The influence of stand structure on ecophysiological leaf characteristics of Pinus ponderosa in western Montana

2001 ◽  
Vol 31 (12) ◽  
pp. 2173-2182 ◽  
Author(s):  
Linda M Nagel ◽  
Kevin L O'Hara

The effects of vertical arrangement of foliage in even-aged and multiaged stand structures of ponderosa pine (Pinus ponderosa Dougl. ex P. Laws. & C. Laws.) on overall stand growth, light interception, and physiological leaf properties were tested on five plot pairs in western Montana. The primary structural difference between stand structures involves greater canopy depth and stratification of foliage in the multiaged stands. Both area- and mass-based maximum photosynthetic rates (Aarea and Amass) were relatively constant with canopy depth in both stand structures. Area- and mass-based leaf nitrogen (Narea and Nmass) decreased with increasing canopy depth in the even-aged stand structures but not in the multiaged. Specific leaf area (SLA) tended to increase with increasing canopy depth, although this relationship was only significant in the multiaged stand structures. The typical linear relationship observed for many species between photosynthetic rate and leaf nitrogen was not present in either stand structure; however, Narea was highly correlated to SLA in both even-aged and multiaged stand structures (R2 = 0.66 and R2 = 0.52, respectively). There were no differences in the light extinction coefficient (k), basal area growth or efficiency, or stand-level leaf area index between even-aged and multiaged plot pairs. Relative constancy in leaf physiology combined with similarities in site occupancy and growth rates help explain how different stand structures of ponderosa pine maintain similar rates of woody biomass productivity.

2008 ◽  
Vol 38 (4) ◽  
pp. 844-850 ◽  
Author(s):  
Gregory Peters ◽  
Anna Sala

Thinning and thinning followed by prescribed fire are common management practices intended to restore historic conditions in low-elevation ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) forests of the northern Rocky Mountains. While these treatments generally ameliorate the physiology and growth of residual trees, treatment-specific effects on reproductive output are not known. We examined reproductive output of second-growth ponderosa pine in western Montana 9 years after the application of four treatments: thinning, thinning followed by spring prescribed fire, thinning followed by fall prescribed fire, and unthinned control stands. Field and greenhouse observations indicated that reproductive traits vary depending on the specific management treatment. Cone production was significantly higher in trees from all actively managed stands relative to control trees. Trees subjected to prescribed fire produced cones with higher numbers of filled seeds than trees in unburned treatments. Seed mass, percentage germination, and seedling biomass were significantly lower for seeds from trees in spring burn treatments relative to all others and were generally higher in trees from fall burn treatments. We show for the first time that thinning and prescribed-burning treatments can influence reproductive output in ponderosa pine.


1999 ◽  
Vol 124 (1) ◽  
pp. 99-105 ◽  
Author(s):  
Claudio M. Dunan ◽  
Philip Westra ◽  
Frank D. Moore

A simulation model was built as a decision aid for management of five weed species in direct seeded irrigated onion (Allium cepa L.). The model uses the state variable approach and simulations are driven by temperature and sunlight as photosynthetically active radiation (PAR). It predicts yield reduction caused by competition for PAR according to the ratio of crop leaf area index (LAI) to weed LAI and respective light extinction coefficients (k). Input variables are plant density by species and average number of leaves by species. Number of leaves per plant is used by the model to provide an estimate of initial leaf area per plant. The model calculates initial species LAIs by multiplying species density times average leaf area per plant. The model accurately describes competitive interactions, taking into account respective plant densities, time of emergence, and time of weed removal. It permits economic evaluation of management factors such as handweeding, chemical weed control, herbicide phytotoxicity due to early application, and control of weed flushes during the season. The model is also used to evaluate mechanisms of plant competition for sunlight. In a sensitivity analysis, onion yield loss was more sensitive to weed PAR interception than to PAR use efficiency, the latter a species-dependent constant in the model.


2003 ◽  
Vol 33 (1) ◽  
pp. 164-170 ◽  
Author(s):  
John D Marshall ◽  
Robert A Monserud

Specific leaf area (SLA), the ratio of projected leaf area to leaf dry mass, is a critical parameter in many forest process models. SLA describes the efficiency with which the leaf captures light relative to the biomass invested in the leaf. It increases from top to bottom of a canopy, but it is unclear why. We sampled stands with low and elevated canopies (young and old stands) to determine whether SLA is related to water potential, as inferred from branch height and length, or shade, as inferred from branch position relative to the rest of the canopy, or both. We studied western white pine (Pinus monticola Dougl. ex D. Don), ponderosa pine (Pinus ponderosa Dougl. ex P. & C. Laws.), and interior Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco var. glauca) in northern Idaho. SLA decreased with branch height (P < 0.0001) at rates that varied among species (P < 0.0001). Branch length had no influence on SLA (P = 0.85). We detected no differences with canopy elevation (P = 0.90), but the slopes of lines relating SLA to branch height may have differed between the canopy elevation classes (P = 0.039). The results are consistent with predictions based on the hypothesis that SLA decreases as the gravitational component of water potential falls. The lack of a strong shading effect simplifies the estimation of canopy SLA for process models, requiring only species and branch heights.


1999 ◽  
Vol 14 (3) ◽  
pp. 137-143 ◽  
Author(s):  
Dayna M. Ayers ◽  
Donald J. Bedunah ◽  
Michael G. Harrington

Abstract In many western Montana ponderosa pine (Pinus ponderosa) stands, fire suppression and past selective logging of large trees have resulted in conditions favoring succession to dense stands of shade-tolerant, but insect- and disease-prone Douglas-fir (Pseudotsuga menziesii). Stand thinning and understory prescribed burning have been proposed as surrogates for pre-Euro-American settlement ecological processes and as potential treatments to improve declining forest condition and reduce the probability of severe wildfire. To test the effectiveness of these silvicultural techniques on overstory and understory conditions, research is ongoing in the Lick Creek Demonstration Site in the Bitterroot National Forest, Montana. Our research examined the response (mortality and vigor) of the dominant browse species, antelope bitterbrush (Purshia tridentata) and Scouler's willow (Salix scouleriana), to a ponderosa pine stand restoration project utilizing four treatments: (1) a shelterwood cut that removed 53% of the tree basal area; (2) a shelterwood cut with a low fuel consumption burn; (3) a shelterwood cut with a high fuel consumption burn; and (4) a control. Prior to the application of treatments, 1,856 bitterbrush and 871 willow were located, and their survival and vigor subsequently monitored for 2 yr posttreatment. The cut and burn treatments resulted in the greatest reduction in antelope bitterbrush and Scouler's willow density averaging 66% and 24% of pretreatment density, respectively. The shelterwood cut reduced bitterbrush and Scouler's willow density by 35% and 14%, respectively. On treatments receiving a shelterwood cut (all treatments but the control), but where antelope bitterbrush and Scouler's willow did not have fire damage, mortality was 45% for bitterbrush and 20% for willow, respectively. For bitterbrush and Scouler's willow plants that received fire damage, mortality was 72% for bitterbrush and 19% for willow. Although the burn and shelterwood harvest treatments resulted in reduced density of antelope bitterbrush and Scouler's willow 2 yr posttreatment, these treatments increased vigor of both species and created mineral seedbeds that may be necessary for establishment of seedlings. West. J. Appl. For. 14(3):137-143.


1992 ◽  
Vol 18 (4) ◽  
pp. 275-282 ◽  
Author(s):  
Peng Gong ◽  
Ruiliang Pu ◽  
John R. Miller

2003 ◽  
Vol 18 (3) ◽  
pp. 149-154 ◽  
Author(s):  
Chad E. Keyser ◽  
Kelsey S. Milner

Abstract This study examined long-term diameter and height response of ponderosa pine (Pinus ponderosa) and lodgepole pine (Pinus contorta var. latifolia) to a one-time application of herbicide during seedling establishment in western Montana. Two herbicide trials initiated in 1981 and 1983 by Champion International Corporation were resurrected during the summer of 1997. Analysis of variance (ANOVA) F-tests along with Duncan's Multiple Range tests (DMR) were performed on mean tree dbh, tree height, and competing vegetation percent cover estimates. In addition, a graphical analysis of mean annual height growth over time was performed. Significant increases (alpha = 0.05) in ponderosa pine dbh (25 to 44%) and height (11 to 28%) were found after 16 yr of growth on Velpar L. (hexazinone) treated plots, while lodgepole pine had significant increases in dbh (70 to 118%) and height (41 to 82%) after 15 yr of growth on Velpar L. treated plots. Graphical analyses indicated a positive height growth effect for a period of 11 to 13 yr following treatment for ponderosa pine; however, total height gains had decreased in recent years. Positive increases in height growth for lodgepole pine were continuing after 15 yr of growth. These results indicate that a one-time application of herbicide applied during seedling establishment will promote faster tree growth in western Montana. We also note that proper density management will be necessary early in the life of the stand to maintain height gains. West. J. Appl. For. 18(3):149–154.


1995 ◽  
Vol 25 (9) ◽  
pp. 1553-1557 ◽  
Author(s):  
Kevin L. O'Hara ◽  
Narayanan I. Valappil

Ponderosa pine (Pinusponderosa Dougl. ex Laws.) frequently grows in pure, multi-aged stands throughout its range. Sapwood–leaf area prediction equations were developed for multi-aged, multi-strata ponderosa pine stands in western Montana and central Oregon. No significant differences were found between equations for trees from lower or upper strata, or between equations for all trees and equations for upper or lower strata trees in either study location. These results indicate overstory ponderosa pine trees do not require significantly greater sapwood conducting tissue per unit of leaf area than understory trees. Single variable models using only sapwood area at breast height are recommended.


2007 ◽  
Vol 37 (11) ◽  
pp. 2096-2105 ◽  
Author(s):  
Kelsey Sherich ◽  
Amy Pocewicz ◽  
Penelope Morgan

Trees respond to edge-to-interior microclimate differences in fragmented forests. To better understand tree physiological responses to fragmentation, we measured ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws) and Douglas-fir ( Pseudotsuga menziesii (Mirbel) Franco) leaf area, crown ratios, sapwood area, basal area (BA) growth rates, and BA growth efficiency at 23 long-established (>50 year) forest edges in northern Idaho. Trees located at forest edges had more leaf area, deeper crowns, higher BA growth rates, and more sapwood area at breast height than interior trees. Ponderosa pine had significantly higher BA growth efficiency at forest edges than interiors, but Douglas-fir BA growth efficiency did not differ, which may relate to differences in photosynthetic capacity and drought and shade tolerance. Edge orientation affected BA growth efficiency, with higher values at northeast-facing edges for both species. Edge effects were significant even after accounting for variation in stand density, which did not differ between the forest edge and interior. Although edge trees had significantly greater canopy depth on their edge-facing than forest-facing side, sapwood area was evenly distributed. We found no evidence that growing conditions at the forest edge were currently subjecting trees to stress, but higher leaf area and deeper crowns could result in lower tolerance to future drought conditions.


Sign in / Sign up

Export Citation Format

Share Document