Influence of repeated fertilization on forest ecosystems: relative habitat use by mule deer and moose

2006 ◽  
Vol 36 (6) ◽  
pp. 1395-1406 ◽  
Author(s):  
Thomas P Sullivan ◽  
Druscilla S Sullivan ◽  
Pontus M.F Lindgren ◽  
Douglas B Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Mule Deer ◽  
Odocoileus Hemionus ◽  
Old Growth ◽  
Winter Habitat ◽  
South Central ◽  
Mature Stands

This study was designed to test the hypothesis that large-scale precommercial thinning (PCT) and repeated fertilization of young lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) stands would enhance relative habitat use by mule deer (Odocoileus hemionus Rafinesque) and moose (Alces alces L.) in summer and winter periods, compared to that in mature and old-growth stands. Replicate study areas were located near Summerland, Kelowna, and Williams Lake in south central British Columbia, Canada. Each study area had a range of PCT densities, with and without fertilization, and mature and old-growth stands. Habitat use in summer and winter was measured by pellet-group counts of deer and moose from 1999 to 2003, 6–10 years after the onset of treatments. During summer months, habitat use by deer was enhanced by PCT with fertilization, and the 1000 stems/ha fertilized stands experienced greater use than the unthinned or mature stands. Winter habitat use by deer was similar in the 1000 stems/ha fertilized stands and old-growth stands. In both summer and winter, moose preferred fertilized to unfertilized stands and low-density to high-density stands. Intensive management of young lodgepole pine forests has considerable potential to develop summer and winter ranges for these ungulates.

Response of northwestern chipmunks (Tamias amoenus) to variable habitat structure in young lodgepole pine forest

2000 ◽  
Vol 78 (2) ◽  
pp. 283-293 ◽  
Author(s):  
Thomas P Sullivan ◽  
Walt Klenner
Keyword(s):  
Pinus Contorta ◽  
Habitat Structure ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Volume Index ◽  
Low Density ◽  
Old Growth ◽  
Ecological Zones ◽  
South Central ◽  
Tamias Amoenus

This study was designed to test the hypothesis that large-scale habitat alteration by stand thinning over a range of densities would increase the abundance and related population dynamics of northwestern chipmunks (Tamias amoenus) in young lodgepole pine (Pinus contorta) forest. Replicate study areas were located near Penticton, Kamloops, and Prince George in south-central British Columbia, Canada. Each study area had three stands thinned to densities of approximately 500 (low), 1000 (medium), and 2000 (high) stems/ha, with an unthinned young pine stand and an old-growth pine stand for comparison. Chipmunk populations were sampled intensively in thinned stands from 1989 to 1991 and in the unthinned and old-growth stands from 1990 to 1991. Habitat structure was sampled in all stands in 1990. For herbs and shrubs, the crown volume index values were similar among stands; for trees, this index was lowest for the low-density stands, with the index for all three thinned stands being lower than that for the unthinned stands. Species diversity and the structural diversity of vegetation were similar among stands. The abundance of chipmunks was significantly higher in low-density than in high-density thinned stands at Penticton (1.3-1.9 times higher) and Prince George (2.4-3.8 times higher) but not at Kamloops. Chipmunks were less abundant in old-growth stands than in the other four treatment stands. Breeding performance and recruitment followed the same pattern as abundance. Chipmunk survival was generally similar among stands. There were heavier chipmunks in the low-density stands in some years at Penticton. Northwestern chipmunks appear to prefer "open" habitats generated by heavy thinning of young lodgepole pine stands. This result was achieved in three different forest ecological zones and may enhance the overall forest ecosystem.

Long-term responses of mammalian herbivores to stand thinning and fertilization in young lodgepole pine (Pinus contorta var. latifolia) forest

2010 ◽  
Vol 40 (12) ◽  
pp. 2302-2312 ◽  
Author(s):  
Thomas P. Sullivan ◽  
Druscilla S. Sullivan ◽  
Pontus M.F. Lindgren ◽  
Douglas B. Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
South Central ◽  
Commercial Thinning ◽  
Snowshoe Hares ◽  
Understory Shrubs

Snowshoe hares (Lepus americanus Exrleben, 1777), mule deer (Odocoileus hemionus (Rafinesque, 1817)), and moose (Alces alces (L., 1758)) commonly occur in young coniferous forests. This study was designed to test the hypothesis that large-scale pre-commercial thinning (PCT) and repeated fertilization 15–20 years after the onset of treatments in young lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.) stands would enhance relative habitat use by hares, deer, and moose compared with unmanaged stands. Study areas were located in south-central British Columbia, Canada. Habitat use was measured by fecal pellet and pellet-group counts. Understory vegetation and coniferous stand structure were measured in all stands. Habitat use by deer and moose was highest in heavily thinned stands, probably due to the higher levels of forage and cover provided by understory shrubs and conifers in thinned stands. Habitat use by snowshoe hares was highest in high-density stands, but also in lower-density (≤1000 stems·ha–1) stands where an increase in understory conifers provided essential cover for hares. Managers should consider the long-term nature of understory development in young stands managed for timber production. Heavy thinning (≤1000 stems·ha–1) will generate suitable understory habitat for these herbivores sooner than conventional PCT at higher stand densities.

Influence of repeated fertilization on forest ecosystems: relative habitat use by snowshoe hares (Lepus americanus)

2006 ◽  
Vol 36 (9) ◽  
pp. 2080-2089 ◽  
Author(s):  
Thomas P Sullivan ◽  
Druscilla S Sullivan ◽  
Pontus MF Lindgren ◽  
Douglas B Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Structural Diversity ◽  
Lepus Americanus ◽  
Volume Index ◽  
South Central ◽  
Crown Volume ◽  
Snowshoe Hares

This study was designed to test the hypothesis that large-scale precommercial thinning (PCT) and repeated fertilization of lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) would enhance relative habitat use by snowshoe hares (Lepus americanus Erxleben) in managed stands. Study areas were located near Summerland, Kelowna, and Williams Lake in south-central British Columbia, Canada. Each study area had nine treatments: four pairs of stands thinned to densities of 250, 500, 1000, and 2000 stems/ha, with one stand of each pair fertilized five times at 2-year intervals, and an unthinned stand. Understory vegetation and relative habitat use by snowshoe hares were measured annually from 1999 to 2003, 6–10 years after the onset of treatments. Mean crown volume index of herbs was significantly higher in fertilized than unfertilized stands, but density had no effect. Shrub volume was not affected by either treatment. Mean crown volume index of trees was significantly greater in the fertilized and high-density stands. Mean total richness of vascular plants was significantly reduced by fertilization. Mean total structural diversity of vegetation was highest in the low-density stands but was not affected by fertilization. Relative habitat use by hares, based on fecal pellet counts, was highest in the 2000 stems/ha and unthinned stands in summer. This pattern also occurred in winter when hare use was higher in fertilized than unfertilized stands. Overall, fertilized 2000 stems/ha stands provided habitat for hares to a degree comparable with unthinned stands of lodgepole pine.

scholarly journals Silviculture and Wildlife: Snowshoe Hare Abundance across a Successional Sequence of Natural and Intensively Managed Forests

ISRN Ecology ◽  
2012 ◽  
Vol 2012 ◽  
pp. 1-10 ◽  
Author(s):  
Thomas P. Sullivan ◽  
Druscilla S. Sullivan ◽  
Pontus M. F. Lindgren ◽  
Douglas B. Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
Bimodal Distribution ◽  
Managed Forests ◽  
Old Growth ◽  
South Central ◽  
Intensively Managed ◽  
Stand Attributes

We tested the hypotheses H1 that relative habitat use by snowshoe hares (Lepus americanus) would have a bimodal distribution with the highest abundance in young lodgepole pine (Pinus contorta) stands (both managed and unmanaged), minimal numbers in mature forests, and moderate abundance in old-growth forests and H2 that habitat use would increase in response to enhanced stand attributes from PCT (precommercial thinning) and fertilization treatments. Habitat use was measured by counts of fecal pellets of hares from 1999 to 2003 in forest stands in south-central British Columbia, Canada. Our results did not support the bimodal distribution of hares among coniferous stands, such that old-growth stands, at least in our region, do not provide sufficient habitat for hare populations. High-density (5000 to 13000 stems/ha) unthinned young lodgepole pine stands provide optimum habitat for hares in terms of overstory and stand structure. Thinned and fertilized stands may also provide habitat, particularly at densities ≤1000 stems/ha, and over time as understory conifers develop. Managed stands provided habitat for hares at the same level as mature stands, at 6–10 years after PCT. Maintenance of a range of managed and unmanaged stands in a landscape mosaic would be ideal for integration of silvicultural and wildlife management goals.

scholarly journals Late-Winter Habitat Use by Mule Deer, Odocoileus hemionus, in Central Interior British Columbia

2008 ◽  
Vol 122 (3) ◽  
pp. 205
Author(s):  
Gilbert Proulx
Keyword(s):  
British Columbia ◽  
Habitat Use ◽  
Populus Tremuloides ◽  
Negative Impact ◽  
Lodgepole Pine ◽  
Mule Deer ◽  
Odocoileus Hemionus ◽  
Late Winter ◽  
High Quality ◽  
Winter Habitat

In central interior British Columbia, extensive cut blocks to recover timber killed by the Mountain Pine Beetle (Dendroctonus ponderosae) could impact negatively on Mule Deer (Odocoileus hemionus) late-winter habitat. This study assessed the possibility of predicting the distribution of potential late-winter habitat for Mule Deer with the BC Vegetation Resources Inventory (VRI) dataset used to produce forestry maps. On the basis of literature review and roadside inventories in December 2004, I predicted that high-quality Mule Deer late-winter habitat would correspond to mature and old conifer-dominated stands with ≤ 20% deciduous species, a canopy closure ≥ 45%, tree heights ≥ 23 m, tree diameter at breast height ≥ 24 cm, and basal area ≥ 45 m2/ha, and would be located on < 60% slopes on south, southeast, southwest or west aspects, or on flat ground. I allocated weight values to these criteria to classify map polygons into high-, medium- and low-quality polygons, and produce predictive maps of late-winter habitat use by Mule Deer. I tested my predictive habitat rating by snowtracking along 18 km of transects in February-March 2006, and 15.6 km of transects in February 2007. I recorded 31 and 12 Mule Deer tracks in 2006 and 2007, respectively, all in high-quality polygons. The observed frequency of tracks per polygon type was significantly (P < 0.001) different from expected. All tracks were in mature and old conifer-dominated stands including 10-60% Lodgepole Pine (Pinus contorta) and 10-20% Trembling Aspen (Populus tremuloides). This study showed that it was possible to predict the distribution of potential late-winter habitat for Mule Deer using a series of habitat criteria and the VRI dataset. The extensive harvesting of Lodgepole Pine in mixed coniferous stands will undoubtedly have a negative impact on Mule Deer late-winter habitat quality and quantity. The rating of habitat types developed in this study should be used in forest management plans to determine sites that should be protected from logging.

scholarly journals Long-term influence of stand thinning and repeated fertilization on forage production in young lodgepole pine forests

2017 ◽  
Vol 47 (8) ◽  
pp. 1123-1130 ◽  
Author(s):  
Pontus M.F. Lindgren ◽  
Thomas P. Sullivan ◽  
Douglas B. Ransome ◽  
Druscilla S. Sullivan ◽  
Lisa Zabek
Keyword(s):  
Pinus Contorta ◽  
Large Scale ◽  
Bos Taurus ◽  
Lodgepole Pine ◽  
Stand Density ◽  
Stand Age ◽  
Forage Production ◽  
South Central ◽  
Stand Thinning

Integration of trees with forage and livestock production as silvopastoralism is another potential component of intensive forest management. Stand thinning and fertilization may enhance growth of crop trees and understory forage for livestock. We tested the hypothesis that large-scale precommercial thinning (PCT) (particularly heavy thinning to ≤1000 stems·ha−1) and repeated fertilization, up to 20 years after the onset of treatments, would enhance production of graminoids, forbs, and shrubs as cattle (Bos taurus L.) forage. Results are from two long-term studies: (1) PCT (1988–2013) and (2) PCT with fertilization (PCT–FERT) (1993–2013) of lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) stands in south-central British Columbia, Canada. Mean biomass estimates of graminoids, forbs, total herbs, and shrubs were not affected by stand density. However, fertilization enhanced mean biomass estimates of graminoids, forbs, and total herbs, but not shrubs. Thus, the density part of our hypothesis was not supported, but the nutrient addition part was supported at least for the herbaceous components. Biomass of the herbaceous understory was maintained as a silvopasture component for up to 20 years (stand age 13 to 33 years) in fertilized heavily thinned stands prior to canopy closure.

scholarly journals Winter Habitat Use by Moose, Alces alces, in Central Interior British Columbia

2005 ◽  
Vol 119 (2) ◽  
pp. 186 ◽  
Author(s):  
Gilbert Proulx ◽  
Rhonda M. Kariz
Keyword(s):  
British Columbia ◽  
Habitat Use ◽  
Pinus Contorta ◽  
Mountain Pine Beetle ◽  
Lodgepole Pine ◽  
Alces Alces ◽  
Late Winter ◽  
Winter Habitat ◽  
Pine Beetle ◽  
Pine Stands

In central British Columbia, recent epidemics of Mountain Pine Beetle (Dendroctonus ponderosae) have resulted in the use of expansive clearcut areas to remove infested mature and old Lodgepole Pine (Pinus contorta) stands. This study aimed to determine if Moose (Alces alces) use late-successional Lodgepole Pine stands in mid- to late-winter. Moose activity and habitat use was determined during February-March track surveys in 2000 (60 km) and 2001 (55.7 km). In 2000 (69 tracks) and 2001 (313 tracks), Moose track distribution differed significantly (P < 0.05) from random. They were significantly more abundant than predicted in young stands (dominated by Picea spp.), or early seral stages; they were less abundant than predicted in mature and old Lodgepole Pine stands. It is unlikely that harvesting late-successional Lodgepole Pine stands would affect Moose winter habitat supply.

Influence of fire severity and vegetation treatments on mule deer (Odocoileus hemionus) winter habitat use on the Kaibab Plateau, Arizona

2020 ◽  
Vol 60 (10) ◽  
pp. 1292
Author(s):  
Kirby D. Bristow ◽  
Larisa E. Harding ◽  
Richard W. Lucas ◽  
Thomas C. McCall
Keyword(s):  
Habitat Use ◽  
Fire Regime ◽  
Fire Severity ◽  
Mule Deer ◽  
Odocoileus Hemionus ◽  
Winter Habitat ◽  
Similar Treatment ◽  
Natural Fire ◽  
Winter Range ◽  
Kaibab Plateau

Context Wildfire and vegetation treatments affect mule deer (Odocoileus hemionus) populations across the western United States. However, the relative influence of fire and treatments on habitat use by mule deer in Arizona is not well defined. Aims We examined locations of mule deer on the Kaibab Plateau in northern Arizona, so as to determine the influence of vegetation treatments and wildfire severity on deer habitat-use patterns across their winter range where fires and treatments had occurred previously. Methods We used locations (n = 11297) from 21 adult female mule deer fitted with global positioning system collars to model probability of use as a function of habitat covariates. Key results The best model describing winter-range habitat use by mule deer on the Kaibab Plateau included covariates describing the age of vegetation treatments and fire severity. Increased deer use in winter was associated with areas of lower terrain ruggedness and reduced snow depths. Deer use also increased in areas that experienced a higher average fire severity, resulting in decreased vegetation heights. Among treatment age classes, deer use was greatest in areas containing vegetation treatments that were ≤6 years old, but negatively associated with treatments that were &gt;6 years old. Conclusions Vegetation treatments designed to remove or reduce less palatable tree and shrub species to improve forage conditions may increase the use of winter habitats by deer on the Kaibab Plateau. Similarly, prescribed fire and rangeland treatments designed to return areas to a more natural fire regime and, thereby, generate new plant growth, may improve winter-range habitat conditions for mule deer. Implications Similar treatment strategies may also benefit mule deer populations throughout the western USA, by improving forage conditions on critical habitats and reducing the potential for catastrophic wildfire.

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