Leaf area index of an old-growth Douglas-fir forest estimated from direct structural measurements in the canopy

2000 ◽  
Vol 30 (12) ◽  
pp. 1922-1930 ◽  
Author(s):  
Sean C Thomas ◽  
William E Winner

Leaf area index (LAI) in old-growth Douglas-fir (Pseudotsuga menziesii var menziesii (Mirb.) Franco) forests exceeds that of any other forest ecosystem by some estimates; however, LAI determinations in coniferous forests have generally been indirect, involving extrapolations of patterns observed in younger stands. Aided by a 75-m construction crane for canopy access, we used a vertical line-intercept method to estimate LAI for a [Formula: see text]450-year-old Douglas-fir - western hemlock (Tsuga heterophylla (Raf.) Sarg.) forest in southwestern Washington state. LAI was calculated as the product of foliage contact frequency and an "extinction coefficient" accounting for foliage angular distribution, geometry, and the ratio of "interceptable" to total leaf area. LAI estimates were 9.3 ± 2.1 (estimate ± 95% confidence interval), 8.5 ± 2.2, and 8.2 ± 1.8 in 1997, 1998, and 1999, respectively, or 8.6 ± 1.1 pooled across years. Understory vegetation, including foliage of woody stems <5 cm diameter, represented 20% of this total. Sample points in which Douglas-fir was dominant had a higher total LAI than points dominated by western hemlock, including a higher LAI of understory vegetation. Our results do not support the contention that old-growth Douglas-fir - western hemlock forests maintain an appreciably higher LAI than do other forest ecosystems. Moreover, LAI in very old stands may decline as western hemlock replaces Douglas-fir through the course of succession.

Ecology ◽  
1986 ◽  
Vol 67 (4) ◽  
pp. 975-979 ◽  
Author(s):  
J. D. Marshall ◽  
R. H. Waring

2002 ◽  
Vol 32 (6) ◽  
pp. 1057-1070 ◽  
Author(s):  
Linda E Winter ◽  
Linda B Brubaker ◽  
Jerry F Franklin ◽  
Eric A Miller ◽  
Donald Q DeWitt

The history of canopy disturbances over the lifetime of an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) stand in the western Cascade Range of southern Washington was reconstructed using tree-ring records of cross-dated samples from a 3.3-ha mapped plot. The reconstruction detected pulses in which many western hemlock (Tsuga heterophylla (Raf.) Sarg.) synchronously experienced abrupt and sustained increases in ringwidth, i.e., "growth-increases", and focused on medium-sized or larger ([Formula: see text]0.8 ha) events. The results show that the stand experienced at least three canopy disturbances that each thinned, but did not clear, the canopy over areas [Formula: see text]0.8 ha, occurring approximately in the late 1500s, the 1760s, and the 1930s. None of these promoted regeneration of the shade-intolerant Douglas-fir, all of which established 1500–1521. The disturbances may have promoted regeneration of western hemlock, but their strongest effect on tree dynamics was to elicit western hemlock growth-increases. Canopy disturbances are known to create patchiness, or horizontal heterogeneity, an important characteristic of old-growth forests. This reconstructed history provides one model for restoration strategies to create horizontal heterogeneity in young Douglas-fir stands, for example, by suggesting sizes of areas to thin in variable-density thinnings.


2005 ◽  
Vol 35 (4) ◽  
pp. 990-1001 ◽  
Author(s):  
David C Shaw ◽  
Jiquan Chen ◽  
Elizabeth A Freeman ◽  
David M Braun

We investigated the distribution and severity of trees infected with western hemlock dwarf mistletoe (Arceuthobium tsugense (Rosendahl) G.N. Jones subsp. tsugense) in an old-growth Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) – western hemlock (Tsuga heterophylla (Raf.) Sarg.) forest. With the use of Hawksworth six-class dwarf mistletoe rating system, infection status was assessed for 3516 hemlock and true firs ≥5 cm diameter on a 12-ha stem-mapped plot located in the Cascade Mountains of southwest Washington State. Within the plot, 33% of the area had some level of infection and 25% (719) of western hemlocks, 2.2% (12) of Pacific silver fir (Abies amabilis (Dougl.) Forbes), and 29% (2) of noble fir (Abies procera Rehd.) trees were infected. Infected trees are larger than uninfected trees, on average, and within the infected tree population, the severely infected trees averaged larger than lightly infected trees. Abundant dwarf mistletoe in larger trees definitely positions the dwarf mistletoe population for future spread. Ripley's K analysis indicates a negative association between infected and uninfected hemlock trees, confirming that the infected trees form distinct dwarf mistletoe infection centers. The infection centers are actively spreading at their margins, which was confirmed by nearest neighbor analysis. Heavily infected trees had a negative association with uninfected trees, while lightly infected trees had a positive association with uninfected trees.


1991 ◽  
Vol 21 (7) ◽  
pp. 1127-1132 ◽  
Author(s):  
N. J. Smith ◽  
G. A. Borstad ◽  
D. A. Hill ◽  
R. C. Kerr

Techniques are developed to estimate stand leaf area index using high-resolution airborne spectral imagery. Leaf area index on 8 m radius plots was found to be strongly related to the normalized ratio of wavelengths in the red (674–687 nm) and near infrared (751–755 nm) part of the spectrum. Data from 17 stands were used. Leaf area index estimates included both the overstory (Douglas-fir, Pseudotsugamenziesii Mirb. (Franco)) and the understory (salal, Gaultheriashallon Pursh) over a range of stem densities.


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