are proposed for three homynyms: Limonius schimmeli Etzler new name for Neoathousius ferrugineus Schimmel and Platia, 1991; Elathous malatyanus Etzler new name for Elathous bicolor Platia, 2010, not Elathous bicolor (LeConte, 1853); and Microdesmes carteri Etzler new name for Limonius angulatus Carter, 1939 (= Microdesmes angulatus). Limonius kondratieffi Al Dhafer, 2009 is a new synonymy of Elathous bicolor (LeConte, 1853). A key to genera, generic descriptions, notes on species, and definitions of important characters are provided. 


1990 ◽  
Vol 68 (11) ◽  
pp. 2338-2351 ◽  
Author(s):  
S. A. Marshall ◽  
Ian P. Smith

All macropterous species of Aptilotus Mik are keyed, with descriptions of two new macropterous North American species, Aptilotus pogophallus and A. nigriphallus. New distributional records are given for other North American species, and brachyptery is noted for the first time in A. luctuosus (Spuler). Four new macropterous species of Aptilotus (glabrifrons, spinistylus, rufiscapus, and binotatus are described from Nepal. The relationships between the North American and Nepalese species are discussed. Minocellina Papp is synonomized with Aptilotus, and the two species formerly in Minocellina, A. thaii (Papp) and A. besucheti (Papp), are thus given as new combinations. Limosina carbonicolor Richards, from Ethiopia, is redescribed and transferred to Aptilotus.


Sign in / Sign up

Export Citation Format

Share Document