Long-term responses of mammalian herbivores to stand thinning and fertilization in young lodgepole pine (Pinus contorta var. latifolia) forest

2010 ◽  
Vol 40 (12) ◽  
pp. 2302-2312 ◽  
Author(s):  
Thomas P. Sullivan ◽  
Druscilla S. Sullivan ◽  
Pontus M.F. Lindgren ◽  
Douglas B. Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
South Central ◽  
Commercial Thinning ◽  
Snowshoe Hares ◽  
Understory Shrubs

Snowshoe hares (Lepus americanus Exrleben, 1777), mule deer (Odocoileus hemionus (Rafinesque, 1817)), and moose (Alces alces (L., 1758)) commonly occur in young coniferous forests. This study was designed to test the hypothesis that large-scale pre-commercial thinning (PCT) and repeated fertilization 15–20 years after the onset of treatments in young lodgepole pine (Pinus contorta var. latifolia Engelm. ex S. Wats.) stands would enhance relative habitat use by hares, deer, and moose compared with unmanaged stands. Study areas were located in south-central British Columbia, Canada. Habitat use was measured by fecal pellet and pellet-group counts. Understory vegetation and coniferous stand structure were measured in all stands. Habitat use by deer and moose was highest in heavily thinned stands, probably due to the higher levels of forage and cover provided by understory shrubs and conifers in thinned stands. Habitat use by snowshoe hares was highest in high-density stands, but also in lower-density (≤1000 stems·ha–1) stands where an increase in understory conifers provided essential cover for hares. Managers should consider the long-term nature of understory development in young stands managed for timber production. Heavy thinning (≤1000 stems·ha–1) will generate suitable understory habitat for these herbivores sooner than conventional PCT at higher stand densities.

Influence of repeated fertilization on forest ecosystems: relative habitat use by snowshoe hares (Lepus americanus)

2006 ◽  
Vol 36 (9) ◽  
pp. 2080-2089 ◽  
Author(s):  
Thomas P Sullivan ◽  
Druscilla S Sullivan ◽  
Pontus MF Lindgren ◽  
Douglas B Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Structural Diversity ◽  
Lepus Americanus ◽  
Volume Index ◽  
South Central ◽  
Crown Volume ◽  
Snowshoe Hares

This study was designed to test the hypothesis that large-scale precommercial thinning (PCT) and repeated fertilization of lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) would enhance relative habitat use by snowshoe hares (Lepus americanus Erxleben) in managed stands. Study areas were located near Summerland, Kelowna, and Williams Lake in south-central British Columbia, Canada. Each study area had nine treatments: four pairs of stands thinned to densities of 250, 500, 1000, and 2000 stems/ha, with one stand of each pair fertilized five times at 2-year intervals, and an unthinned stand. Understory vegetation and relative habitat use by snowshoe hares were measured annually from 1999 to 2003, 6–10 years after the onset of treatments. Mean crown volume index of herbs was significantly higher in fertilized than unfertilized stands, but density had no effect. Shrub volume was not affected by either treatment. Mean crown volume index of trees was significantly greater in the fertilized and high-density stands. Mean total richness of vascular plants was significantly reduced by fertilization. Mean total structural diversity of vegetation was highest in the low-density stands but was not affected by fertilization. Relative habitat use by hares, based on fecal pellet counts, was highest in the 2000 stems/ha and unthinned stands in summer. This pattern also occurred in winter when hare use was higher in fertilized than unfertilized stands. Overall, fertilized 2000 stems/ha stands provided habitat for hares to a degree comparable with unthinned stands of lodgepole pine.

scholarly journals Long-term influence of stand thinning and repeated fertilization on forage production in young lodgepole pine forests

2017 ◽  
Vol 47 (8) ◽  
pp. 1123-1130 ◽  
Author(s):  
Pontus M.F. Lindgren ◽  
Thomas P. Sullivan ◽  
Douglas B. Ransome ◽  
Druscilla S. Sullivan ◽  
Lisa Zabek
Keyword(s):  
Pinus Contorta ◽  
Large Scale ◽  
Bos Taurus ◽  
Lodgepole Pine ◽  
Stand Density ◽  
Stand Age ◽  
Forage Production ◽  
South Central ◽  
Stand Thinning

Integration of trees with forage and livestock production as silvopastoralism is another potential component of intensive forest management. Stand thinning and fertilization may enhance growth of crop trees and understory forage for livestock. We tested the hypothesis that large-scale precommercial thinning (PCT) (particularly heavy thinning to ≤1000 stems·ha−1) and repeated fertilization, up to 20 years after the onset of treatments, would enhance production of graminoids, forbs, and shrubs as cattle (Bos taurus L.) forage. Results are from two long-term studies: (1) PCT (1988–2013) and (2) PCT with fertilization (PCT–FERT) (1993–2013) of lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) stands in south-central British Columbia, Canada. Mean biomass estimates of graminoids, forbs, total herbs, and shrubs were not affected by stand density. However, fertilization enhanced mean biomass estimates of graminoids, forbs, and total herbs, but not shrubs. Thus, the density part of our hypothesis was not supported, but the nutrient addition part was supported at least for the herbaceous components. Biomass of the herbaceous understory was maintained as a silvopasture component for up to 20 years (stand age 13 to 33 years) in fertilized heavily thinned stands prior to canopy closure.

scholarly journals Silviculture and Wildlife: Snowshoe Hare Abundance across a Successional Sequence of Natural and Intensively Managed Forests

ISRN Ecology ◽  
2012 ◽  
Vol 2012 ◽  
pp. 1-10 ◽  
Author(s):  
Thomas P. Sullivan ◽  
Druscilla S. Sullivan ◽  
Pontus M. F. Lindgren ◽  
Douglas B. Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
Bimodal Distribution ◽  
Managed Forests ◽  
Old Growth ◽  
South Central ◽  
Intensively Managed ◽  
Stand Attributes

We tested the hypotheses H1 that relative habitat use by snowshoe hares (Lepus americanus) would have a bimodal distribution with the highest abundance in young lodgepole pine (Pinus contorta) stands (both managed and unmanaged), minimal numbers in mature forests, and moderate abundance in old-growth forests and H2 that habitat use would increase in response to enhanced stand attributes from PCT (precommercial thinning) and fertilization treatments. Habitat use was measured by counts of fecal pellets of hares from 1999 to 2003 in forest stands in south-central British Columbia, Canada. Our results did not support the bimodal distribution of hares among coniferous stands, such that old-growth stands, at least in our region, do not provide sufficient habitat for hare populations. High-density (5000 to 13000 stems/ha) unthinned young lodgepole pine stands provide optimum habitat for hares in terms of overstory and stand structure. Thinned and fertilized stands may also provide habitat, particularly at densities ≤1000 stems/ha, and over time as understory conifers develop. Managed stands provided habitat for hares at the same level as mature stands, at 6–10 years after PCT. Maintenance of a range of managed and unmanaged stands in a landscape mosaic would be ideal for integration of silvicultural and wildlife management goals.

Influence of repeated fertilization on forest ecosystems: relative habitat use by mule deer and moose

2006 ◽  
Vol 36 (6) ◽  
pp. 1395-1406 ◽  
Author(s):  
Thomas P Sullivan ◽  
Druscilla S Sullivan ◽  
Pontus M.F Lindgren ◽  
Douglas B Ransome
Keyword(s):  
Habitat Use ◽  
Pinus Contorta ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Mule Deer ◽  
Odocoileus Hemionus ◽  
Old Growth ◽  
Winter Habitat ◽  
South Central ◽  
Mature Stands

This study was designed to test the hypothesis that large-scale precommercial thinning (PCT) and repeated fertilization of young lodgepole pine (Pinus contorta Dougl. ex Loud. var. latifolia Engelm.) stands would enhance relative habitat use by mule deer (Odocoileus hemionus Rafinesque) and moose (Alces alces L.) in summer and winter periods, compared to that in mature and old-growth stands. Replicate study areas were located near Summerland, Kelowna, and Williams Lake in south central British Columbia, Canada. Each study area had a range of PCT densities, with and without fertilization, and mature and old-growth stands. Habitat use in summer and winter was measured by pellet-group counts of deer and moose from 1999 to 2003, 6–10 years after the onset of treatments. During summer months, habitat use by deer was enhanced by PCT with fertilization, and the 1000 stems/ha fertilized stands experienced greater use than the unthinned or mature stands. Winter habitat use by deer was similar in the 1000 stems/ha fertilized stands and old-growth stands. In both summer and winter, moose preferred fertilized to unfertilized stands and low-density to high-density stands. Intensive management of young lodgepole pine forests has considerable potential to develop summer and winter ranges for these ungulates.

scholarly journals Twenty-Five Years after Stand Thinning and Repeated Fertilization in Lodgepole Pine Forest: Implications for Tree Growth, Stand Structure, and Carbon Sequestration

Forests ◽  
2020 ◽  
Vol 11 (3) ◽  
pp. 337
Author(s):  
Thomas P. Sullivan ◽  
Druscilla S. Sullivan ◽  
Pontus M.F. Lindgren ◽  
Douglas B. Ransome ◽  
Lisa Zabek
Keyword(s):  
Carbon Sequestration ◽  
Carbon Storage ◽  
Large Scale ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
Wood Products ◽  
Partial Support ◽  
South Central ◽  
Commercial Thinning ◽  
Total Crop

Silvicultural practices such as pre-commercial thinning (PCT) and repeated fertilization have been used successfully around the world to increase forest biomass for conventional wood products, biofuels, and carbon sequestration. Two complementary studies were designed to test the hypotheses (H) that large-scale PCT and PCT with repeated fertilization of young (13–17 years old) lodgepole pine (Pinus contorta var. latifolia) stands, at 25 years after the onset of treatments, would enhance (H1) productivity and structural features (diameter and height growth, crown volume and dimensions) of crop trees; (H2) merchantable volume of crop trees, (H3) abundance of understory conifer, herb and shrub layers, and (H4) carbon storage (sequestration) of all layers. Results are from two studies in south-central British Columbia, Canada. The PCT study had three densities: 500, 1000, and 2000 stems/ha, an unthinned, and old-growth stand replicated at three areas. The PCT-FERT study had four densities: 250, 500, 1000, and 2000 stems/ha with and without a repeated fertilization treatment, and an unthinned-unfertilized stand, replicated at two areas. Heavily thinned (≤1000 stems/ha) and fertilized stands generally had larger mean diameters and crown dimensions than lightly thinned or unfertilized stands, whereas mean heights of crop trees remained relatively unaffected, and hence partial support for H1. Despite differences in stand density (4- and 8-fold) of original crop trees, ingress of intermediate crop trees over the 25 years changed density dramatically. There was no support for H2, as mean merchantable volume of total crop trees was similar across stand densities owing to ingress, and was also similar in fertilized and unfertilized stands. In stands thinned to ≤500 trees/ha, there is an anticipated increase in crop tree density of 2.2 to 4.3 times over 25 years compared with the original post-thinning densities. Mean abundance of understory conifers was generally similar among stands owing to successional development towards canopy closure, whereas herbs and shrubs persisted only in canopy gaps in heavily thinned stands, thereby providing no support for H3. Mean carbon storage was similar across densities in both studies with ingress contributing a considerable amount to carbon sequestration at the lower densities. Fertilization increased (1.4 times) mean carbon storage significantly among total crop trees, total conifers, and the grand total of all layers, thereby providing partial support for H4. Thus, much of the potential above-ground carbon storage lost by thinning was restored in the heavily thinned stands at 25 years post-treatment.

Response of northwestern chipmunks (Tamias amoenus) to variable habitat structure in young lodgepole pine forest

2000 ◽  
Vol 78 (2) ◽  
pp. 283-293 ◽  
Author(s):  
Thomas P Sullivan ◽  
Walt Klenner
Keyword(s):  
Pinus Contorta ◽  
Habitat Structure ◽  
Large Scale ◽  
Lodgepole Pine ◽  
Volume Index ◽  
Low Density ◽  
Old Growth ◽  
Ecological Zones ◽  
South Central ◽  
Tamias Amoenus

This study was designed to test the hypothesis that large-scale habitat alteration by stand thinning over a range of densities would increase the abundance and related population dynamics of northwestern chipmunks (Tamias amoenus) in young lodgepole pine (Pinus contorta) forest. Replicate study areas were located near Penticton, Kamloops, and Prince George in south-central British Columbia, Canada. Each study area had three stands thinned to densities of approximately 500 (low), 1000 (medium), and 2000 (high) stems/ha, with an unthinned young pine stand and an old-growth pine stand for comparison. Chipmunk populations were sampled intensively in thinned stands from 1989 to 1991 and in the unthinned and old-growth stands from 1990 to 1991. Habitat structure was sampled in all stands in 1990. For herbs and shrubs, the crown volume index values were similar among stands; for trees, this index was lowest for the low-density stands, with the index for all three thinned stands being lower than that for the unthinned stands. Species diversity and the structural diversity of vegetation were similar among stands. The abundance of chipmunks was significantly higher in low-density than in high-density thinned stands at Penticton (1.3-1.9 times higher) and Prince George (2.4-3.8 times higher) but not at Kamloops. Chipmunks were less abundant in old-growth stands than in the other four treatment stands. Breeding performance and recruitment followed the same pattern as abundance. Chipmunk survival was generally similar among stands. There were heavier chipmunks in the low-density stands in some years at Penticton. Northwestern chipmunks appear to prefer "open" habitats generated by heavy thinning of young lodgepole pine stands. This result was achieved in three different forest ecological zones and may enhance the overall forest ecosystem.

Forest structure altered by mountain pine beetle outbreaks affects subsequent attack in a Wyoming lodgepole pine forest, USA

2011 ◽  
Vol 41 (12) ◽  
pp. 2403-2412 ◽  
Author(s):  
Daniel M. Kashian ◽  
Rebecca M. Jackson ◽  
Heather D. Lyons
Keyword(s):  
Pinus Contorta ◽  
Mountain Pine Beetle ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
Climate Change Scenarios ◽  
Cone Production ◽  
Mountain Pine ◽  
Large Trees ◽  
Pine Beetle ◽  
The 1960S

Extensive outbreaks of the mountain pine beetle ( Dendroctonus ponderosae Hopkins) will alter the structure of many stands that will likely be attacked again before experiencing a stand-replacing fire. We examined a stand of lodgepole pine ( Pinus contorta var. latifolia Engelm. ex S. Watson) in Grand Teton National Park currently experiencing a moderate-level outbreak and previously attacked by mountain pine beetle in the 1960s. Consistent with published studies, tree diameter was the main predictor of beetle attack on a given tree, large trees were preferentially attacked, and tree vigor, age, and cone production were unimportant variables for beetle attack at epidemic levels. Small trees killed in the stand were killed based mainly on their proximity to large trees and were likely spatially aggregated with large trees as a result of the previous outbreak. We concluded that the driving factors of beetle attack and their spatial patterns are consistent across outbreak severities but that stand structure altered by the previous outbreak had implications for the current outbreaks in the same location. This study should catalyze additional research that examines how beetle-altered stand structure affects future outbreaks — an important priority for predicting their impacts under climate change scenarios that project increases in outbreak frequency and extent.

Recovery time of snowshoe hare habitat after commercial thinning in boreal Quebec

2012 ◽  
Vol 42 (1) ◽  
pp. 123-133 ◽  
Author(s):  
Guylaine Bois ◽  
Louis Imbeau ◽  
Marc J. Mazerolle
Keyword(s):  
Habitat Use ◽  
Black Spruce ◽  
Keystone Species ◽  
Snowshoe Hare ◽  
Negative Effects ◽  
Boreal Ecosystems ◽  
Commercial Thinning ◽  
Long Term Impact ◽  
Habitat Parameters

As short-term effects of partial cuts generally decrease available cover for snowshoe hare ( Lepus americanus Erxleben), most studies have shown negative effects of such treatments on this keystone species in boreal ecosystems. This study aims to determine the long-term impact of commercial thinning on snowshoe hare habitat, and we hypothesized that habitat quality, as well as habitat use, recovers with time since treatment. We selected stands aged 50–90 years dominated by black spruce ( Picea mariana (Mill.) Britton, Sterns & Poggenb.) in Abitibi (Quebec). We used models of habitat parameters to explain the abundance of snowshoe hare tracks and pellets in 20 commercially thinned stands treated between 1989 and 1999 and 12 control stands. Lateral cover was the dominant parameter influencing snowshoe hare habitat use. On average, commercially thinned stands had a lower lateral cover than controls (–18%). We also found that snowshoe hare use of commercially thinned stands increases with time since treatment. However, 11–18 years are needed before commercially thinned stands return to the same level of lateral cover and snowshoe hare signs as control stands. Commercial thinning is generally followed by harvesting all merchantable stems 15 years after treatment. Thus, we suggest that commercial thinning as currently practiced should be avoided if the objective is to maintain quality habitat for snowshoe hare and its associated predators.

Simulated fire behaviour in young, postfire lodgepole pine forests

2017 ◽  
Vol 26 (10) ◽  
pp. 852 ◽  
Author(s):  
Kellen N. Nelson ◽  
Monica G. Turner ◽  
William H. Romme ◽  
Daniel B. Tinker
Keyword(s):  
Pinus Contorta ◽  
Stand Structure ◽  
Lodgepole Pine ◽  
Pine Forests ◽  
Fuel Moisture ◽  
Crown Fire ◽  
Fire Behaviour ◽  
Fire Initiation ◽  
Moisture Conditions ◽  
Simulated Fire

Early-seral forests are expanding throughout western North America as fire frequency and annual area burned increase, yet fire behaviour in young postfire forests is poorly understood. We simulated fire behaviour in 24-year-old lodgepole pine (Pinus contorta var. latifolia) stands in Yellowstone National Park, Wyoming, United States using operational models parameterised with empirical fuel characteristics, 50–99% fuel moisture conditions, and 1–60kmhr−1 open winds to address two questions: [1] How does fireline intensity, and crown fire initiation and spread vary among young, lodgepole pine stands? [2] What are the contributions of fuels, moisture and wind on fire behaviour? Sensitivity analysis indicated the greatest contributors to output variance were stand structure mediated wind attenuation, shrub fuel loads and 1000-h fuel moisture for fireline intensity; crown base height for crown fire initiation; and crown bulk density and 1-h fuel moisture for crown fire spread. Simulation results predicted crown fire (e.g. passive, conditional or active types) in over 90% of stands at 50th percentile moisture conditions and wind speeds greater than 3kmhr−1. We conclude that dense canopy characteristics heighten crown fire potential in young, postfire lodgepole pine forests even under less than extreme wind and fuel moisture conditions.

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