On the causes which influence the direction of the growth of roots. By Thomas Andrew Knight, Esq. F. R. S. In a letter to the Right Hon. Sir Joseph Banks, Bart. K. B. P. R. S
In a former paper Mr. Knight showed the influence of gravitation on the plumule and radicle of germinating seeds; in the present he considers the fibrous roots, which, with little comparative regard to gravity, extend themselves in whatever direction the greatest nutriment or moisture is to be found, with an appearance of predilection, which some naturalists have been disposed to compare to animal instinct. Mr. Knight examines whether this comparison has any foundation. He remarks, that when trees which require moisture are near to water, by far the greatest part of their roots extend themselves in that direction; but when those to which a dry soil is more congenial are placed in the same situation, their roots take an opposite course. When a tree is placed upon a wall, at a distance from the ground, its powers appear directed exclusively to one object; its roots are extended with well-directed efforts to reach the soil beneath, while its branches are in the mean time stinted in their growth; but as soon as the roots have entered the soil, the branches then grow with vigour and rapidity.